treatments-xml/data/7B/A5/7C/7BA57CE8E1BF50FDA5B874973727F170.xml
2024-06-21 12:41:18 +02:00

272 lines
27 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.895.38576" ID-GBIF-Dataset="f9f1c191-c386-4c8c-9abd-4b4e6fecd2a6" ID-PMC="PMC6906171" ID-Pensoft-Pub="1313-2970-895-1" ID-Pensoft-UUID="99AB66C9ACF058A581D9036697D415F3" ID-PubMed="31844411" ID-ZooBank="11350FE921E64D7AB1F916CE4639F406" ModsDocID="1313-2970-895-1" checkinTime="1575565024213" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas &amp; Gattolliat, Jean-Luc" docDate="2019" docId="7BA57CE8E1BF50FDA5B874973727F170" docLanguage="en" docName="ZooKeys 895: 1-117" docOrigin="ZooKeys 895" docSource="http://dx.doi.org/10.3897/zookeys.895.38576" docTitle="Labiobaetis roulade Kaltenbach &amp; Gattolliat, 2019, sp. nov." docType="treatment" docUuid="7C7F4A79-E767-4EEE-8ADF-FE2CB67F0309" docUuidSource="ZooBank" docVersion="5" id="99AB66C9ACF058A581D9036697D415F3" lastPageNumber="1" masterDocId="99AB66C9ACF058A581D9036697D415F3" masterDocTitle="The tremendous diversity of Labiobaetis Novikova &amp; Kluge in Indonesia (Ephemeroptera, Baetidae)" masterLastPageNumber="117" masterPageNumber="1" pageNumber="1" updateTime="1668168188677" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The tremendous diversity of Labiobaetis Novikova &amp; Kluge in Indonesia (Ephemeroptera, Baetidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kaltenbach, Thomas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>895</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>117</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.895.38576</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.895.38576</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-895-1</mods:identifier>
<mods:identifier type="ZooBank">11350FE921E64D7AB1F916CE4639F406</mods:identifier>
<mods:identifier type="Pensoft-UUID">99AB66C9ACF058A581D9036697D415F3</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="160631144" LSID="urn:lsid:zoobank.org:act:7C7F4A79-E767-4EEE-8ADF-FE2CB67F0309" httpUri="http://treatment.plazi.org/id/7BA57CE8E1BF50FDA5B874973727F170" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
5.
<taxonomicName LSID="7BA57CE8-E1BF-50FD-A5B8-74973727F170" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis roulade" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roulade">Labiobaetis roulade</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Labiobaetis roulade sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium." figureDoi="10.3897/zookeys.895.38576.figure9" httpUri="https://binary.pensoft.net/fig/361899" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure9">Figures 9</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Labiobaetis roulade sp. nov., larva morphology: a Foreleg b Femur dorsal seta c Femur ventral seta d Tibia dorsal seta e Femur dorsal margin f Fore claw g Tergum IV h Gill IV i Paraproct j Scape k Metanotum." figureDoi="10.3897/zookeys.895.38576.figure10" httpUri="https://binary.pensoft.net/fig/361900" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure10">10</figureCitation>
,
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Habitus, larvae, dorsal view: a Labiobaetis batakorum sp. nov. b Labiobaetis sulawesiensis sp. nov. c Labiobaetis sumbensis sp. nov. d Labiobaetis roulade sp. nov." figureDoi="10.3897/zookeys.895.38576.figure47" httpUri="https://binary.pensoft.net/fig/361938" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure47">47d</figureCitation>
,
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Larvae: a Labiobaetis batakorum sp. nov., ventral view, legs b Labiobaetis sumbensis sp. nov., ventral view, legs c Labiobaetis roulade sp. nov., ventral view, legs d Labiobaetis weifangae sp. nov., ventral view, legs e Labiobaetis gueuningi sp. nov., ventral view f Labiobaetis jonasi sp. nov., ventral view." figureDoi="10.3897/zookeys.895.38576.figure52" httpUri="https://binary.pensoft.net/fig/361943" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure52">52c</figureCitation>
,
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Distribution of Labiobaetis in Indonesia." figureDoi="10.3897/zookeys.895.38576.figure55" httpUri="https://binary.pensoft.net/fig/361946" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure55">55a</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
Following combination of characters: A) dorsal surface of labrum with submarginal arc of 1 + 8-9 feathered setae; B) maxillary palp approx. as long as galea-lacinia; C) labial palp segment II with broad thumb-like distomedial protuberance, segment III conical; D) fore femur rather slender, length 3.7
<normalizedToken originalValue="×">x</normalizedToken>
maximum width, dorsal margin with ca. 15 spine-like setae, apically rounded and with minute serration; E) paraproct distally not expanded, with ca. 21 stout marginal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva</emphasis>
(
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Labiobaetis roulade sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium." figureDoi="10.3897/zookeys.895.38576.figure9" httpUri="https://binary.pensoft.net/fig/361899" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure9">Figs 9</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Labiobaetis roulade sp. nov., larva morphology: a Foreleg b Femur dorsal seta c Femur ventral seta d Tibia dorsal seta e Femur dorsal margin f Fore claw g Tergum IV h Gill IV i Paraproct j Scape k Metanotum." figureDoi="10.3897/zookeys.895.38576.figure10" httpUri="https://binary.pensoft.net/fig/361900" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure10">10</figureCitation>
