treatments-xml/data/9B/47/4B/9B474B53FFEB3D69FF01B1FBFC455569.xml
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<document id="511A31E719B856F4341A1195E6F6C952" ID-CLB-Dataset="29524" ID-DOI="10.11646/zootaxa.4446.1.4" ID-GBIF-Dataset="d35cedde-4442-4a14-be2d-d5213a12ab29" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437384" ID-ZooBank="E5ABBD4C-C643-4091-9AD4-D9D4028A218A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538121156330" checkinUser="plazi" docAuthor="Ban, Teruaki" docDate="2018" docId="9B474B53FFEB3D69FF01B1FBFC455569" docLanguage="en" docName="zootaxa.4446.1.4.pdf" docOrigin="Zootaxa 4446 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sadoletus laosensis Ban 2018, sp. nov." docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="677E332BFFEE3D6FFF96B264FFD05011" masterDocTitle="The lygaeoid bug family Heterogastridae (Hemiptera: Heteroptera: Lygaeoidea) from Laos and Thailand, with descriptions of three new species" masterLastPageNumber="67" masterPageNumber="51" pageNumber="56" updateTime="1698555679930" updateUser="ExternalLinkService">
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<mods:title id="651B93F5551C7EAB30742447B6FB876D">The lygaeoid bug family Heterogastridae (Hemiptera: Heteroptera: Lygaeoidea) from Laos and Thailand, with descriptions of three new species</mods:title>
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<mods:namePart id="2AA91C2DE112E8045F821F296653AF2D">Ban, Teruaki</mods:namePart>
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<mods:date id="0ACCE813BB6F0AE640B54168C14CB6A0">2018</mods:date>
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<mods:number id="D3F140DC5B1EF8843365D1E3DB7255C3">2018-07-16</mods:number>
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<treatment id="9B474B53FFEB3D69FF01B1FBFC455569" ID-DOI="http://doi.org/10.5281/zenodo.5974608" ID-GBIF-Taxon="148403378" ID-Zenodo-Dep="5974608" LSID="urn:lsid:plazi:treatment:9B474B53FFEB3D69FF01B1FBFC455569" httpUri="http://treatment.plazi.org/id/9B474B53FFEB3D69FF01B1FBFC455569" lastPageId="6" lastPageNumber="57" pageId="5" pageNumber="56">
<subSubSection id="5BF4A9CEFFEB3D6AFF01B1FBFE6A53CA" pageId="5" pageNumber="56" type="nomenclature">
<paragraph id="1351FA45FFEB3D6AFF01B1FBFE3653AB" blockId="5.[151,486,927,987]" box="[151,486,927,954]" pageId="5" pageNumber="56">
<heading id="48194D29FFEB3D6AFF01B1FBFE3653AB" bold="true" box="[151,486,927,954]" fontSize="11" level="1" pageId="5" pageNumber="56" reason="1">
<emphasis id="219A2657FFEB3D6AFF01B1FBFE3653AB" bold="true" box="[151,486,927,954]" pageId="5" pageNumber="56">
<taxonomicName id="D4EE81C6FFEB3D6AFF01B1FBFE5053A8" ID-CoL="8QM2N" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[151,384,927,953]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="219A2657FFEB3D6AFF01B1FBFE5053A8" bold="true" box="[151,384,927,953]" italics="true" pageId="5" pageNumber="56">Sadoletus laosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFEB3D6AFE1EB1C4FE3653AB" box="[392,486,928,954]" pageId="5" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1351FA45FFEB3D6AFF01B1A6FE6A53CA" blockId="5.[151,486,927,987]" box="[151,442,962,987]" pageId="5" pageNumber="56">
