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<document ID-CLB-Dataset="187404" ID-DOI="http://dx.doi.org/10.3897/zookeys.1129.87706" ID-GBIF-Dataset="d2ef8eec-1ffa-4d5d-ba17-fb7eba1e850b" ID-PMC="PMC9836715" ID-Pensoft-Pub="1313-2970-1129-85" ID-Pensoft-UUID="4AB339E1FB775670A7A051080595E617" ID-PubMed="36761843" ID-ZooBank="CF6FB3C5FD724040913B371386CA9B9A" ModsDocID="1313-2970-1129-85" checkinTime="1668554634167" checkinUser="pensoft" docAuthor="Wang, Ning, Huang, Huateng &amp; Ma, Li-Bin" docDate="2022" docId="B188F7B0633655AD91F91F3BA1430361" docLanguage="en" docName="ZooKeys 1129: 85-107" docOrigin="ZooKeys 1129" docPubDate="2022-11-15" docSource="http://dx.doi.org/10.3897/zookeys.1129.87706" docTitle="Gymnogryllus odonopetalus Xie &amp; Zheng 2003" docType="treatment" docVersion="5" id="4AB339E1FB775670A7A051080595E617" lastPageNumber="85" masterDocId="4AB339E1FB775670A7A051080595E617" masterDocTitle="The intraspecific variation of morphology and coloration of field crickets: a taxonomic revision of Chinese Gymnogryllus Saussure, 1877 and Phonarellus Gorochov, 1983 (Orthoptera, Gryllidae, Gryllini)" masterLastPageNumber="107" masterPageNumber="85" pageNumber="85" updateTime="1732787025633" updateUser="ExternalLinkService">
<mods:mods id="C92B636C12DD3B3E4F858D1FAF2B51C4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DC5E43197E030206C5657A080410E16A">
<mods:title id="1B0A87D3D94E204FA873969BD77DCE67">The intraspecific variation of morphology and coloration of field crickets: a taxonomic revision of Chinese Gymnogryllus Saussure, 1877 and Phonarellus Gorochov, 1983 (Orthoptera, Gryllidae, Gryllini)</mods:title>
</mods:titleInfo>
<mods:name id="3F4A9029C81E7206B73D2F5FD0D48841" type="personal">
<mods:role id="C68F9B78160E68DA7B258B5327158F80">
<mods:roleTerm id="E654FCE2421BB4E5C98351627D1C8331">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4ADC5198677F332AFFE8A8FDF2A47FE3">Wang, Ning</mods:namePart>
<mods:nameIdentifier id="C444AAF2EB042E7BB51A45083B12CC4E" type="ORCID">https://orcid.org/0000-0002-2367-4388</mods:nameIdentifier>
<mods:affiliation id="62B56428A9DD38A478978385050B5D25">College of Life Sciences, Shaanxi Normal University, Xi'an, China, 710119</mods:affiliation>
</mods:name>
<mods:name id="26A4CAB8453B02F0E2B5EFA08048AEC8" type="personal">
<mods:role id="5646D7B88C26A667A7ADBF0E5945D0D5">
<mods:roleTerm id="BDB7F7A9F35CBD5E982B56E614AB44CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B7C4591813289327A247F5E4EA27E40">Huang, Huateng</mods:namePart>
<mods:nameIdentifier id="BF63AD6291C520466D1AB11004596868" type="ORCID">https://orcid.org/0000-0002-0011-2091</mods:nameIdentifier>
<mods:affiliation id="6DD621015DEB49A41D0FD1C2B87A51CF">College of Life Sciences, Shaanxi Normal University, Xi'an, China, 710119</mods:affiliation>
<mods:nameIdentifier id="EE326A6D3DBF3CD75FC2C9C5F55E04A5" type="email">huanghuateng@snnu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="46C02CDC35A9F5D2F80196676A319EF4" type="personal">
<mods:role id="8D4755C57B4C2E0D232CE115A252F390">
<mods:roleTerm id="5F6F963745B15527452FAE2220AED9B1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8A5F7BFD6A9914A17F9269577ADDD671">Ma, Li-Bin</mods:namePart>
<mods:nameIdentifier id="A547694F00DA03A4405F71B2EA30F8AF" type="ORCID">https://orcid.org/0000-0002-8556-7158</mods:nameIdentifier>
<mods:affiliation id="66D5F92E7E1530CCDE0C4909AB1B9509">College of Life Sciences, Shaanxi Normal University, Xi'an, China, 710119</mods:affiliation>
<mods:nameIdentifier id="C9BA55A68C53116F2594B7736D83CCE9" type="email">libinma@foxmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A8AEA487222A36E35880859C0AEC70E7">text</mods:typeOfResource>
<mods:relatedItem id="56B8F6593E93E3B27881EC83B19FDBEB" type="host">
<mods:titleInfo id="DC9A129E07993BC2E6D87C31B621D9D7">
<mods:title id="F09813A3F16753D74DBF066346AC89A8">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="720F036F0182D60B82780D534E8B7059">
<mods:date id="2E8C929A38244A0C8EC951FEF1649EE2">2022</mods:date>
<mods:detail id="FA549112A804514058FE609E28F8E338" type="pubDate">
<mods:number id="B294081CCB03502A71BA653439DA7921">2022-11-15</mods:number>
</mods:detail>
<mods:detail id="93A0B41D308B0FA6673D255DE60D952E" type="volume">
<mods:number id="69FD23EFB89AD9E09DCFB7CB7EAD714A">1129</mods:number>
</mods:detail>
<mods:extent id="892DF2CB2CBC78071588751B3800379E" unit="page">
<mods:start id="1F58691E2E7C3F3FB5ADA8E16D9445BE">85</mods:start>
<mods:end id="715F4FC321070C3FAE38BD2AD643C760">107</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C6FEE87BEA01A695406DF5C602559511">
<mods:url id="52E6CD9F23DE85A0C7775419E00398A6">http://dx.doi.org/10.3897/zookeys.1129.87706</mods:url>
</mods:location>
<mods:classification id="2F18F263A2295A962A6CE74F6D1EB65C">journal article</mods:classification>
<mods:identifier id="99DF1C27F6901431F951A0232E9A5EF0" type="DOI">http://dx.doi.org/10.3897/zookeys.1129.87706</mods:identifier>
<mods:identifier id="FC25F4AEFBAB49E656D9BAA381427C15" type="Pensoft-Pub">1313-2970-1129-85</mods:identifier>
<mods:identifier id="E60C443A48C41DB4790BF39458D95E16" type="ZooBank">CF6FB3C5FD724040913B371386CA9B9A</mods:identifier>
<mods:identifier id="1B0BF9D8D824ACEBFB0F0A708D5BAE89" type="Pensoft-UUID">4AB339E1FB775670A7A051080595E617</mods:identifier>
</mods:mods>
<treatment id="B188F7B0633655AD91F91F3BA1430361" ID-GBIF-Taxon="205021276" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361" httpUri="http://treatment.plazi.org/id/B188F7B0633655AD91F91F3BA1430361" lastPageNumber="85" pageId="0" pageNumber="85" scope_family="Gryllidae" scope_order="Orthoptera" scope_tribe="Gryllini">
<subSubSection id="E9CF815F09CC6D01EEB6F507FBBD3143" pageId="0" pageNumber="85" type="nomenclature">
<paragraph id="22C92D380CE17ED1CA47EAAE9BF509FA" pageId="0" pageNumber="85">
<taxonomicName id="D81D344CC9E3D38092E5EA738A8DD673" ID-CoL="3HSLJ" LSID="B188F7B0-6336-55AD-91F9-1F3BA1430361" authority="Xie &amp; Zheng, 2003" authorityName="Xie &amp; Zheng" authorityYear="2003" class="Insecta" family="Gryllidae" genus="Gymnogryllus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gymnogryllus odonopetalus" order="Orthoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="odonopetalus">Gymnogryllus odonopetalus Xie &amp; Zheng, 2003</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="02FE118BAD03484442615FC31600765A" pageId="0" pageNumber="85" type="description">
<paragraph id="C8A6AA009B2E0D696A5F15FB98CC1F6A" pageId="0" pageNumber="85">
<figureCitation id="E69AE7E8758D66A274B7FE2BC55D177F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Living photos of some Phonarellus and Gymnogryllus species A, B male of P. minor (with varing proportion of black areas on hind legs) C female of P. minor D male of P. ritsemae E male of G. odonopetalus F female of G. odonopetalus (Photos A-D were provided by Zhang, Tao, and E, F were photographed by He, Zhixin)." figureDoi="10.3897/zookeys.1129.87706.figure1" httpUri="https://binary.pensoft.net/fig/770016" pageId="0" pageNumber="85">Figs 1E-F</figureCitation>
<figureCitation id="DC1224096FF8A911AE185AB7F4EFCEE2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bodies of G. odonopetalus A male B female. Scale bar: 10 mm." figureDoi="10.3897/zookeys.1129.87706.figure3" httpUri="https://binary.pensoft.net/fig/770018" pageId="0" pageNumber="85">, 3</figureCitation>
<figureCitation id="6622610A8D2FABA8D295A63670A580FF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Intraspecific variation of genitalia in G. odonopetalus A-F genitalia from six individuals in lateral view (Note: the red arrows point to the posterior of epiphallus, highlighting variations among individuals; the blue circles indicate the protuberance of mid-ectoparamere) G-L genitalia of the same six individuals in dorsal view (Note: the green circles indicate the median lobe of the epiphallus posterior)." figureDoi="10.3897/zookeys.1129.87706.figure4" httpUri="https://binary.pensoft.net/fig/770019" pageId="0" pageNumber="85">, 4</figureCitation>
<figureCitation id="4128302405853F6F1307E68C2DD406E2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Intraspecific variation of genitalia in G. odonopetalus A-C and G-I photos of genitalia from six individuals in caudal view D-F and J-L drawings of genitalia from the same six individuals in caudal view." figureDoi="10.3897/zookeys.1129.87706.figure5" httpUri="https://binary.pensoft.net/fig/770020" pageId="0" pageNumber="85">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7D8E0D5BD646216198B707E02BB9097E" pageId="0" pageNumber="85" type="reference_group">
<paragraph id="488434C29786FFE4D1B827110FD66FD3" pageId="0" pageNumber="85">
<taxonomicName id="FF932D5FF5B08D8D16EFDAC0C737DC05" authorityName="Xie &amp; Zheng" authorityYear="2003" class="Insecta" family="Gryllidae" genus="Gymnogryllus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gymnogryllus odonopetalus" order="Orthoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="odonopetalus">Gymnogryllus odonopetalus</taxonomicName>
Xie &amp; Zheng, 2003: 496, 498.
</paragraph>
<paragraph id="F3AD59CC9484FBCAC6809184514EB2B4" pageId="0" pageNumber="85">
<taxonomicName id="B2CED73C291D1E65880A38236A4B0400" authorityName="Ma, Libin &amp; Yalin Zhang" authorityYear="2011" class="Insecta" family="Gryllidae" genus="Gymnogryllus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gymnogryllus yunnanensis" order="Orthoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="yunnanensis">Gymnogryllus yunnanensis</taxonomicName>
Ma &amp; Zhang, 2011: 31-40, syn. nov.
</paragraph>
<paragraph id="F186CFDB1841C8C04953EF79C9A9FB34" pageId="0" pageNumber="85">
<taxonomicName id="B3EA223D18C52A9EE8F69DD89A621CC6" authorityName="Ma, Libin &amp; Yalin Zhang" authorityYear="2011" class="Insecta" family="Gryllidae" genus="Gymnogryllus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gymnogryllus longus" order="Orthoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="longus">Gymnogryllus longus</taxonomicName>
Ma &amp; Zhang, 2011: 31-40, syn. nov.
</paragraph>
<paragraph id="8532368F5798A5C601B5EE82992C1855" pageId="0" pageNumber="85">
<taxonomicName id="D54C859527A68F0B16301EBFC203D4AD" authorityName="Ma, Libin &amp; Yalin Zhang" authorityYear="2011" class="Insecta" family="Gryllidae" genus="Gymnogryllus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gymnogryllus tumidulus" order="Orthoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="tumidulus">Gymnogryllus tumidulus</taxonomicName>
Ma &amp; Zhang, 2011: 31-40, syn. nov.
</paragraph>
<paragraph id="B067DC8C016C72CE5DD740A3942D7F7A" pageId="0" pageNumber="85">
<taxonomicName id="DA4E1A6FB62DD31998EB33A0C3102FDC" authorityName="Ma, Libin &amp; Yalin Zhang" authorityYear="2011" class="Insecta" family="Gryllidae" genus="Gymnogryllus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gymnogryllus striatus" order="Orthoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="striatus">Gymnogryllus striatus</taxonomicName>
Ma &amp; Zhang, 2011: 31-40, syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="BFDDC4A3FA1D1FB13B9E9191FC2A787C" pageId="0" pageNumber="85" type="holotype">
<paragraph id="66EE635012EBC3FCA75956ABF9246499" pageId="0" pageNumber="85">Holotype.</paragraph>
<paragraph id="849C186CEF2C2F81686C23660D33B5B6" pageId="0" pageNumber="85">
Type locality: Menglun, Xishuangbanna, Yunnan, China. Deposited at Museum of Flora and Fauna of Shaanxi Normal University,
<normalizedToken id="EAE65BFE4E9A21AA5F6A86CD23C49F39" originalValue="Xi’an">Xi'an</normalizedToken>
, China (SNNU).
</paragraph>
</subSubSection>
<subSubSection id="62B9EB32CBE251EFA12B4E17A2B3B297" pageId="0" pageNumber="85" type="materials_examined">
<paragraph id="54B2E3F15C077C1FDEB244F2CDE6BABD" pageId="0" pageNumber="85">Specimens examined.</paragraph>
<paragraph id="0C726C7E505490E611FA87FC70EC48A1" pageId="0" pageNumber="85">
<materialsCitation id="5168108FCA14C8ACE8DBDB862FDA0D19" collectingDate="1999-09-08" collectorName="Xie, Lingde" country="China" location="Menglun" municipality="Xishuangbanna" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="C0A311DDBBEE5623F8EA3438C68021D7" bold="true" pageId="0" pageNumber="85">
<collectingCountry id="A966695C8AE480C81D9F5CBA7A2684D5" name="China">China</collectingCountry>
</emphasis>
:
<specimenCount id="D011FBBE7175466E50B06D0848DA9E6C" type="male">1 male</specimenCount>
(
<typeStatus id="C6EC5FDCCF7C1B477EE446B57734BBD9">holotype</typeStatus>
),
<collectingRegion id="FBC8D608F02F55D2F142252DE67D9180" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingMunicipality id="C2AEA4D4C9A7078E891BD74D9BE6D5B3">Xishuangbanna</collectingMunicipality>
,
<location id="47614EDBEC8B9214CD00AFB406257F69" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361:47614EDBEC8B9214CD00AFB406257F69" country="China" municipality="Xishuangbanna" name="Menglun" stateProvince="Yunnan">Menglun</location>
,
<collectingDate id="E35941F9D36BB16E5E8DA4825694E8BE" value="1999-09-08">Sept. 8, 1999</collectingDate>
,
<collectorName id="B2F00FB6149E767AEBD78431064F1D7E">Xie</collectorName>
,
<collectorName id="9E8795CDAEC91C6064D6AB5B56D5A873">Lingde</collectorName>
coll. (SNNU)
</materialsCitation>
;
<materialsCitation id="4FE298B55C8E905DB9063541E47A8DF3" collectingDate="2009-06-11" collectorName="Ma, Libin" country="China" location="Wengdang" municipality="Honghe" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="2D058D5E4F6E70879C7C48FB9BFCE2AA" type="male">2 males</specimenCount>
and
<specimenCount id="AAE109723AF8774456075140846DD9F5" type="female">1 female</specimenCount>
,
<collectingRegion id="7DE38AFEF15E62D8CDA2168DF23ED72A" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingMunicipality id="1E7978D6D2314003B77F643AE8357ED0">Honghe</collectingMunicipality>
,
<location id="FEEBA21393E6ED3369873C9B60318D0F" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361:FEEBA21393E6ED3369873C9B60318D0F" country="China" municipality="Honghe" name="Wengdang" stateProvince="Yunnan">Wengdang</location>
,
<collectingDate id="EBD96AEF260A710CE05608E676704D0B" value="2009-06-11">Jun. 11, 2009</collectingDate>
,
<collectorName id="7AF88082AD77EB8F04F550CF918EF1E6">Ma</collectorName>
,
<collectorName id="8159D4C09D7BDD14CAEBFD71F044BB95">Libin</collectorName>
coll. (SNNU)
</materialsCitation>
;
<materialsCitation id="CC053F1974F8034E75C41BA36DED40BD" collectingDate="2013-05-13" collectorName="Ma, Libin" country="China" county="Mengla" location="Longmen" municipality="Shangyong" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="1EDA1CC1574F50153A31F8487F27E346" type="female">1 female</specimenCount>
,
<collectingRegion id="BE74A3B89E2EE9B79B4063D3201B61F2" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingCounty id="D8CC55F94DAB228CEC5432999992A345">Mengla</collectingCounty>
,
<collectingMunicipality id="F4E73C7A3CD437F5B9B231031AB4D84F">Shangyong</collectingMunicipality>
,
<location id="16B0CD68CA0BCF95F64CA9F171626B8B" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361:16B0CD68CA0BCF95F64CA9F171626B8B" country="China" county="Mengla" municipality="Shangyong" name="Longmen" stateProvince="Yunnan">Longmen</location>
,
<collectingDate id="556CB931259DA368BF94A6A5FAD4BD6E" value="2013-05-13">May 13, 2013</collectingDate>
,
<collectorName id="2C6EA9B22C1B85D5CE412CF1F54A0F3B">Ma</collectorName>
,
<collectorName id="80E9D9A6217D03EFCECE40B6AE73CC0C">Libin</collectorName>
coll. (SNNU)
</materialsCitation>
;
<materialsCitation id="E6A0175C9C942B5A1C1E32CA926E5CF9" collectingDate="2014-10-02" collectorName="Zhang, Tao" country="China" location="Wangtianshu" municipality="Mengla" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="1A3FE23B925BB9A1DEEC48DDE0577666" type="male">1 male</specimenCount>
,
<collectingRegion id="7DD2FA5444E7614D5B15B52DD4402EED" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingMunicipality id="7384B97F4CE2A7377FCF471E6B5D9FA9">Mengla</collectingMunicipality>
(or
<location id="F72931355012C2BFA1D7FB50B4009D71" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361:F72931355012C2BFA1D7FB50B4009D71" country="China" municipality="Mengla" name="Wangtianshu" stateProvince="Yunnan">Wangtianshu</location>
),
<collectingDate id="A76D605AF49256FEB5AFB28B3B06B408" value="2014-10-02">Oct. 2, 2014</collectingDate>
,
<collectorName id="6430B7B47927CC21E8CA9F6811456911">Zhang</collectorName>
,
<collectorName id="9E3278F27C1C3CD5177E104AA3EAB955">Tao</collectorName>
coll. (SNNU)
</materialsCitation>
;
<materialsCitation id="92471FEAF9992C44D9A9A7F32EBDBB57" collectingDate="2018-07-11" collectorName="Peng, Zhong" country="China" location="Jinghong" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="EC1CEF86745A0BD1F8C35D57BDADBAEE" type="male">3 males</specimenCount>
,
<collectingRegion id="655213CDF76CF15B55C6A720558B10B2" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<location id="8249D4A0803D65045FB991294BEB8121" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361:8249D4A0803D65045FB991294BEB8121" country="China" name="Jinghong" stateProvince="Yunnan">Jinghong</location>
,
<collectingDate id="AA836491CB13E04B139ECDAD786D655F" value="2018-07-11">Jul. 11, 2018</collectingDate>
,
<collectorName id="FA48F02A996A32A7A4B5BD80AFD3CC9C">Peng</collectorName>
,
<collectorName id="003C3773FE3F1DF399861F1F66940764">Zhong</collectorName>
coll. (SNNU)
</materialsCitation>
;
<materialsCitation id="F4882F0E458062D09F0EA7F878449A0C" collectingDate="2021-08-17" collectorName="He, Zhixin" country="China" location="Jinggu" municipality="Pu'er" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="1BD2E19F64F87D21DB50D6E81A402523" type="male">2 males</specimenCount>
,
<collectingRegion id="AC72B245C8B872BEDA359571C1982EBA" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<normalizedToken id="AD30F5F14F92A763B9C93211DA8F1833" originalValue="Pu’er">
<collectingMunicipality id="205E8A028374A0A1DDFF66B4D5849D67">Pu'er</collectingMunicipality>
</normalizedToken>
,
<location id="10608ECFF08356140B9A5B33C6D90E17" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361:10608ECFF08356140B9A5B33C6D90E17" country="China" municipality="Pu'er" name="Jinggu" stateProvince="Yunnan">Jinggu</location>
,
<collectingDate id="5D75E797CB8704FB62791D6D62CA0A6A" value="2021-08-17">Aug. 17, 2021</collectingDate>
,
<collectorName id="0F277C5F3A5C74E8BCD94D3A71689A1E">He</collectorName>
,
<collectorName id="6EC88C06060B6ABF0CE4B2EC569B7120">Zhixin</collectorName>
coll. (SNNU)
</materialsCitation>
;
<materialsCitation id="4F058F2B3C41105D869E6225B8953D78" collectingDate="2021-08-18" collectorName="He, Zhixin" country="China" location="Simaoqu" municipality="Pu'er" specimenCount="7" specimenCount-male="7" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="F25CDEFC6C7ED6783AE27E54CC7132FC" type="male">7 males</specimenCount>
,
<collectingRegion id="190D550EED2D83A3E830C452E3ED1AEE" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<normalizedToken id="9898F92A3A384BB7F40ACE47C8F9960A" originalValue="Pu’er">
<collectingMunicipality id="01BB0ECCA0B5601D22A141013B78F6A7">Pu'er</collectingMunicipality>
</normalizedToken>
,
<location id="28BE08BFA40901969D7157DC66322C29" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361:28BE08BFA40901969D7157DC66322C29" country="China" municipality="Pu'er" name="Simaoqu" stateProvince="Yunnan">Simaoqu</location>
,
<collectingDate id="C269D20692F23A4CEAFD9FC3D2247B92" value="2021-08-18">Aug. 18, 2021</collectingDate>
,
<collectorName id="9757FE6DC6A9D3347EF755EE5F621617">He</collectorName>
,
<collectorName id="C5130E7C68B31C7308CA07434953F0B0">Zhixin</collectorName>
coll. (SNNU)
</materialsCitation>
;
<materialsCitation id="7E950E0BF7B6BEAE95BA7762AF2E61EB" collectingDate="2021-08-25" collectorName="He, Zhixin" country="China" location="Menglun" municipality="Mengla" specimenCount="15" specimenCount-male="15" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="F31A0EC3D52AC2E4F4E42602875C9BE2" type="male">15 males</specimenCount>
,
<collectingRegion id="608A67174EA3063801CCD0AA56DE88FD" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingMunicipality id="53023AF327036AAAA785870F11C18BA9">Mengla</collectingMunicipality>
,
<location id="5CD33C6C670B04B737DA2335F90D5E2F" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361:5CD33C6C670B04B737DA2335F90D5E2F" country="China" municipality="Mengla" name="Menglun" stateProvince="Yunnan">Menglun</location>
,
<collectingDate id="DD78F0742136FC26E1936B8DD80AC729" value="2021-08-25">Aug. 25, 2021</collectingDate>
,
<collectorName id="46362187A90DB038FC01213332306EFA">He</collectorName>
,
<collectorName id="80F41D02388F932A2062BF5BC9FFED0C">Zhixin</collectorName>
coll. (SNNU)
</materialsCitation>
;
<materialsCitation id="F9D31D57179C8BD0AF1DF3F8C7897BA2" collectingDate="2021-08-25" collectorName="He, Zhixin" country="China" location="Menglun" municipality="Mengla" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="985E96FFD5CF64FC0BC45B424AB2460D" type="female">3 females</specimenCount>
,
<collectingRegion id="032A8EA08A7AFC7F6525E65ECE08D350" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingMunicipality id="9540CE2D39A0099C605A09EA59B2AB59">Mengla</collectingMunicipality>
,
<location id="5226CBEDB55306999EC0EEE32BCAA6F2" LSID="urn:lsid:plazi:treatment:B188F7B0633655AD91F91F3BA1430361:5226CBEDB55306999EC0EEE32BCAA6F2" country="China" municipality="Mengla" name="Menglun" stateProvince="Yunnan">Menglun</location>
,
<collectingDate id="B0AD2E4F7A6ABE935B9202DF22F96E3B" value="2021-08-25">Aug. 25, 2021</collectingDate>
,
<collectorName id="AB7D681FA8F3072CC313A422B031147D">He</collectorName>
,
<collectorName id="ADEB9B6984989710EB030B6A63A4849F">Zhixin</collectorName>
coll. (SNNU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9D4B5ECB11EC8B81D4E06FE194F8FFB2" pageId="0" pageNumber="85" type="distribution">
<paragraph id="6AD919023CF7E88FB3827DA2247DFB37" pageId="0" pageNumber="85">Distribution.</paragraph>
<paragraph id="F318C55A7F73E8349DDE3B1CD480CF75" pageId="0" pageNumber="85">
(Fig.
<figureCitation id="6EB2F9315FFD53A2B9295535D5D39594" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distributions of Phonarellus and Gymnogryllus in China (Note: the locality of P. zebripes cited from Liu and He (2022), and the remaining location information is based on the data of examined materials)." figureDoi="10.3897/zookeys.1129.87706.figure2" httpUri="https://binary.pensoft.net/fig/770017" pageId="0" pageNumber="85">2</figureCitation>
). China (Yunnan, Guangxi, Guangdong).
</paragraph>
</subSubSection>
<subSubSection id="312DA9B5E326D33FC39F8AF71FF8F625" pageId="0" pageNumber="85" type="description">
<paragraph id="DC7F5BF3B8AA68365F52EDF33187EB90" pageId="0" pageNumber="85">Measurements (mm).</paragraph>
<paragraph id="A0BBD2C8CFAA95A74BF72D838CBC19B0" pageId="0" pageNumber="85">
<emphasis id="6F6E9A29FA8C2C9A07EBA0552A956EDF" bold="true" pageId="0" pageNumber="85">
Male (
<emphasis id="95999949E89022395013EE0950033B23" italics="true" pageId="0" pageNumber="85">N</emphasis>
= 30)
</emphasis>
: BL 27.73-29.42; HW 6.42-6.84; PL 4.86-5.32; FWL 18.62-21.32; HLL 16.27-17.12; HTL 9.43-10.14; EW 1.56-1.79; PW 7.62-7.98; HWL 6.04-6.84; DVL 4.62-4.96; ML 3.72-3.98; CL 8.15-8.54; FTL 5.42-5.76; MTL 6.29-6.45.
</paragraph>
</subSubSection>
<subSubSection id="E279A792C04693F59D45F0F14A292A2D" pageId="0" pageNumber="85" type="description">
<paragraph id="5C82FA911369218D0D5EA61D979F9554" pageId="0" pageNumber="85">Description.</paragraph>
<paragraph id="8937252CB9D0112D1D35C847EE2F72D1" pageId="0" pageNumber="85">
<emphasis id="77CA4AD35E76460C3816005C5D99BFD4" bold="true" italics="true" pageId="0" pageNumber="85">Male</emphasis>
(Figs
<figureCitation id="AF8427BF29E7F0FC649E9DC6939EAF22" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Living photos of some Phonarellus and Gymnogryllus species A, B male of P. minor (with varing proportion of black areas on hind legs) C female of P. minor D male of P. ritsemae E male of G. odonopetalus F female of G. odonopetalus (Photos A-D were provided by Zhang, Tao, and E, F were photographed by He, Zhixin)." figureDoi="10.3897/zookeys.1129.87706.figure1" httpUri="https://binary.pensoft.net/fig/770016" pageId="0" pageNumber="85">1E</figureCitation>
,
<figureCitation id="94121DE301FE9175CADA0F38E0A4973C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bodies of G. odonopetalus A male B female. Scale bar: 10 mm." figureDoi="10.3897/zookeys.1129.87706.figure3" httpUri="https://binary.pensoft.net/fig/770018" pageId="0" pageNumber="85">3A</figureCitation>
). Head longer than wide, as wide as fore margin of pronotum. Vertex smooth and broad. Occiput slightly inclined. Frontal rostrum as wide as one eye and flattened. Scape of antennae flat, shield-like, and as wide as half of frontal rostrum. With three small ocelli, arranged in a straight line, median ocellus semilunar. Eyes about 1/4 length of head. Epistomal suture straight and close to eyes. Labrum slightly convex and rhombus shaped. Last segment of maxillary palpi slightly narrower than the third. Labial palpi the middle segment longer than the others.
</paragraph>
<caption id="557DD762A2A9184EF2AEFD4C8644E2DC" doi="10.3897/zookeys.1129.87706.figure3" httpUri="https://binary.pensoft.net/fig/770018" pageId="0" pageNumber="85" start="Figure 3" startId="F3">
<paragraph id="BA208C7860AB71909C788EC6433F6B24" pageId="0" pageNumber="85">
<emphasis id="82782806CDDE66FC740050D2800494E7" bold="true" pageId="0" pageNumber="85">Figure 3.</emphasis>
Bodies of
<taxonomicName id="50515713DBBF0E8C35CCC1B7485A1499" lsidName="G. odonopetalus" pageId="0" pageNumber="85" rank="species" species="odonopetalus">
<emphasis id="E29542E09944F2F94A00C6940A25FD3C" italics="true" pageId="0" pageNumber="85">G. odonopetalus</emphasis>
</taxonomicName>
<emphasis id="67D0EE4158C2271995B3D7C441B29CCB" bold="true" pageId="0" pageNumber="85">A</emphasis>
male
<emphasis id="788DE11AA33D506AB62F3C36010A501D" bold="true" pageId="0" pageNumber="85">B</emphasis>
female. Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="424FD6EB607D5D18849F604C663688DB" pageId="0" pageNumber="85">Pronotum disc rectanglur, anterior margin concave, posterior margin sinuated; a longitudinal groove in middle of pronotum, about 2/3 the length of the pronotum. Fore margin of pronotum rough and densely pubescent. Tegmen reaching tip of abdomen; with three oblique veins, outmost nearly vertical and straight, two internal inclined and longer than outmost; and them converging diagonal vein. Diagonal vein curved and anteriorly forked. Chord veins three, the internal two extremely curved, connected at bottom. Between diagonal vein and the most internal chord vein armed with two transverse veins. Mirror large, inclined rectangular. Apical field triangular, about 2/5 the length of tegmina, variable among individuals, armed with rectangular cells.</paragraph>
<paragraph id="2618653D8E0896D3B3B273B3806C45C2" pageId="0" pageNumber="85">Fore tibiae with inner and outer tympanum, inner tympanum small and ovoid, outer longer-oval. Hind femora brown with light stripes. Distal of hind tibiae with five dorsal spurs on both sides; apical spurs six, the inner apical spurs three (the dorsal one longest, the ventral one shortest and 1/4 length of the longest one, the middle one about 2/3 length of the longest), and the outer apical spurs three (equal length of the dorsal one and the ventral one, about 2/3 length of the middle one). Subgenital plate hook-like. Cercus straight and short; with long hair sparse and short hair dense.</paragraph>
<paragraph id="1A356F4DE286E285D267D5F508B1214E" pageId="0" pageNumber="85">
<emphasis id="D1AACA7F2CE207E3D813E3E243068AD8" bold="true" italics="true" pageId="0" pageNumber="85">Genitalia</emphasis>
(Figs
<figureCitation id="473CD14933C4454CBDDC890A204DF49A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Intraspecific variation of genitalia in G. odonopetalus A-F genitalia from six individuals in lateral view (Note: the red arrows point to the posterior of epiphallus, highlighting variations among individuals; the blue circles indicate the protuberance of mid-ectoparamere) G-L genitalia of the same six individuals in dorsal view (Note: the green circles indicate the median lobe of the epiphallus posterior)." figureDoi="10.3897/zookeys.1129.87706.figure4" httpUri="https://binary.pensoft.net/fig/770019" pageId="0" pageNumber="85">4</figureCitation>
,
<figureCitation id="1385DD19DC8455AC428618D73B87772D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Intraspecific variation of genitalia in G. odonopetalus A-C and G-I photos of genitalia from six individuals in caudal view D-F and J-L drawings of genitalia from the same six individuals in caudal view." figureDoi="10.3897/zookeys.1129.87706.figure5" httpUri="https://binary.pensoft.net/fig/770020" pageId="0" pageNumber="85">5</figureCitation>
). Male genitalia robust, epiphallus arch-shaped in front view; inner side of medial lobe armed with a pair of teeth. Median notch between paired apical teeth and length of teeth varyable between individuals. End of middle lobes of epiphallus bent upward, for less than 90°, varyable between individuals. Notch of epiphallus anterior variable, with bottom angular or broad and arc-shaped. Outer edge of end of ectoparamere armed with irregular numbers of teeth.
</paragraph>
<caption id="3446B051F31A09264C5B9B9BBEF7781B" doi="10.3897/zookeys.1129.87706.figure4" httpUri="https://binary.pensoft.net/fig/770019" pageId="0" pageNumber="85" start="Figure 4" startId="F4">
<paragraph id="CE4EE19270D937B89A0CD1FA8F70F8FF" pageId="0" pageNumber="85">
<emphasis id="0F94186BDA82CCFF4B6CA6CE3C0A3296" bold="true" pageId="0" pageNumber="85">Figure 4.</emphasis>
Intraspecific variation of genitalia in
<taxonomicName id="051B00E36EF5EA5F2795ED5A71B3D4A7" lsidName="G. odonopetalus" pageId="0" pageNumber="85" rank="species" species="odonopetalus">
<emphasis id="EDB8E22A3091063247F7606017338A64" italics="true" pageId="0" pageNumber="85">G. odonopetalus</emphasis>
</taxonomicName>
<emphasis id="DCE90116D15D495CF2AD818E13EAB60F" bold="true" pageId="0" pageNumber="85">A-F</emphasis>
genitalia from six individuals in lateral view (Note: the red arrows point to the posterior of epiphallus, highlighting variations among individuals; the blue circles indicate the protuberance of mid-ectoparamere)
<emphasis id="DDD58625D8E9C8B7EEDDAD7E53606D92" bold="true" pageId="0" pageNumber="85">G-L</emphasis>
genitalia of the same six individuals in dorsal view (Note: the green circles indicate the median lobe of the epiphallus posterior).
</paragraph>
</caption>
<caption id="737192B9E88DF94BD068020A41CFB378" doi="10.3897/zookeys.1129.87706.figure5" httpUri="https://binary.pensoft.net/fig/770020" pageId="0" pageNumber="85" start="Figure 5" startId="F5">
<paragraph id="C0E29DA3301790DFFCC74D2EC051B722" pageId="0" pageNumber="85">
<emphasis id="063ED229846CDF879C2C9C2914AB6506" bold="true" pageId="0" pageNumber="85">Figure 5.</emphasis>
Intraspecific variation of genitalia in
<taxonomicName id="B4B162AE22128AA5DC558E138DE96D8E" lsidName="G. odonopetalus" pageId="0" pageNumber="85" rank="species" species="odonopetalus">
<emphasis id="1B91E20E6184F1BB6392868545018A0A" italics="true" pageId="0" pageNumber="85">G. odonopetalus</emphasis>
</taxonomicName>
<emphasis id="C3D9C13B30EB9331EEA77E313BF898AA" bold="true" pageId="0" pageNumber="85">A-C</emphasis>
and
<emphasis id="C5109D1868888480D3A2C78B4820E950" bold="true" pageId="0" pageNumber="85">G-I</emphasis>
photos of genitalia from six individuals in caudal view
<emphasis id="97C8EC80E34FB59EA76D7D6087A2E81A" bold="true" pageId="0" pageNumber="85">D-F</emphasis>
and
<emphasis id="059F9939346DA96FC1B3B44334FA6125" bold="true" pageId="0" pageNumber="85">J-L</emphasis>
drawings of genitalia from the same six individuals in caudal view.
</paragraph>
</caption>
<paragraph id="E2DB4E10FDD2C35E8B2269EC45573337" pageId="0" pageNumber="85">
<emphasis id="7D51AE213387062710EE2595220D9794" bold="true" italics="true" pageId="0" pageNumber="85">Coloration</emphasis>
(Fig.
<figureCitation id="3F6BF4FE3A5ABA568E7CC91582FB25AB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bodies of G. odonopetalus A male B female. Scale bar: 10 mm." figureDoi="10.3897/zookeys.1129.87706.figure3" httpUri="https://binary.pensoft.net/fig/770018" pageId="0" pageNumber="85">3</figureCitation>
). Head, pronotum and much of the hind femur blackish brown. Occiput uniformly blackish brown. 1/3 part of cheek bottom brown. Frons black, ventral quarter of lateral lobe with a light stripe. Hind femur black.
</paragraph>
<paragraph id="AC910B9C4BB41BBA8FE455C92DBB726D" pageId="0" pageNumber="85">
<emphasis id="E2674553483D1E57AACEEA05758F811D" bold="true" pageId="0" pageNumber="85">Female</emphasis>
(Figs
<figureCitation id="B984218C73135C3AFE04912425A4F211" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Living photos of some Phonarellus and Gymnogryllus species A, B male of P. minor (with varing proportion of black areas on hind legs) C female of P. minor D male of P. ritsemae E male of G. odonopetalus F female of G. odonopetalus (Photos A-D were provided by Zhang, Tao, and E, F were photographed by He, Zhixin)." figureDoi="10.3897/zookeys.1129.87706.figure1" httpUri="https://binary.pensoft.net/fig/770016" pageId="0" pageNumber="85">1F</figureCitation>
,
<figureCitation id="D3AF18BF1B76F6778DA5307FE423F459" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bodies of G. odonopetalus A male B female. Scale bar: 10 mm." figureDoi="10.3897/zookeys.1129.87706.figure3" httpUri="https://binary.pensoft.net/fig/770018" pageId="0" pageNumber="85">3B</figureCitation>
). Slightly smaller than male, with tegmina dark brown. Ovipositor as long as pronotum.
</paragraph>
</subSubSection>
<subSubSection id="7AEBC97C4E13949E267494821E2792D4" pageId="0" pageNumber="85" type="remarks">
<paragraph id="4C4B2EA7517342C36639BABF616AF753" pageId="0" pageNumber="85">Remarks.</paragraph>
<paragraph id="890A1A11A71EC31BC04FE43FBAE155D3" pageId="0" pageNumber="85">
<bibRefCitation id="0B81B0C64013D44D6CA86197BA80A223" DOI="https://doi.org/10.11646/zootaxa.2733.1.3" author="Ma, LB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="85" pagination="31 - 40" refId="B20" refString="Ma, LB, Zhang, YL, 2011. The cricket genus Gymnogryllus (Grylloidea: Gryllidae: Gryllinae: Gryllini) from China with description of six new species. Zootaxa 2733 (1): 31 - 40, DOI: https://doi.org/10.11646/zootaxa.2733.1.3" title="The cricket genus Gymnogryllus (Grylloidea: Gryllidae: Gryllinae: Gryllini) from China with description of six new species." url="https://doi.org/10.11646/zootaxa.2733.1.3" volume="2733" year="2011">Ma and Zhang (2011)</bibRefCitation>
proposed four species,
<taxonomicName id="3C6BD9A2F5051108498C38FE7C5D325A" lsidName="G. yunnanensis" pageId="0" pageNumber="85" rank="species" species="yunnanensis">
<emphasis id="28FA30B7F0AB21F6EEC8F881ADD96A90" italics="true" pageId="0" pageNumber="85">G. yunnanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="B110197A2E5C709E7C9D61931D3E601F" lsidName="G. striatus" pageId="0" pageNumber="85" rank="species" species="striatus">
<emphasis id="4B1869A784B390F9500FB5DEE92A3036" italics="true" pageId="0" pageNumber="85">G. striatus</emphasis>
</taxonomicName>
,
<taxonomicName id="75CA9E3A6DEB95CCF5BB3626B817A46F" lsidName="G. longus" pageId="0" pageNumber="85" rank="species" species="longus">
<emphasis id="C4135AD7931026D3F67D6D5ECD67129A" italics="true" pageId="0" pageNumber="85">G. longus</emphasis>
</taxonomicName>
, and
<taxonomicName id="F0DAA6DFFDA4659473C12F7ED03A8A04" lsidName="G. tumidulus" pageId="0" pageNumber="85" rank="species" species="tumidulus">
<emphasis id="375900E85394805CFE11E07CB9A621A4" italics="true" pageId="0" pageNumber="85">G. tumidulus</emphasis>
</taxonomicName>
, based on the angle between the apical epiphallic teeth, the posterior edge of the medial lobe of epiphallus, and the apical field of the forewing. We examined our specimens and determined that these characters present intraspecific variation. In particular, the angle between the apical teeth of epiphallus shows tremendous variation between specimens (Fig.
<figureCitation id="D81F5F6F493632CA4A63CE6754A60821" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Intraspecific variation of genitalia in G. odonopetalus A-F genitalia from six individuals in lateral view (Note: the red arrows point to the posterior of epiphallus, highlighting variations among individuals; the blue circles indicate the protuberance of mid-ectoparamere) G-L genitalia of the same six individuals in dorsal view (Note: the green circles indicate the median lobe of the epiphallus posterior)." figureDoi="10.3897/zookeys.1129.87706.figure4" httpUri="https://binary.pensoft.net/fig/770019" pageId="0" pageNumber="85">4A-F</figureCitation>
). Besides, the epiphalli of these four species are armed with two pairs of teeth inside, which is consistent with the holotype of
<taxonomicName id="B4E95F41A0C66D06FC98BC7A5FA74315" lsidName="G. odonopetalus" pageId="0" pageNumber="85" rank="species" species="odonopetalus">
<emphasis id="ED9C6C7E2A77BAAE4519F8BBFF5F649D" italics="true" pageId="0" pageNumber="85">G. odonopetalus</emphasis>
</taxonomicName>
deposited in SNNU (in fact, we found that one pair of teeth is formed by the ectoparamere curved at the tip on the type specimen of
<taxonomicName id="98EA0BC56A85A7238B1DE4489F6A4943" lsidName="G. odonopetalus" pageId="0" pageNumber="85" rank="species" species="odonopetalus">
<emphasis id="E3F1C664B2CA0B735BCCF39E91BE2408" italics="true" pageId="0" pageNumber="85">G. odonopetalus</emphasis>
</taxonomicName>
). Therefore, we regard
<taxonomicName id="CDAD790C5C295555DC3ED49E95E70E65" lsidName="G. yunnanensis" pageId="0" pageNumber="85" rank="species" species="yunnanensis">
<emphasis id="3CA88738F9F0F0D12E8505C83726183C" italics="true" pageId="0" pageNumber="85">G. yunnanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="82BF77ED23250900522CCC020EEA1F30" lsidName="G. striatus" pageId="0" pageNumber="85" rank="species" species="striatus">
<emphasis id="12E7132180AA0E3DE6A32EFF491B2667" italics="true" pageId="0" pageNumber="85">G. striatus</emphasis>
</taxonomicName>
,
<taxonomicName id="6291098D48F4BB63769CCC95B816E9CF" lsidName="G. longus" pageId="0" pageNumber="85" rank="species" species="longus">
<emphasis id="F29BAEE02299DAA85B3C686A25EEEF00" italics="true" pageId="0" pageNumber="85">G. longus</emphasis>
</taxonomicName>
, and
<taxonomicName id="112509C4CAD983D17A124A9B3A496533" lsidName="G. tumidulus" pageId="0" pageNumber="85" rank="species" species="tumidulus">
<emphasis id="13DB07F998BFCA6A592B4CDA20D564D8" italics="true" pageId="0" pageNumber="85">G. tumidulus</emphasis>
</taxonomicName>
as junior synonyms of
<taxonomicName id="47B4B27C16914EDC202943F607EDC968" lsidName="G. odonopetalus" pageId="0" pageNumber="85" rank="species" species="odonopetalus">
<emphasis id="935375D4C0F35A3CB81A0A08870B552F" italics="true" pageId="0" pageNumber="85">G. odonopetalus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="27B373DB2A5F7B25BC881AD739E160C6" pageId="0" pageNumber="85">
Hence, the species
<taxonomicName id="21D5BA53AA37F0E465CB67DB490C97B4" lsidName="G. odonopetalus" pageId="0" pageNumber="85" rank="species" species="odonopetalus">
<emphasis id="0D5C7816EC16168A98E140AD6056D8E6" italics="true" pageId="0" pageNumber="85">G. odonopetalus</emphasis>
</taxonomicName>
has some charecters showing intraspecific variation even within specimens collected from the same place and time. In lateral view, the angle between the apical teeth and the posterior edge of the medial lobe of epiphallus is variable among individuals (red arrows in Fig.
<figureCitation id="8CC7BEC7A4511B87C6E5BC6E081C3AE5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Intraspecific variation of genitalia in G. odonopetalus A-F genitalia from six individuals in lateral view (Note: the red arrows point to the posterior of epiphallus, highlighting variations among individuals; the blue circles indicate the protuberance of mid-ectoparamere) G-L genitalia of the same six individuals in dorsal view (Note: the green circles indicate the median lobe of the epiphallus posterior)." figureDoi="10.3897/zookeys.1129.87706.figure4" httpUri="https://binary.pensoft.net/fig/770019" pageId="0" pageNumber="85">4</figureCitation>
); the protuberances in the middle of the ectoparamere vary in size (blue circles indicate the protuberance of mid-ectoparamere in Fig.
<figureCitation id="F4AEC60C4527FC63829A6C31E7C1CA6F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Intraspecific variation of genitalia in G. odonopetalus A-F genitalia from six individuals in lateral view (Note: the red arrows point to the posterior of epiphallus, highlighting variations among individuals; the blue circles indicate the protuberance of mid-ectoparamere) G-L genitalia of the same six individuals in dorsal view (Note: the green circles indicate the median lobe of the epiphallus posterior)." figureDoi="10.3897/zookeys.1129.87706.figure4" httpUri="https://binary.pensoft.net/fig/770019" pageId="0" pageNumber="85">4</figureCitation>
). In dorsal view, the end of the epiphallus is variable (shown by green circles in Fig.
<figureCitation id="6F042795E70FD9286506122D0565F240" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Intraspecific variation of genitalia in G. odonopetalus A-F genitalia from six individuals in lateral view (Note: the red arrows point to the posterior of epiphallus, highlighting variations among individuals; the blue circles indicate the protuberance of mid-ectoparamere) G-L genitalia of the same six individuals in dorsal view (Note: the green circles indicate the median lobe of the epiphallus posterior)." figureDoi="10.3897/zookeys.1129.87706.figure4" httpUri="https://binary.pensoft.net/fig/770019" pageId="0" pageNumber="85">4G-L</figureCitation>
); the anterior notch of epiphallus is morphologically also diverse. In caudal view, shape and space between apical teeth of epiphallus are variable (drawing of epiphallus in Fig.
<figureCitation id="17662DFCA1FFA88D4F5F814D9EE45B9A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Intraspecific variation of genitalia in G. odonopetalus A-C and G-I photos of genitalia from six individuals in caudal view D-F and J-L drawings of genitalia from the same six individuals in caudal view." figureDoi="10.3897/zookeys.1129.87706.figure5" httpUri="https://binary.pensoft.net/fig/770020" pageId="0" pageNumber="85">5D-F, J-L</figureCitation>
); the length of the apical area of the ectoparamere varies among individuals.
</paragraph>
</subSubSection>
</treatment>
</document>
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