,
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Habitus, larvae, dorsal view: a Labiobaetis batakorum sp. nov. b Labiobaetis sulawesiensis sp. nov. c Labiobaetis sumbensis sp. nov. d Labiobaetis roulade sp. nov." figureDoi="10.3897/zookeys.895.38576.figure47" httpUri="https://binary.pensoft.net/fig/361938" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure47">47d</figureCitation>
,
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Larvae: a Labiobaetis batakorum sp. nov., ventral view, legs b Labiobaetis sumbensis sp. nov., ventral view, legs c Labiobaetis roulade sp. nov., ventral view, legs d Labiobaetis weifangae sp. nov., ventral view, legs e Labiobaetis gueuningi sp. nov., ventral view f Labiobaetis jonasi sp. nov., ventral view." figureDoi="10.3897/zookeys.895.38576.figure52" httpUri="https://binary.pensoft.net/fig/361943" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure52">52c</figureCitation>
). Body length 5.2-6.2 mm. Cerci ca. 1/2 of body length. Antenna approximately 2.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as head length.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
. Head, thorax, and abdomen dorsally brown, head and thorax with bright median, dorsal suture, thorax and abdomen with bright pattern as in
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Habitus, larvae, dorsal view: a Labiobaetis batakorum sp. nov. b Labiobaetis sulawesiensis sp. nov. c Labiobaetis sumbensis sp. nov. d Labiobaetis roulade sp. nov." figureDoi="10.3897/zookeys.895.38576.figure47" httpUri="https://binary.pensoft.net/fig/361938" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure47">Fig. 47d</figureCitation>
, forewing pads with bright striation. Head, thorax, and abdomen ventrally brown, legs transparent, femur ventrodistomedially and distally with brown spot and dorsally with brown streak along margin (
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Larvae: a Labiobaetis batakorum sp. nov., ventral view, legs b Labiobaetis sumbensis sp. nov., ventral view, legs c Labiobaetis roulade sp. nov., ventral view, legs d Labiobaetis weifangae sp. nov., ventral view, legs e Labiobaetis gueuningi sp. nov., ventral view f Labiobaetis jonasi sp. nov., ventral view." figureDoi="10.3897/zookeys.895.38576.figure52" httpUri="https://binary.pensoft.net/fig/361943" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure52">Fig. 52c</figureCitation>
), caudal filaments transparent with brown band at 1/3 of cerci.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Antenna</emphasis>
with scape and pedicel subcylindrical, with well-developed distolateral process at scape; flagellum with broad, apically blunt spines and fine, simple setae on apex of each segment.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Labrum</emphasis>
(
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Labiobaetis roulade sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium." figureDoi="10.3897/zookeys.895.38576.figure9" httpUri="https://binary.pensoft.net/fig/361899" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure9">Fig. 9a</figureCitation>
). Rectangular, length 0.8
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of 1 + 8-9 feathered setae. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid, pectinate setae; ventral surface with four short, spine-like setae near lateral and anterolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Right mandible</emphasis>
(
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Labiobaetis roulade sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium." figureDoi="10.3897/zookeys.895.38576.figure9" httpUri="https://binary.pensoft.net/fig/361899" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure9">Fig. 9b, c</figureCitation>
). Incisors fused. Outer and inner sets of denticles with 4 + 4 denticles. Inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Left mandible</emphasis>
(
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Labiobaetis roulade sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium." figureDoi="10.3897/zookeys.895.38576.figure9" httpUri="https://binary.pensoft.net/fig/361899" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure9">Fig. 9d, e</figureCitation>
). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles and one minute intermediate denticle. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola slightly convex. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="1">Both mandibles with lateral margins almost straight. Basal half with fine, simple setae and slightly lanceolate setae scattered over surface.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName genus="Hypopharynx" lsidName="Hypopharynx" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hypopharynx</emphasis>
</taxonomicName>
(
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Labiobaetis roulade sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium." figureDoi="10.3897/zookeys.895.38576.figure9" httpUri="https://binary.pensoft.net/fig/361899" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure9">Fig. 9f</figureCitation>