(
<figureCitation id="8BD5E6C0FFEB3D6AFF09B1A6FEDB53CA" box="[159,267,962,987]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Figs. 67</figureCitation>
,
<figureCitation id="8BD5E6C0FFEB3D6AFE80B1A7FEE453CA" box="[278,308,963,987]" captionStart="FIGURES 1416" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 1416. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="5" pageNumber="56">14</figureCitation>
,
<figureCitation id="8BD5E6C0FFEB3D6AFED6B1A7FE8E53CA" box="[320,350,963,987]" captionStart="FIGURES 1719" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 1719. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="5" pageNumber="56">17</figureCitation>
,
<figureCitation id="8BD5E6C0FFEB3D6AFEFCB1A7FE6253CA" box="[362,434,963,987]" captionStart="FIGURES 2024" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[173,1319,245,1801]" captionTargetId="figure@12.[151,1436,215,1822]" captionTargetPageId="12" captionText="FIGURES 2024. Sadoletus laosensis sp. nov. 20, Pygophore, dorsal view (setae omitted from left half); 21, left paramere, dorsal view; 2223, phallus (endosoma not everted), ventral (22) and lateral (23) views; 24, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437396/files/figure.png" pageId="5" pageNumber="56">2024</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFEB3D6AFF01B66EFAC5547A" pageId="5" pageNumber="56" type="materials_examined">
<paragraph id="1351FA45FFEB3D6AFF01B66EFAC5547A" blockId="5.[151,1437,1034,2031]" pageId="5" pageNumber="56">
<materialsCitation id="A386F018FFEB3D6AFF01B66EFCA35456" ID-GBIF-Occurrence="1913858568" collectingDate="2009-10-12" collectorName="Teruaki Ban" country="Laos" county="Lak" elevation="837" latitude="20.616667" location="Oudomxay Prov." longLatPrecision="1267" longitude="102.0" municipality="Waterfall" pageId="5" pageNumber="56" specimenCount="1" specimenCount-female="1" stateProvince="Oudomxay" typeStatus="holotype">
<emphasis id="219A2657FFEB3D6AFF01B66EFE855432" bold="true" box="[151,341,1034,1059]" pageId="5" pageNumber="56">
<typeStatus id="CC5544E7FFEB3D6AFF01B66EFF025432" box="[151,210,1034,1059]" pageId="5" pageNumber="56">Type</typeStatus>
Materials.
</emphasis>
<typeStatus id="CC5544E7FFEB3D6AFECBB66EFE1A5432" box="[349,458,1034,1059]" pageId="5" pageNumber="56" type="holotype">Holotype</typeStatus>
:
<specimenCount id="05E831CCFFEB3D6AFE40B66FFE3B5433" box="[470,491,1035,1058]" pageId="5" pageNumber="56" type="female"></specimenCount>
,
<collectingCountry id="6BF9BAD5FFEB3D6AFE6DB66EFD975432" box="[507,583,1034,1059]" name="Laos" pageId="5" pageNumber="56">
<emphasis id="219A2657FFEB3D6AFE6DB66EFD975432" bold="true" box="[507,583,1034,1059]" pageId="5" pageNumber="56">LAOS</emphasis>
</collectingCountry>
:
<geoCoordinate id="76DA9C82FFEB3D6AFDCEB66FFD675432" box="[600,695,1035,1059]" direction="north" orientation="latitude" pageId="5" pageNumber="56" precision="925" value="20.616667">20°37'N</geoCoordinate>
,
<geoCoordinate id="76DA9C82FFEB3D6AFD52B66FFCFE5432" box="[708,814,1035,1059]" direction="east" orientation="longitude" pageId="5" pageNumber="56" precision="925" value="102.0">102°00'E</geoCoordinate>
, alt.
