182 lines
22 KiB
XML
182 lines
22 KiB
XML
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<mods:title id="42145332411841642AEC68362F6D166E">A monograph of the Afrotropical Cassidinae (Coleoptera: Chrysomelidae). Part 6. Revision of the tribe Cassidini 3, the genus Cassida L.</mods:title>
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<mods:namePart id="FD02FDA608EE77AD9B04170AC8A4B92E">Borowiec, Lech</mods:namePart>
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<mods:nameIdentifier id="B220B92D9C1548C36999D9291BE777FD" type="email">lech.borowiec@uwr.edu.pl</mods:nameIdentifier>
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<mods:namePart id="82B4043822B6E8960D9CAD5F2A1A8D03">Świętojańska, Jolanta</mods:namePart>
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<mods:nameIdentifier id="AF4091D2DC52B7B2D008675271D8E299" type="email">jolanta.swietojanska@uwr.edu.pl</mods:nameIdentifier>
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<mods:title id="F38B5BBA7249BEE204F45D0508597DE9">Zootaxa</mods:title>
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<mods:date id="C78BE6C2FE8737D9582A1F4D0FE19479">2022</mods:date>
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<mods:number id="DB96808147C1381907D5E605A5A6D3DB">2022-08-04</mods:number>
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<treatment id="E417E526B1CEA9B9FF6CF8A50922941B" ID-DOI="http://doi.org/10.5281/zenodo.6966579" ID-GBIF-Taxon="198839245" ID-Zenodo-Dep="6966579" LSID="urn:lsid:plazi:treatment:E417E526B1CEA9B9FF6CF8A50922941B" httpUri="http://treatment.plazi.org/id/E417E526B1CEA9B9FF6CF8A50922941B" lastPageId="146" lastPageNumber="147" pageId="145" pageNumber="146">
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<subSubSection id="24A407BBB1CEA9BAFF6CF8A50E7397AA" box="[151,528,726,753]" pageId="145" pageNumber="146" type="nomenclature">
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<paragraph id="6C015430B1CEA9BAFF6CF8A50E7397AA" blockId="145.[151,528,726,789]" box="[151,528,726,753]" pageId="145" pageNumber="146">
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<heading id="3749E35CB1CEA9BAFF6CF8A50E7397AA" bold="true" box="[151,528,726,753]" fontSize="11" level="2" pageId="145" pageNumber="146" reason="4">
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<taxonomicName id="ABBE2FB3B1CEA9BAFF6CF8A50E7397AA" ID-CoL="7RYXD" authority="Spaeth, 1941" authorityName="Spaeth" authorityYear="1941" box="[151,528,726,753]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="145" pageNumber="146" phylum="Arthropoda" rank="species" species="somalica">
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<emphasis id="5ECA8822B1CEA9BAFF6CF8A50E7397AA" bold="true" box="[151,528,726,753]" pageId="145" pageNumber="146">
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<emphasis id="5ECA8822B1CEA9BAFF6CF8A50D0A97AB" bold="true" box="[151,361,726,753]" italics="true" pageId="145" pageNumber="146">Cassida somalica</emphasis>
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<bibRefCitation id="082F29C1B1CEA9BAFE8BF8A50E7397AA" author="Spaeth, F." box="[368,528,726,752]" pageId="145" pageNumber="146" pagination="315 - 318" refId="ref188404" refString="Spaeth, F. (1941) Neue Cassidinen aus Italienish-Ost-Afrika des Museo Civico di Storia Naturale - Trieste. Atti del Museo Civico di Storia Naturale di Trieste, 14, 315 - 318." type="journal article" year="1941">Spaeth, 1941</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="24A407BBB1CEA9BAFF6CF8880D0D964F" box="[151,366,763,789]" pageId="145" pageNumber="146" type="description">
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<paragraph id="6C015430B1CEA9BAFF6CF8880D0D964F" blockId="145.[151,528,726,789]" box="[151,366,763,789]" pageId="145" pageNumber="146">
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(
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<figureCitation id="F48548B5B1CEA9BAFF64F8880C94964F" box="[159,247,763,789]" captionStart="FIGURE 48" captionStartId="166.[151,250,1607,1631]" captionTargetBox="[159,1426,393,1573]" captionTargetId="figure-129@166.[151,1436,381,1584]" captionTargetPageId="166" captionText="FIGURE 48. Distribution of Cassida troglodytes (black circles) and Cassida somalica (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6966380" httpUri="https://zenodo.org/record/6966380/files/figure.png" pageId="145" pageNumber="146">figs. 48</figureCitation>
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,