). Lingua longer than superlingua. Lingua longer than broad; medial tuft of stout setae poorly developed; distal half laterally expanded. Superlingua rounded; lateral margin rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Maxilla</emphasis>
(
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Labiobaetis roulade sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium." figureDoi="10.3897/zookeys.895.38576.figure9" httpUri="https://binary.pensoft.net/fig/361899" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure9">Fig. 9g</figureCitation>
). Galea-lacinia with one simple, robust apical seta under crown. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid, and pectinate. Medially with one bipectinate, spine-like seta and five long, simple setae. Maxillary palp approx. as long as length of galea-lacinia; two segmented; palp segment II 1.6
<normalizedToken originalValue="×">x</normalizedToken>
length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment slightly pointed, with excavation at inner distolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Labium</emphasis>
(
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Labiobaetis roulade sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium." figureDoi="10.3897/zookeys.895.38576.figure9" httpUri="https://binary.pensoft.net/fig/361899" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure9">Fig. 9h</figureCitation>
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with nine spine-like setae increasing in length distally; apex with two long and one medium, robust, pectinate setae; outer margin with seven spine-like setae increasing in length distally; ventral surface with short, fine, simple and short, spine-like setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and 2-3 medium, simple setae in anteromedial area; dorsally with a row of four long, spine-like setae near inner margin. Labial palp with segment I 0.8
<normalizedToken originalValue="×">x</normalizedToken>
length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with broad thumb-like distomedial protuberance; distomedial protuberance 0.5
<normalizedToken originalValue="×">x</normalizedToken>
width of base of segment III; inner and outer margin with short, fine, simple setae; dorsally with a row of four long, spine-like, simple setae near outer margin. Segment III conical; apex slightly pointed; length 0.8
<normalizedToken originalValue="×">x</normalizedToken>
width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<caption doi="10.3897/zookeys.895.38576.figure9" httpUri="https://binary.pensoft.net/fig/361899" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
<taxonomicName class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis roulade" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roulade">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis roulade</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
Labrum
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
Right mandible
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
Right prostheca
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
Left mandible
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
Left prostheca
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
<taxonomicName genus="Hypopharynx" lsidName="Hypopharynx" pageId="0" pageNumber="1" rank="genus">Hypopharynx</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
Maxilla
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
Labium.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Hind wing pads</emphasis>
absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Foreleg</emphasis>
(
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Labiobaetis roulade sp. nov., larva morphology: a Foreleg b Femur dorsal seta c Femur ventral seta d Tibia dorsal seta e Femur dorsal margin f Fore claw g Tergum IV h Gill IV i Paraproct j Scape k Metanotum." figureDoi="10.3897/zookeys.895.38576.figure10" httpUri="https://binary.pensoft.net/fig/361900" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure10">
Fig. 10
<normalizedToken originalValue="af">a-f</normalizedToken>
</figureCitation>
). Ratio of foreleg segments 1.3:1.0:0.6:0.2.
<emphasis italics="true" pageId="0" pageNumber="1">Femur</emphasis>
. Length ca. 4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with a row of ca. 15 curved, spine-like setae, apically rounded and with minute serration, and many fine, simple setae; length of setae 0.17
<normalizedToken originalValue="×">x</normalizedToken>
maximum width of femur. Apex rounded; with one pair of curved, spine-like setae, apically rounded and with minute serration, and some short, stout setae. Short, stout, lanceolate setae scattered along the ventral margin; femoral patch reduced to a few setae.
<emphasis italics="true" pageId="0" pageNumber="1">Tibia.</emphasis>
Dorsal margin with a row of short, lanceolate, apically rounded setae and many short, lanceolate, apically rounded setae scattered along the dorsal margin. Ventral margin with a row of short, curved, spine-like setae, on apex some spine-like, partly bipectinate setae and a tuft of fine, simple setae. Anterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal 1/2.
<emphasis italics="true" pageId="0" pageNumber="1">Tarsus.</emphasis>
Dorsal margin with a row of short, lanceolate, apically rounded setae and many short, lanceolate, apically rounded setae scattered along the dorsal margin. Ventral margin with a row of curved, spine-like setae. Tarsal claw with one row of 15 denticles; distally pointed; with 7-8 stripes; subapical setae absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Tergum</emphasis>
(
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Labiobaetis roulade sp. nov., larva morphology: a Foreleg b Femur dorsal seta c Femur ventral seta d Tibia dorsal seta e Femur dorsal margin f Fore claw g Tergum IV h Gill IV i Paraproct j Scape k Metanotum." figureDoi="10.3897/zookeys.895.38576.figure10" httpUri="https://binary.pensoft.net/fig/361900" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure10">Fig. 10g</figureCitation>
). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae and micropores. Posterior margin of tergum IV with rounded or triangular spines, wider than long.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gills</emphasis>
(
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Labiobaetis roulade sp. nov., larva morphology: a Foreleg b Femur dorsal seta c Femur ventral seta d Tibia dorsal seta e Femur dorsal margin f Fore claw g Tergum IV h Gill IV i Paraproct j Scape k Metanotum." figureDoi="10.3897/zookeys.895.38576.figure10" httpUri="https://binary.pensoft.net/fig/361900" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure10">Fig. 10h</figureCitation>
). Present on segments
<normalizedToken originalValue="IVII">I-VII</normalizedToken>
. Margin with small denticles intercalating fine simple setae. Tracheae partly extending from main trunk towards outer and inner margins. Gill I ca. 2/3 length of segment II. Gill IV as long as length of segments V and 2/3 VI combined. Gill VII as long as length of segments VIII and 1/3 IX combined.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paraproct</emphasis>
(
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Labiobaetis roulade sp. nov., larva morphology: a Foreleg b Femur dorsal seta c Femur ventral seta d Tibia dorsal seta e Femur dorsal margin f Fore claw g Tergum IV h Gill IV i Paraproct j Scape k Metanotum." figureDoi="10.3897/zookeys.895.38576.figure10" httpUri="https://binary.pensoft.net/fig/361900" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure10">Fig. 10i</figureCitation>
). Distally not expanded, with ca. 21 stout marginal spines. Surface scattered with U-shaped scale bases, fine, simple setae and micropores. Cercotractor with numerous small marginal spines.
</paragraph>
<caption doi="10.3897/zookeys.895.38576.figure10" httpUri="https://binary.pensoft.net/fig/361900" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
<taxonomicName class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis roulade" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roulade">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis roulade</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
Foreleg
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
Femur dorsal seta
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
Femur ventral seta
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
Tibia dorsal seta
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
Femur dorsal margin
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
Fore claw
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
Tergum IV
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
Gill IV
<emphasis bold="true" pageId="0" pageNumber="1">i</emphasis>
Paraproct
<emphasis bold="true" pageId="0" pageNumber="1">j</emphasis>
Scape
<emphasis bold="true" pageId="0" pageNumber="1">k</emphasis>
Metanotum.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Dedicated to Jean-Marc Elouard, the collector of the specimens and well-known ephemerologist. The name is an anagram of Elouard.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Indonesia: Sumatra.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biological aspects">
<paragraph pageId="0" pageNumber="1">Biological aspects.</paragraph>
<paragraph pageId="0" pageNumber="1">The specimens were collected at an altitude of 530 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type-material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Holotype.</emphasis>
Larva (on slide, GBIFCH 00592234), Indonesia, Sumatra Barat, Payakumbuh, south of Payakumbuh,
<geoCoordinate degrees="00" direction="south" minutes="16.55" orientation="latitude" precision="9" value="-0.2758333">00°16.55'S</geoCoordinate>
,
<geoCoordinate degrees="100" direction="east" minutes="36.28" orientation="longitude" precision="9" value="100.60467">100°36.28'E</geoCoordinate>
, 530 m, 28.V.2010, J.-M. Elouard leg. Temporary deposited in MZL before definitely housed in MZB.
<emphasis bold="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
9 larvae (2 on slides, GBIFCH 00592235, GBIFCH 00592250, 4 in alcohol, GBIFCH 00515364, deposited in MZL; 3 in alcohol, GBIFCH 00515363, GBIFCH 00657757, GBIFCH 00657756, deposited in ZSM), same data as holotype.
</paragraph>
</subSubSection>
</treatment>
</document>