<elevation id="98C31D76FFEB3D6AFCF1B66FFC625433" box="[871,946,1035,1059]" metricMagnitude="2" metricUnit="m" metricValue="8.37" pageId="5" pageNumber="56" unit="m" value="837.0">
<quantity id="D41657A0FFEB3D6AFCF1B66FFC625433" box="[871,946,1035,1059]" metricMagnitude="2" metricUnit="m" metricValue="8.37" pageId="5" pageNumber="56" unit="m" value="837.0">837 m</quantity>
</elevation>
,
<collectingCounty id="FA3082C9FFEB3D6AFC56B66EFC3C5432" box="[960,1004,1034,1059]" pageId="5" pageNumber="56">Lak</collectingCounty>
11
<collectingMunicipality id="F335603FFFEB3D6AFB8EB66EFB525432" box="[1048,1154,1034,1059]" pageId="5" pageNumber="56">Waterfall</collectingMunicipality>
,
<quantity id="D41657A0FFEB3D6AFB19B66FFB075432" box="[1167,1239,1034,1059]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="56" unit="km" value="11.0">11 km</quantity>
South-east From
<collectingRegion id="D12A34A7FFEB3D6AFF01B64AFEC55456" box="[151,277,1070,1095]" country="Laos" name="Oudomxai" pageId="5" pageNumber="56">Oudomxay</collectingRegion>
,
<location id="1631AC9EFFEB3D6AFEB3B64AFE3C5457" LSID="urn:lsid:plazi:treatment:9B474B53FFEB3D69FF01B1FBFC455569:1631AC9EFFEB3D6AFEB3B64AFE3C5457" box="[293,492,1070,1095]" country="Laos" county="Lak" latitude="20.616667" longLatPrecision="1267" longitude="102.0" municipality="Waterfall" name="Oudomxay Prov." pageId="5" pageNumber="56" stateProvince="Oudomxay">Oudomxay Prov.</location>
,
<collectingDate id="7714256DFFEB3D6AFE6BB64BFDA65456" box="[509,630,1071,1095]" pageId="5" pageNumber="56" value="2009-10-12">
<date id="6750DC85FFEB3D6AFE6BB64BFDA65456" box="[509,630,1071,1095]" pageId="5" pageNumber="56" value="2009-10-12">12.X.2009</date>
</collectingDate>
,
<collectorName id="BE1B9F93FFEB3D6AFD11B64AFCC65457" box="[647,790,1070,1095]" pageId="5" pageNumber="56">Teruaki Ban</collectorName>
(TUA)
</materialsCitation>
.
<materialsCitation id="A386F018FFEB3D6AFC12B64BFC6D547A" ID-GBIF-Occurrence="1913858562" collectingDate="2009-10-04" collectorName="Genki Oishi &amp; Same" country="Laos" county="Lak" elevation="837" latitude="20.616667" location="Same" longLatPrecision="1267" longitude="102.0" municipality="Waterfall" pageId="5" pageNumber="56" specimenCount="1" specimenCount-female="1" stateProvince="Oudomxay" typeStatus="paratype">
<typeStatus id="CC5544E7FFEB3D6AFC12B64BFC255456" box="[900,1013,1071,1095]" pageId="5" pageNumber="56" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="6BF9BAD5FFEB3D6AFB93B64AFB815456" box="[1029,1105,1070,1095]" name="Laos" pageId="5" pageNumber="56">
<emphasis id="219A2657FFEB3D6AFB93B64AFB815456" bold="true" box="[1029,1105,1070,1095]" pageId="5" pageNumber="56">LAOS</emphasis>
</collectingCountry>
:
<specimenCount id="05E831CCFFEB3D6AFBF0B64BFB475457" box="[1126,1175,1071,1095]" pageId="5" pageNumber="56" type="female">1 ♀</specimenCount>
,
<location id="1631AC9EFFEB3D6AFB3DB64BFB395456" LSID="urn:lsid:plazi:treatment:9B474B53FFEB3D69FF01B1FBFC455569:1631AC9EFFEB3D6AFB3DB64BFB395456" box="[1195,1257,1071,1095]" country="Laos" name="Same" pageId="5" pageNumber="56">Same</location>
locality as for holotype,
<collectingDate id="7714256DFFEB3D6AFE91B637FEA1547A" box="[263,369,1107,1131]" pageId="5" pageNumber="56" value="2009-10-04">
<date id="6750DC85FFEB3D6AFE91B637FEA1547A" box="[263,369,1107,1131]" pageId="5" pageNumber="56" value="2009-10-04">4.X.2009</date>
</collectingDate>
,
<collectorName id="BE1B9F93FFEB3D6AFEEBB636FDD5547A" box="[381,517,1106,1131]" pageId="5" pageNumber="56">Genki Ôishi</collectorName>
leg. (TUA);
<collectorName id="BE1B9F93FFEB3D6AFD01B637FD05547A" box="[663,725,1107,1131]" pageId="5" pageNumber="56">Same</collectorName>
date as for
<typeStatus id="CC5544E7FFEB3D6AFCCEB636FC6D547A" box="[856,957,1106,1131]" pageId="5" pageNumber="56" type="holotype">holotype</typeStatus>
</materialsCitation>
, 3 ♂ 3 ♀, Teruaki Ban (TUA).