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<figureCitation id="F48548B5B1CEA9BAFEF8F8880D05964F" box="[259,358,763,789]" captionStart="FIGURES 340–341" captionStartId="244.[151,264,1330,1354]" captionTargetBox="[182,1402,191,1288]" captionTargetId="figure-36@244.[168,1420,181,1305]" captionTargetPageId="244" captionText="FIGURES 340–341. Cassida somalica. 340. Dorsal; 341. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966645" httpUri="https://zenodo.org/record/6966645/files/figure.png" pageId="145" pageNumber="146">340–341</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="24A407BBB1CEA9BAFF6CF94C0F67960D" box="[151,772,831,855]" pageId="145" pageNumber="146" type="reference_group">
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<paragraph id="6C015430B1CEA9BAFF6CF94C0F67960D" blockId="145.[151,772,831,855]" box="[151,772,831,855]" pageId="145" pageNumber="146">
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<treatmentCitationGroup id="4CAE731EB1CEA9BAFF6CF94C0F67960D" box="[151,772,831,855]" pageId="145" pageNumber="146">
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<treatmentCitation id="ED1F7221B1CEA9BAFF6CF94C0E6F960D" author="Spaeth, F." box="[151,524,831,855]" page="317" pageId="145" pageNumber="146" year="1941">
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<taxonomicName id="ABBE2FB3B1CEA9BAFF6CF94C0E6F960D" ID-CoL="7RYXD" authority="Spaeth, 1941: 317" authorityName="Spaeth" authorityPageNumber="317" authorityYear="1941" box="[151,524,831,855]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="145" pageNumber="146" phylum="Arthropoda" rank="species" species="somalica">
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<emphasis id="5ECA8822B1CEA9BAFF6CF94C0D29960D" box="[151,330,831,855]" italics="true" pageId="145" pageNumber="146">Cassida somalica</emphasis>
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<bibRefCitation id="082F29C1B1CEA9BAFEAAF94C0E6F960D" author="Spaeth, F." box="[337,524,831,855]" pageId="145" pageNumber="146" pagination="315 - 318" refId="ref188404" refString="Spaeth, F. (1941) Neue Cassidinen aus Italienish-Ost-Afrika des Museo Civico di Storia Naturale - Trieste. Atti del Museo Civico di Storia Naturale di Trieste, 14, 315 - 318." type="journal article" year="1941">Spaeth, 1941: 317</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
;
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<bibRefCitation id="082F29C1B1CEA9BAFDEDF94C0EDB960D" author="Borowiec, L." box="[534,696,831,855]" pageId="145" pageNumber="146" refId="ref184613" refString="Borowiec, L. (1999) A world catalogue of the Cassidinae (Coleoptera: Chrysomelidae). Biologica Silesiae, Wroclaw, 476 pp." type="book" year="1999">Borowiec, 1999</bibRefCitation>
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a: 285.
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="24A407BBB1CEA9BAFF6CF9F6087196C5" box="[151,1042,901,927]" pageId="145" pageNumber="146" type="etymology">
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<paragraph id="6C015430B1CEA9BAFF6CF9F6087196C5" blockId="145.[151,1437,901,2008]" box="[151,1042,901,927]" pageId="145" pageNumber="146">
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<emphasis id="5ECA8822B1CEA9BAFF6CF9F60D7D96C5" bold="true" box="[151,286,901,927]" pageId="145" pageNumber="146">Etymology.</emphasis>
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Named after its locus typicus,
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<collectingRegion id="AE7A9AD2B1CEA9BAFD82F9F60F4296C5" box="[633,801,901,927]" country="Ethiopia" name="Sumale" pageId="145" pageNumber="146">Somali Region</collectingRegion>
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in southern
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<collectingCountry id="14A914A0B1CEA9BAFC57F9F6086C96C5" box="[940,1039,901,927]" name="Ethiopia" pageId="145" pageNumber="146">Ethiopia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="24A407BBB1CEA9BAFF3CF9DA0E71921D" pageId="145" pageNumber="146" type="description">
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<paragraph id="6C015430B1CEA9BAFF3CF9DA0D6B96BD" blockId="145.[151,1437,901,2008]" pageId="145" pageNumber="146">
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<emphasis id="5ECA8822B1CEA9BAFF3CF9DA0D3A9699" bold="true" box="[199,345,937,963]" pageId="145" pageNumber="146">Description.</emphasis>
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L: 4.40 mm, W: 3.55 mm, Lp: 1.70 mm, Wp: 2.90 mm, L/W: 1.24, Wp/Lp: 1.71. Body short–oval (
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<figureCitation id="F48548B5B1CEA9BAFF64F9BE0C9F96BD" box="[159,252,973,999]" captionStart="FIGURES 340–341" captionStartId="244.[151,264,1330,1354]" captionTargetBox="[182,1402,191,1288]" captionTargetId="figure-36@244.[168,1420,181,1305]" captionTargetPageId="244" captionText="FIGURES 340–341. Cassida somalica. 340. Dorsal; 341. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966645" httpUri="https://zenodo.org/record/6966645/files/figure.png" pageId="145" pageNumber="146">fig. 340</figureCitation>
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).