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFEB3D6AFF51B612FD6D5576" pageId="5" pageNumber="56" type="diagnosis">
<paragraph id="1351FA45FFEB3D6AFF51B612FD6D5576" blockId="5.[151,1437,1034,2031]" pageId="5" pageNumber="56">
<emphasis id="219A2657FFEB3D6AFF51B612FE91549E" bold="true" box="[199,321,1142,1167]" pageId="5" pageNumber="56">Diagnosis.</emphasis>
Recognized among congeners of
<taxonomicName id="D4EE81C6FFEB3D6AFD42B612FC91549F" authorityName="Distant" authorityYear="1903" box="[724,833,1142,1166]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="219A2657FFEB3D6AFD42B612FC91549F" box="[724,833,1142,1166]" italics="true" pageId="5" pageNumber="56">Sadoletus</emphasis>
</taxonomicName>
by the following characters: head, anterior lobe of pronotum and basal half of scutellum black (
<figureCitation id="8BD5E6C0FFEB3D6AFD18B6FEFD2A54A2" box="[654,762,1178,1203]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Figs. 6, 7</figureCitation>
); labium exceeding posterior margin of prothorax (
<figureCitation id="8BD5E6C0FFEB3D6AFAACB6FEFA5E54A2" box="[1338,1422,1178,1203]" captionStart="FIGURES 1416" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 1416. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="5" pageNumber="56">Fig. 14</figureCitation>
); posterior lobe of pronotum pale yellow, with 2 pale brownish longitudinal markings (
<figureCitation id="8BD5E6C0FFEB3D6AFBF4B6DAFB0154C6" box="[1122,1233,1214,1239]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Figs. 6, 7</figureCitation>
); corium white in basal two-thirds and dark brown in apical third, with pale brown markings around inner angle; clavus white (
<figureCitation id="8BD5E6C0FFEB3D6AFAF2B686FF14550E" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Figs. 6, 7</figureCitation>
); femora and tibiae generally pale yellow, profemur with 2 spines; abdominal sterna reddish brown; posterior margin of sternite VI weakly sinuate in female (
<figureCitation id="8BD5E6C0FFEB3D6AFD56B74EFCC55552" box="[704,789,1322,1347]" captionStart="FIGURES 1719" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 1719. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="5" pageNumber="56">Fig. 17</figureCitation>
); posterior margin of pygophore and cuplike sclerite not fused (
<figureCitation id="8BD5E6C0FFEB3D6AFF75B72AFEE75576" box="[227,311,1358,1383]" captionStart="FIGURES 2024" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[173,1319,245,1801]" captionTargetId="figure@12.[151,1436,215,1822]" captionTargetPageId="12" captionText="FIGURES 2024. Sadoletus laosensis sp. nov. 20, Pygophore, dorsal view (setae omitted from left half); 21, left paramere, dorsal view; 2223, phallus (endosoma not everted), ventral (22) and lateral (23) views; 24, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437396/files/figure.png" pageId="5" pageNumber="56">Fig. 20</figureCitation>
); paramere acute at apex (
<figureCitation id="8BD5E6C0FFEB3D6AFDCAB72AFD7F5576" box="[604,687,1358,1383]" captionStart="FIGURES 2024" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[173,1319,245,1801]" captionTargetId="figure@12.[151,1436,215,1822]" captionTargetPageId="12" captionText="FIGURES 2024. Sadoletus laosensis sp. nov. 20, Pygophore, dorsal view (setae omitted from left half); 21, left paramere, dorsal view; 2223, phallus (endosoma not everted), ventral (22) and lateral (23) views; 24, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437396/files/figure.png" pageId="5" pageNumber="56">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFEB3D69FF51B716FA835424" lastPageId="6" lastPageNumber="57" pageId="5" pageNumber="56" type="description">
<paragraph id="1351FA45FFEB3D6AFF51B716FAB75706" blockId="5.[151,1437,1034,2031]" pageId="5" pageNumber="56">
<emphasis id="219A2657FFEB3D6AFF51B716FE89559A" bold="true" box="[199,345,1394,1419]" pageId="5" pageNumber="56">Description.</emphasis>
Female (
<typeStatus id="CC5544E7FFEB3D6AFE53B716FDFE559A" box="[453,558,1394,1419]" pageId="5" pageNumber="56" type="holotype">holotype</typeStatus>
): Short decumbent setae on body silvery. Punctures on pronotum, scutellum, and corium brown. Head (
<figureCitation id="8BD5E6C0FFEB3D6AFE5AB7F2FDC355BE" box="[460,531,1430,1455]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Fig. 7</figureCitation>
) black. Clypeus yellowish brown. Antenna generally brownish yellow, apical three-quarters of segment IV tinged with brown. Labium brownish yellow; basal three-quarters of segment I and apex of segment IV brown. Pronotum (
<figureCitation id="8BD5E6C0FFEB3D6AFDD8B7BAFD4155E6" box="[590,657,1502,1527]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Fig. 7</figureCitation>
) black in anterior lobe, anterior margin reddish brown; posterior lobe of pronotum pale yellow, with darker sublateral stripes (
<figureCitation id="8BD5E6C0FFEB3D6AFC99B466FC85560A" box="[783,853,1538,1563]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Fig. 7</figureCitation>
). Venter of thorax black to dark brown for the most part; posterior margin of metepimeron brownish yellow; peritreme yellowish. Scutellum (
<figureCitation id="8BD5E6C0FFEB3D6AFB3EB442FB3E562E" box="[1192,1262,1574,1599]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Fig. 7</figureCitation>
) black to dark brown in basal half and yellowish in apical half; apical half of scutellum with pale yellowish white longitudinal stripe along meson. Corium (
<figureCitation id="8BD5E6C0FFEB3D6AFE74B40AFDF55696" box="[482,549,1646,1671]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Fig. 7</figureCitation>
) pale in basal two-thirds and dark brown in remaining part, with small brown marking around inner angle; clavus pale; hemelytral membrane hyaline, with pale brown transverse marking at middle. Coxae pale reddish brown; trochanters, femora and tibiae pale yellow; metafemur and metatibia with brown annulations; tarsomere I pale yellow, tarsomeres II and III pale brown. Abdomen (
<figureCitation id="8BD5E6C0FFEB3D6AFBBDB4BEFBAB56E2" box="[1067,1147,1754,1779]" captionStart="FIGURES 1416" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 1416. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="5" pageNumber="56">Fig. 14</figureCitation>
) reddish brown; posterior margin of segments V and VI, middle of sternites VII and VIII dark brownish yellow; each laterotergite brown.
</paragraph>
<paragraph id="1351FA45FFEB3D6AFF51B546FDC757FE" blockId="5.[151,1437,1034,2031]" pageId="5" pageNumber="56">
Body length (
<figureCitation id="8BD5E6C0FFEB3D6AFEF6B546FE74572A" box="[352,420,1826,1851]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Fig. 7</figureCitation>
) approximately 3 times width across humeri. Head (
<figureCitation id="8BD5E6C0FFEB3D6AFC64B546FBE5572A" box="[1010,1077,1826,1851]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="5" pageNumber="56">Fig. 7</figureCitation>
) 1.8 times as wide as its length, covered with both rough and fine punctures; tylus tumid upwards; clypeus weakly projecting anteriad; juga with decumbent setae. Eyes occupying nearly half area of head in lateral view, not reaching anterior margin of pronotum. Antennae covered with suberect setae; proportional lengths of segments I to IV 1.0: 1.5: 1.3: 2.2. Labium slightly exceeding posterior margin of prothorax, not reaching mesocoxae; proportional lengths of segments I to IV 1.2: 1.3: 1.1: 1.0.
</paragraph>
<paragraph id="1351FA45FFE83D69FF51B2FCFBC95196" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="57">
Pronotum (
<figureCitation id="8BD5E6C0FFE83D69FED2B2F3FE5750A1" box="[324,391,151,176]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Fig. 7</figureCitation>
) slightly more than 1.4 times as wide as its length, nearly trapezoidal, lacking collar, and with lateral margin of anterior lobe weakly serrate; posterior lobe 1.5 times as long as anterior lobe, slightly more convex than anterior lobe, covered with dense punctures; posterior margin almost straight. Venter of prothorax, mesepisterna and metepisterna covered with decumbent setae. Scutellum (
<figureCitation id="8BD5E6C0FFE83D69FB81B360FBB2510D" box="[1047,1122,260,285]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Fig. 7</figureCitation>
) triangular, weakly tumid transversely in basal half, covered with decumbent setae. Hemelytra (
<figureCitation id="8BD5E6C0FFE83D69FC25B343FC285151" box="[947,1016,295,320]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Fig. 7</figureCitation>
) exceeding apex of abdomen; apical margin of corium as long as claval suture. Profemur moderately expanded, armed with 2 spines ventrally at about apical third, the basal spine larger; tibiae and tarsi covered with long erect setae.
</paragraph>
<paragraph id="1351FA45FFE83D69FF51B3F0FC585209" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="57">
Abdomen (
<figureCitation id="8BD5E6C0FFE83D69FED1B3F0FE5D51BD" box="[327,397,404,429]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Fig. 7</figureCitation>
) wider than hemelytral width; sternum entirely covered with decumbent setae. Ovipositor (
<figureCitation id="8BD5E6C0FFE83D69FF09B3D3FF2651C1" box="[159,246,439,464]" captionStart="FIGURES 1719" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 1719. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="6" pageNumber="57">Fig. 17</figureCitation>
) straight, reaching posterior margin of sternite VI, approximate oviposital length
<quantity id="D41657A0FFE83D69FB50B3D3FAE251DE" box="[1222,1330,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="57" unit="mm" value="0.45">0.45 mm</quantity>
; the two hemiventrites of the medially completely bisected sternite VII are short therefore posterior margins of sternites III V straight, posterior margin of sternite VI weakly sinuate (
<figureCitation id="8BD5E6C0FFE83D69FCB1B39BFCAB5209" box="[807,891,511,536]" captionStart="FIGURES 1719" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 1719. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="6" pageNumber="57">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="1351FA45FFE83D69FF51B040FDA5522D" blockId="6.[151,1437,151,1400]" box="[199,629,548,573]" pageId="6" pageNumber="57">
Spermatheca (
<typeStatus id="CC5544E7FFE83D69FEF0B040FE05522D" box="[358,469,548,572]" pageId="6" pageNumber="57" type="paratype">paratypes</typeStatus>
) as in
<figureCitation id="8BD5E6C0FFE83D69FD88B040FDA1522D" box="[542,625,548,573]" captionStart="FIGURES 2024" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[173,1319,245,1801]" captionTargetId="figure@12.[151,1436,215,1822]" captionTargetPageId="12" captionText="FIGURES 2024. Sadoletus laosensis sp. nov. 20, Pygophore, dorsal view (setae omitted from left half); 21, left paramere, dorsal view; 2223, phallus (endosoma not everted), ventral (22) and lateral (23) views; 24, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437396/files/figure.png" pageId="6" pageNumber="57">Fig. 24</figureCitation>
.