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</paragraph>
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<paragraph id="6C015430B1CEA9BAFF3CF98208B09151" blockId="145.[151,1437,901,2008]" box="[199,1235,1009,1036]" pageId="145" pageNumber="146">
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Entire body, including head, ventrites, legs and antennae ochraceous yellow (
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<figureCitation id="F48548B5B1CEA9BAFBDCF98208A59151" box="[1063,1222,1009,1035]" captionStart="FIGURES 340–341" captionStartId="244.[151,264,1330,1354]" captionTargetBox="[182,1402,191,1288]" captionTargetId="figure-36@244.[168,1420,181,1305]" captionTargetPageId="244" captionText="FIGURES 340–341. Cassida somalica. 340. Dorsal; 341. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966645" httpUri="https://zenodo.org/record/6966645/files/figure.png" pageId="145" pageNumber="146">figs. 340, 341</figureCitation>
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).
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</paragraph>
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<paragraph id="6C015430B1CEA9BAFF3CFE650D62919A" blockId="145.[151,1437,901,2008]" pageId="145" pageNumber="146">Pronotum elliptical, with maximum width in the middle, anterior margin regularly convex, sides narrowly rounded, no basal corners. Disc indistinctly bordered from explanate margin but with short, deep lateral impressions. Surface of disc shiny, with fine and sparse punctation. Distance between punctures several times wider than puncture diameter, interspaces regular. Explanate margin broad, impunctate, shiny, transparent with well visible honeycomb structure.</paragraph>
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<paragraph id="6C015430B1CEA9BAFF3CFEB908AE935E" blockId="145.[151,1437,901,2008]" pageId="145" pageNumber="146">
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Base of elytra only slightly wider than base of pronotum, humeral angles distinctly protruding anterad, angulate. Disc convex in profile (
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<figureCitation id="F48548B5B1CEA9BAFE51FE9E0E6A905D" box="[426,521,1261,1287]" captionStart="FIGURES 340–341" captionStartId="244.[151,264,1330,1354]" captionTargetBox="[182,1402,191,1288]" captionTargetId="figure-36@244.[168,1420,181,1305]" captionTargetPageId="244" captionText="FIGURES 340–341. Cassida somalica. 340. Dorsal; 341. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966645" httpUri="https://zenodo.org/record/6966645/files/figure.png" pageId="145" pageNumber="146">fig. 340</figureCitation>
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), with distinct postscutellar and principal impressions, low H–shaped elevation, slightly elevated second interval and some elevated interspaces between rows 1 and 2 and 3 and 6. Punctation coarse arranged in completely regular rows, dense, distance between punctures mostly narrower than puncture diameter, on sides punctures almost touching each other. Marginal row distinct, its punctures as coarse as punctures in central rows. Intervals mostly as wide as or slightly narrower than rows, their surface shiny, marginal interval as wide as submarginal interval and submarginal row combined, without humeral but with flat lateral folds. Explanate margin narrow, moderately declivous, in the widest part five times narrower than disc, surface shiny with punctation and transverse grooves appears slightly irregular, transparent with well visible honeycomb structure.