</paragraph>
<paragraph id="1351FA45FFE83D69FF51B023FE6D5304" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="57">
Male (
<typeStatus id="CC5544E7FFE83D69FE86B02CFE535271" box="[272,387,584,608]" pageId="6" pageNumber="57" type="paratype">paratypes</typeStatus>
): resembles female in general appearance. Body (
<figureCitation id="8BD5E6C0FFE83D69FC26B023FC245271" box="[944,1012,583,608]" captionStart="FIGURES 69" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 69. Habitus of Heterogastridae species from Laos and Thailand. 67, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 89, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Fig. 6</figureCitation>
) evidently narrower; pygophore (
<figureCitation id="8BD5E6C0FFE83D69FAF9B023FF645295" captionStart="FIGURES 2024" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[173,1319,245,1801]" captionTargetId="figure@12.[151,1436,215,1822]" captionTargetPageId="12" captionText="FIGURES 2024. Sadoletus laosensis sp. nov. 20, Pygophore, dorsal view (setae omitted from left half); 21, left paramere, dorsal view; 2223, phallus (endosoma not everted), ventral (22) and lateral (23) views; 24, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437396/files/figure.png" pageId="6" pageNumber="57">Fig. 20</figureCitation>
) nearly spherical in dorsal view, covered with suberect setae; maximum width of dorsal sinus of posterior aperture more than one-quarter as wide at middle as pygophore, without collar along anterior margin; posterior margin of pygophore and cuplike sclerite not fused; paramere (
<figureCitation id="8BD5E6C0FFE83D69FCC2B0D0FC7552DD" box="[852,933,692,717]" captionStart="FIGURES 2024" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[173,1319,245,1801]" captionTargetId="figure@12.[151,1436,215,1822]" captionTargetPageId="12" captionText="FIGURES 2024. Sadoletus laosensis sp. nov. 20, Pygophore, dorsal view (setae omitted from left half); 21, left paramere, dorsal view; 2223, phallus (endosoma not everted), ventral (22) and lateral (23) views; 24, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437396/files/figure.png" pageId="6" pageNumber="57">Fig. 21</figureCitation>
) narrow, twisted at basal third, acute at apex, sensory processes projecting laterally; phallosoma (
<figureCitation id="8BD5E6C0FFE83D69FD48B0B3FCB752E1" box="[734,871,727,752]" captionStart="FIGURES 2024" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[173,1319,245,1801]" captionTargetId="figure@12.[151,1436,215,1822]" captionTargetPageId="12" captionText="FIGURES 2024. Sadoletus laosensis sp. nov. 20, Pygophore, dorsal view (setae omitted from left half); 21, left paramere, dorsal view; 2223, phallus (endosoma not everted), ventral (22) and lateral (23) views; 24, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437396/files/figure.png" pageId="6" pageNumber="57">Figs. 22, 23</figureCitation>
) nearly twice as long as its width in ventral view, wrinkled at apical margin.
</paragraph>
<paragraph id="1351FA45FFE83D69FF51B17BFA835424" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="57">
<emphasis id="219A2657FFE83D69FF51B17BFEA65329" bold="true" box="[199,374,799,824]" pageId="6" pageNumber="57">Measurements</emphasis>
. [
<specimenCount id="05E831CCFFE83D69FE1AB144FE735326" box="[396,419,800,823]" pageId="6" pageNumber="57" type="male"></specimenCount>
(n=3) /
<specimenCount id="05E831CCFFE83D69FD96B144FDC55326" box="[512,533,800,823]" pageId="6" pageNumber="57" type="female"></specimenCount>
(n=5), value for
<typeStatus id="CC5544E7FFE83D69FD40B17BFCE85329" box="[726,824,799,824]" pageId="6" pageNumber="57" type="holotype">holotype</typeStatus>
female in parentheses]. Body length 4.35.0 / 4.85.2 (5.2); body height between coxae and dorsum 1.11.25 / 1.251.35 (1.25); head length 0.80.95 / 0.751.0 (0.75), head width across eyes 1.21.25 / 1.251.35 (1.35); length of antennal segments IIV respectively 0.350.4 / 0.33 0.38 (0.38), 0.650.69 / 0.530.65 (0.58), 0.60.65 / 0.50.63 (0.5), 0.780.93 / 0.830.88 (0.83); length of rostral segments IIV respectively 0.380.43 / 0.40.43 (0.4), 0.430.48 / 0.380.48 (0.43), 0.350.36 / 0.330.38 (0.35), 0.30.31 / 0.310.35 (0.33); mesal pronotal length 1.11.25 / 1.11.3 (1.25); posterior pronotal width 1.381.5 / 1.51.8 (1.75); mesal scutellal length 0.750.83 / 0.80.95 (0.95); basal scutellal width 0.750.85 / 0.881.0 (1.0); hemelytral length 3.23.38 / 3.253.5 (3.5); maximum width across hemelytra 1.151.38 / 1.581.85 (1.85).
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE83D69FF51B65BFDAC5449" box="[199,636,1087,1112]" pageId="6" pageNumber="57" type="distribution">
<paragraph id="1351FA45FFE83D69FF51B65BFDAC5449" blockId="6.[151,1437,151,1400]" box="[199,636,1087,1112]" pageId="6" pageNumber="57">
<emphasis id="219A2657FFE83D69FF51B65BFEB05449" bold="true" box="[199,352,1087,1112]" pageId="6" pageNumber="57">Distribution.</emphasis>
<collectingCountry id="6BF9BAD5FFE83D69FEF1B624FE4D5446" box="[359,413,1088,1111]" name="Laos" pageId="6" pageNumber="57">Laos</collectingCountry>
(
<collectingRegion id="D12A34A7FFE83D69FE3AB65BFDFA5449" box="[428,554,1087,1112]" country="Laos" name="Oudomxai" pageId="6" pageNumber="57">Oudomxay</collectingRegion>
Prov.).
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE83D69FF51B600FAD5546D" box="[199,1285,1124,1149]" pageId="6" pageNumber="57" type="etymology">
<paragraph id="1351FA45FFE83D69FF51B600FAD5546D" blockId="6.[151,1437,151,1400]" box="[199,1285,1124,1149]" pageId="6" pageNumber="57">
<emphasis id="219A2657FFE83D69FF51B600FE9E546C" bold="true" box="[199,334,1124,1149]" pageId="6" pageNumber="57">Etymology.</emphasis>
The species epithet is the Latin adjective “
<taxonomicName id="D4EE81C6FFE83D69FCB9B600FC46546D" authorityName="Ban" authorityYear="2018" box="[815,918,1124,1148]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="219A2657FFE83D69FCB9B600FC46546D" box="[815,918,1124,1148]" italics="true" pageId="6" pageNumber="57">laosensis</emphasis>
</taxonomicName>
” referring to
<typeStatus id="CC5544E7FFE83D69FBB9B601FB8F546C" box="[1071,1119,1125,1149]" pageId="6" pageNumber="57">type</typeStatus>
locality,
<collectingCountry id="6BF9BAD5FFE83D69FB51B601FAD1546D" box="[1223,1281,1125,1148]" name="Laos" pageId="6" pageNumber="57">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE83D69FF51B6E3FC455569" pageId="6" pageNumber="57" type="discussion">
<paragraph id="1351FA45FFE83D69FF51B6E3FBB75521" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="57">
<emphasis id="219A2657FFE83D69FF51B6E3FEEB54B1" bold="true" box="[199,315,1159,1184]" pageId="6" pageNumber="57">Remarks.</emphasis>
This new species is most similar in general appearance to
<taxonomicName id="D4EE81C6FFE83D69FB87B6ECFB3E54B1" authority="Bergroth" authorityName="Bergroth" box="[1041,1262,1159,1184]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="219A2657FFE83D69FB87B6ECFBAA54B1" box="[1041,1146,1160,1184]" italics="true" pageId="6" pageNumber="57">S. bakeri</emphasis>
Bergroth
</taxonomicName>
, but it can be distinguished from the latter by a combination of the following characters: antennal segment II 1.5 times longer than segment I (vs. antennal segment II twice longer than segment I); labium exceeding posterior margin of prothorax slightly, not reaching mesocoxae (vs. reaching mesocoxae); coxae pale reddish brown (vs. coxae black); ground color of femora and tibiae pale yellow (vs. femora and tibiae yellowish brown).
</paragraph>
<paragraph id="1351FA45FFE83D69FF51B758FC455569" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="57">
<emphasis id="219A2657FFE83D69FF51B758FE5B5544" bold="true" box="[199,395,1340,1365]" pageId="6" pageNumber="57">Biological notes.</emphasis>
The specimens were collected on fruiting trees of
<taxonomicName id="D4EE81C6FFE83D69FC45B759FBC15545" box="[979,1041,1341,1364]" class="Magnoliopsida" family="Moraceae" genus="Ficus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="57" phylum="Tracheophyta" rank="genus">
<emphasis id="219A2657FFE83D69FC45B759FBC15545" box="[979,1041,1341,1364]" italics="true" pageId="6" pageNumber="57">Ficus</emphasis>
</taxonomicName>
sp. in the forest along a mountain stream with several other heterogastrid and rhyparochromid species.
</paragraph>
</subSubSection>
</treatment>
</document>