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</paragraph>
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<paragraph id="6C015430B1CEA9BAFF3CFC7D0EB493ED" blockId="145.[151,1437,901,2008]" pageId="145" pageNumber="146">Eyes very large, gena obsolete. Clypeus broad, approximately 1.3 times as wide as long. Clypeal grooves fine but well marked, converging in regular triangle, surface of clypeus flat, shiny with few very small, setose punctures. Labrum broadly emarginate to 1/5 length. Antennae stout, segments 9–10 slightly transverse. Length ratio of antennal segments: 100:57:64:64:61:50:54:46:46:50:107. Segment 3 approximately 1.1 times as long as segment 2 and approximately as long as segment 4.</paragraph>
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<paragraph id="6C015430B1CEA9BAFF3CFCB10954927E" blockId="145.[151,1437,901,2008]" pageId="145" pageNumber="146">Prosternum broad in the middle, strongly expanded apically, area between coxa canaliculate along sides with rows of setose punctures, central part convex, shiny, expanded apex only slightly convex in the middle, shallowly impressed laterally with several longitudinal wrinkles and small setose punctures, surface appears rugose.</paragraph>
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<paragraph id="6C015430B1CEA9BAFF3CFD5E0E71921D" blockId="145.[151,1437,901,2008]" box="[199,530,1837,1863]" pageId="145" pageNumber="146">Claws with small basal tooth.</paragraph>
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</subSubSection>
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<subSubSection id="24A407BBB1CEA9BAFF3CFD220E249231" box="[199,583,1873,1899]" pageId="145" pageNumber="146" type="distribution">
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||
<paragraph id="6C015430B1CEA9BAFF3CFD220E249231" blockId="145.[151,1437,901,2008]" box="[199,583,1873,1899]" pageId="145" pageNumber="146">
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<emphasis id="5ECA8822B1CEA9BAFF3CFD220D029231" bold="true" box="[199,353,1873,1899]" pageId="145" pageNumber="146">Distribution.</emphasis>
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S
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<collectingCountry id="14A914A0B1CEA9BAFE85FD220DBE9231" box="[382,477,1873,1899]" name="Ethiopia" pageId="145" pageNumber="146">Ethiopia</collectingCountry>
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||
(
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<figureCitation id="F48548B5B1CEA9BAFE17FD220E599231" box="[492,570,1873,1899]" captionStart="FIGURE 48" captionStartId="166.[151,250,1607,1631]" captionTargetBox="[159,1426,393,1573]" captionTargetId="figure-129@166.[151,1436,381,1584]" captionTargetPageId="166" captionText="FIGURE 48. Distribution of Cassida troglodytes (black circles) and Cassida somalica (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6966380" httpUri="https://zenodo.org/record/6966380/files/figure.png" pageId="145" pageNumber="146">fig. 48</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="24A407BBB1CEA9B9FF3CFD060DD99447" lastPageId="146" lastPageNumber="147" pageId="145" pageNumber="146" type="discussion">
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||
<paragraph id="6C015430B1CEA9B9FF3CFD060DD99447" blockId="145.[151,1437,901,2008]" lastBlockId="146.[151,1436,151,321]" lastPageId="146" lastPageNumber="147" pageId="145" pageNumber="146">
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||
<emphasis id="5ECA8822B1CEA9BAFF3CFD060D5892D5" bold="true" box="[199,315,1909,1935]" pageId="145" pageNumber="146">Remarks.</emphasis>
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A distinct species with no close relatives among African members of the genus
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<taxonomicName id="ABBE2FB3B1CEA9BAFB63FD06089192D5" authorityName="Linnaeus" authorityYear="1758" box="[1176,1266,1909,1935]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="145" pageNumber="146" phylum="Arthropoda" rank="genus">
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<emphasis id="5ECA8822B1CEA9BAFB63FD06089192D5" box="[1176,1266,1909,1935]" italics="true" pageId="145" pageNumber="146">Cassida</emphasis>
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</taxonomicName>
|
||
. Its ochraceous yellow dorsal colouration without a pattern, rounded pronotal sides, elytral disc with H–shaped elevation but regularly convex in profile, coarse and dense elytral punctation arranged in completely regular rows and claws with small basal tooth are unique characters (
|
||
<figureCitation id="F48548B5B1CDA9B9FDA5FAE40E9D95EB" box="[606,766,151,177]" captionStart="FIGURES 340–341" captionStartId="244.[151,264,1330,1354]" captionTargetBox="[182,1402,191,1288]" captionTargetId="figure-36@244.[168,1420,181,1305]" captionTargetPageId="244" captionText="FIGURES 340–341. Cassida somalica. 340. Dorsal; 341. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966645" httpUri="https://zenodo.org/record/6966645/files/figure.png" pageId="146" pageNumber="147">figs. 340, 341</figureCitation>
|
||
). The pale form of
|
||
<taxonomicName id="ABBE2FB3B1CDA9B9FC22FAE4083395EB" authorityName="Borowiec & Świętojańska" authorityYear="2022" box="[985,1104,151,177]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="146" pageNumber="147" phylum="Tracheophyta" rank="species" species="villiersi">
|
||
<emphasis id="5ECA8822B1CDA9B9FC22FAE4083395EB" box="[985,1104,151,177]" italics="true" pageId="146" pageNumber="147">C. villiersi</emphasis>
|
||
</taxonomicName>
|
||
appears similar but differs in a slimmer body with L/W ratio 1.27–1.37 (in
|
||
<taxonomicName id="ABBE2FB3B1CDA9B9FD61FAC80F42958F" authorityName="Spaeth" authorityYear="1941" box="[666,801,187,213]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="146" pageNumber="147" phylum="Tracheophyta" rank="species" species="somalica">
|
||
<emphasis id="5ECA8822B1CDA9B9FD61FAC80F42958F" box="[666,801,187,213]" italics="true" pageId="146" pageNumber="147">C. somalica</emphasis>
|
||
</taxonomicName>
|
||
1.24) and in the punctate explanate margin of pronotum (fig. 207). Also a pale form of
|
||
<taxonomicName id="ABBE2FB3B1CDA9B9FE13FAAC0E0195A2" authorityName="Borowiec & Świętojańska" authorityYear="2022" box="[488,610,223,249]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="146" pageNumber="147" phylum="Tracheophyta" rank="species" species="voiensis">
|
||
<emphasis id="5ECA8822B1CDA9B9FE13FAAC0E0195A2" box="[488,610,223,249]" italics="true" pageId="146" pageNumber="147">C. voiensis</emphasis>
|
||
</taxonomicName>
|
||
appears similar but differs in a more elongate and slimmer body with L/W ratio 1.32–1.42 (fig. 196).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="24A407BBB1CDA9B9FF3CFB540922941B" box="[199,1345,294,321]" pageId="146" pageNumber="147" type="materials_examined">
|
||
<paragraph id="6C015430B1CDA9B9FF3CFB540922941B" blockId="146.[151,1436,151,321]" box="[199,1345,294,321]" pageId="146" pageNumber="147">
|
||
<emphasis id="5ECA8822B1CDA9B9FF3CFB540DE0941A" bold="true" box="[199,387,294,321]" pageId="146" pageNumber="147">Type examined.</emphasis>
|
||
<materialsCitation id="DCD65E6DB1CDA9B9FE71FB540922941B" ID-GBIF-Occurrence="3881325135" box="[394,1345,295,321]" collectingDate="1897-10-14" collectionCode="MCSNT" country="Ethiopia" county="Vatova" location="Somalia" municipality="Bogol Magno" pageId="146" pageNumber="147" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="B305EA92B1CDA9B9FE71FB540D94941B" box="[394,503,295,321]" pageId="146" pageNumber="147" type="holotype">Holotype</typeStatus>
|
||
: [
|
||
<collectingCountry id="14A914A0B1CDA9B9FDF1FB540EF0941B" box="[522,659,295,321]" name="Ethiopia" pageId="146" pageNumber="147">ETHIOPIA</collectingCountry>
|
||
]:
|
||
<location id="696102EBB1CDA9B9FD58FB540E9C941B" LSID="urn:lsid:plazi:treatment:E417E526B1CEA9B9FF6CF8A50922941B:696102EBB1CDA9B9FD58FB540E9C941B" box="[675,767,295,321]" country="Ethiopia" county="Vatova" municipality="Bogol Magno" name="Somalia" pageId="146" pageNumber="147">Somalia</location>
|
||
mer.,
|
||
<collectingMunicipality id="8C65CE4AB1CDA9B9FCBEFB540F80941B" box="[837,995,295,321]" pageId="146" pageNumber="147">Bogol Magno</collectingMunicipality>
|
||
,
|
||
<date id="180072F0B1CDA9B9FC15FB540804941B" box="[1006,1127,295,321]" pageId="146" pageNumber="147" value="1897-10-14">
|
||
<collectingDate id="08448B18B1CDA9B9FC15FB540804941B" box="[1006,1127,295,321]" pageId="146" pageNumber="147" value="1897-10-14">14 X 1897</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingCounty id="85602CBCB1CDA9B9FB89FB5408A3941B" box="[1138,1216,295,321]" pageId="146" pageNumber="147">Vatova</collectingCounty>
|
||
(
|
||
<collectionCode id="0AAFCCF5B1CDA9B9FB35FB540956941B" box="[1230,1333,295,321]" pageId="146" pageNumber="147">MCSNT</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |