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<document id="01D815006F8E752C02C8D8CD888EACC3" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFDFE52D534085DB606CFDD5" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) russiceps Assing 2017, spec. nov." docType="treatment" docUuid="0F8B55FE-9FAD-4393-8C10-94FBC35FCD77" docUuidSource="ZooBank" docVersion="13" lastPageNumber="148" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="147" updateTime="1699471407760" updateUser="ExternalLinkService">
<mods:mods id="DFB28742D8160A321B0ED9A0CEA2F7F5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BB2241A10C2C07B879751E15DB56D27E">
<mods:title id="52F56269BE1F48B409B7E425F2C01E29">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
<mods:name id="9F6A942C720B9B2FCB000457872D6BF7" type="personal">
<mods:role id="9AAB6E95BB2C3CD027DA860EA15D344C">
<mods:roleTerm id="B30D462ADD50071003B6A42402C4432F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="435FFF133D5D0DB71C76BD91CC135BE8">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="F3BC203E9C595D95089EAB15C67C767A">text</mods:typeOfResource>
<mods:relatedItem id="F3B0F57E70739C9ECD0A58210A5500D4" type="host">
<mods:titleInfo id="94A3087ACFE387CABF484A79A21E3CBA">
<mods:title id="E9DD999349BAAE742BFDF946F50F2AA8">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="B99C0B04B8EC561602F9719838A07151">
<mods:date id="38F861EC962B24879CD7CB8BE4C95866">2017</mods:date>
<mods:detail id="C2A01A118D610A3E8848608C2AE23CC8" type="pubDate">
<mods:number id="F6B88A7B23DEE91C2F73887EBB4EDB5C">2017-06-30</mods:number>
</mods:detail>
<mods:detail id="784402360C8B9FD6100DE0ABBA7A6F20" type="volume">
<mods:number id="CAF995EA5FB774FB3300C4CAA1CEE632">67</mods:number>
</mods:detail>
<mods:detail id="AA6C50094C4458298AC9AF8AED8160E7" type="issue">
<mods:number id="91D816C5005B12F5E1F6AD5FE495C533">1</mods:number>
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<mods:start id="01725D7B976D465227E8A2AD5535F4C9">117</mods:start>
<mods:end id="875932327A171A54B834811512926CD1">192</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="844EE8312204F943C23B5EFBDC420B25">journal article</mods:classification>
<mods:identifier id="DC6B690BCA2EC2CFF1D00F31ABAC8E00" type="CLB-Dataset">2472</mods:identifier>
<mods:identifier id="B964B170F16F58383CBE94C72385AA29" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
<mods:identifier id="EEECFF2AD68B8B4295AE8AFA1504BDB7" type="GBIF-Dataset">39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43</mods:identifier>
<mods:identifier id="4ACFBA9C85397F368B30324481C0EE78" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="C040E232C4945AF3F89DB46EDDAD5784" type="Zenodo-Dep">5742363</mods:identifier>
<mods:identifier id="0C1A6F492F735E1A409AB6E990EA10C6" type="ZooBank">FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE</mods:identifier>
</mods:mods>
<treatment id="03A787BAFFDFE52D534085DB606CFDD5" ID-DOI="http://doi.org/10.5281/zenodo.5888477" ID-GBIF-Taxon="192498198" ID-Zenodo-Dep="5888477" LSID="urn:lsid:zoobank.org:act:0F8B55FE-9FAD-4393-8C10-94FBC35FCD77" httpUri="http://treatment.plazi.org/id/03A787BAFFDFE52D534085DB606CFDD5" lastPageId="31" lastPageNumber="148" pageId="30" pageNumber="147">
<subSubSection id="C3146527FFDFE52C534085DB6661FCBB" pageId="30" pageNumber="147" type="nomenclature">
<paragraph id="8BB136ACFFDFE52C534085DB67B7FD5A" blockId="30.[869,1386,723,815]" box="[869,1212,723,750]" pageId="30" pageNumber="147">
<heading id="D0F981C0FFDFE52C534085DB67B7FD5A" box="[869,1212,723,750]" fontSize="12" level="2" pageId="30" pageNumber="147" reason="2">
<taxonomicName id="4C0E4D2FFFDFE52C534085DB6747FD5A" ID-CoL="5DDFK" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[869,1100,723,750]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="147" phylum="Arthropoda" rank="species" species="russiceps" status="sp. nov." subGenus="Zyras">
<emphasis id="B97AEABEFFDFE52C534085DB6090FD5A" box="[869,923,723,750]" italics="true" pageId="30" pageNumber="147">Zyras</emphasis>
(
<emphasis id="B97AEABEFFDFE52C538C85DB60D4FD5A" box="[937,991,723,750]" italics="true" pageId="30" pageNumber="147">Zyras</emphasis>
)
<emphasis id="B97AEABEFFDFE52C53C885DB6747FD5A" box="[1005,1100,723,750]" italics="true" pageId="30" pageNumber="147">russiceps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFDFE52C547685DB67B7FD5A" box="[1107,1212,723,750]" pageId="30" pageNumber="147" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BB136ACFFDFE52C534085F36661FCBB" blockId="30.[869,1386,723,815]" box="[869,1386,763,783]" pageId="30" pageNumber="147">
<uri id="FF9F3AAEFFDFE52C534085F36661FCBB" box="[869,1386,763,783]" pageId="30" pageNumber="147">
urn:lsid:zoobank.org:act:
<uuid id="FFA80C79FFDFE52C543685F36661FCBB" box="[1043,1386,763,783]" pageId="30" pageNumber="147">0F8B55FE-9FAD-4393-8C10-94FBC35FCD77</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFDFE52C534084136739FC9B" box="[869,1074,795,815]" pageId="30" pageNumber="147" type="description">
<paragraph id="8BB136ACFFDFE52C534084136739FC9B" blockId="30.[869,1386,723,815]" box="[869,1074,795,815]" pageId="30" pageNumber="147">
(
<figureCitation id="13352A29FFDFE52C534C841360A9FC9B" box="[873,930,795,815]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="30" pageNumber="147">Figs 14</figureCitation>
,
<figureCitation id="13352A29FFDFE52C538C841360B5FC9B" box="[937,958,795,815]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="30" pageNumber="147">61</figureCitation>
,
<figureCitation id="13352A29FFDFE52C53E0841360E8FC9B" box="[965,995,795,815]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="30" pageNumber="147">114</figureCitation>
,
<figureCitation id="13352A29FFDFE52C53CF84136726FC9B" box="[1002,1069,795,815]" captionStart="Figs 174–190" captionStartId="58.[189,229,2020,2042]" captionTargetBox="[190,1430,246,1968]" captionTargetId="figure-107@58.[189,1436,237,1976]" captionText="Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888372" httpUri="https://zenodo.org/record/5888372/files/figure.png" pageId="30" pageNumber="147">187–190</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFDFE52C531A844167BFFBD5" pageId="30" pageNumber="147" type="materials_examined">
<paragraph id="8BB136ACFFDFE52C531A844167BFFBD5" blockId="30.[831,1437,841,1122]" pageId="30" pageNumber="147">
<emphasis id="B97AEABEFFDFE52C531A844160D5FCD5" bold="true" box="[831,990,841,865]" pageId="30" pageNumber="147">Type material</emphasis>
:
<materialsCitation id="3B663CF1FFDFE52C53D784416757FBB5" ID-GBIF-Occurrence="3450162441" collectingDate="1988-04-18" collectingDateMax="1988-04-27" collectingDateMin="1988-04-18" collectionCode="BMNH" collectorName="M. J. D. Brendell." country="Thailand" determinerName="V. Assing" elevation="300" latitude="15.466666" location="Thung Yai Wildlife Sanctuary" longLatPrecision="1285" longitude="98.8" pageId="30" pageNumber="147" specimenCount="1" stateProvince="Tak Province" typeStatus="holotype">
<typeStatus id="54B5880EFFDFE52C53D78441675EFCD5" box="[1010,1109,841,865]" pageId="30" pageNumber="147" type="holotype">Holotype</typeStatus>
<specimenCount id="9D08FD25FFDFE52C544684426773FCD5" box="[1123,1144,842,865]" pageId="30" pageNumber="147">♂</specimenCount>
: “W.
<collectingCountry id="F319763CFFDFE52C549484416636FCD5" box="[1201,1341,841,865]" name="Thailand" pageId="30" pageNumber="147">THAILAND</collectingCountry>
:
<quantity id="4CF69B49FFDFE52C556B8441669FFCD5" box="[1358,1428,841,865]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="147" unit="m" value="300.0">
<elevation id="0023D19FFFDFE52C556B8441669FFCD5" box="[1358,1428,841,865]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="147" unit="m" value="300.0">300 m</elevation>
</quantity>
.,
<location id="8ED16077FFDFE52C531A84616792FC35" LSID="urn:lsid:plazi:treatment:03A787BAFFDFE52D534085DB606CFDD5:8ED16077FFDFE52C531A84616792FC35" box="[831,1177,873,897]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Thung Yai Wildlife Sanctuary" pageId="30" pageNumber="147" stateProvince="Tak Province">Thung Yai Wildlife Sanctuary</location>
.
<geoCoordinate id="EE3A506BFFDFE52C548E8461660BFC35" box="[1195,1280,873,897]" degrees="15" direction="north" minutes="28" orientation="latitude" pageId="30" pageNumber="147" precision="925" value="15.466666">15°28'N</geoCoordinate>
–
<geoCoordinate id="EE3A506BFFDFE52C55088461668BFC35" box="[1325,1408,873,897]" degrees="98" direction="east" minutes="48" orientation="longitude" pageId="30" pageNumber="147" precision="925" value="98.8">98°48'E</geoCoordinate>
. /
<collectingRegion id="49CAF84EFFDFE52C531A848160C6FC15" box="[831,973,905,929]" country="Thailand" name="Tak" pageId="30" pageNumber="147">Tak Province</collectingRegion>
,
<location id="8ED16077FFDFE52C53C6848167BFFC15" LSID="urn:lsid:plazi:treatment:03A787BAFFDFE52D534085DB606CFDD5:8ED16077FFDFE52C53C6848167BFFC15" box="[995,1204,905,929]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Umphang District" pageId="30" pageNumber="147" stateProvince="Tak Province">Umphang District</location>
,
<taxonomicName id="4C0E4D2FFFDFE52C54EF84816696FC15" authority="Bae Stream." authorityName="Bae Stream." box="[1226,1437,905,929]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="147" phylum="Arthropoda" rank="species" species="song">
<location id="8ED16077FFDFE52C54EF84816692FC15" LSID="urn:lsid:plazi:treatment:03A787BAFFDFE52D534085DB606CFDD5:8ED16077FFDFE52C54EF84816692FC15" box="[1226,1433,905,929]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Song Bae Stream" pageId="30" pageNumber="147" stateProvince="Tak Province">Song Bae Stream</location>
.
</taxonomicName>
<date id="FFB0106CFFDFE52C531A84A160DFFC75" box="[831,980,937,961]" pageId="30" pageNumber="147" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">
<collectingDate id="EFF4E984FFDFE52C531A84A160DFFC75" box="[831,980,937,961]" pageId="30" pageNumber="147" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">18–27.iv.1988</collectingDate>
</date>
. / Evergreen rain forest.
<collectorName id="26FB537AFFDFE52C54D784A26696FC75" box="[1266,1437,937,961]" pageId="30" pageNumber="147">M.J.D. Brendell.</collectorName>
B.M. 1988-183 / / Holotypus ♂
<taxonomicName id="4C0E4D2FFFDFE52C54B184C26624FC55" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[1172,1327,970,993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="147" phylum="Arthropoda" rank="species" species="russiceps" status="sp. nov.">
<emphasis id="B97AEABEFFDFE52C54B184C26624FC55" box="[1172,1327,970,993]" italics="true" pageId="30" pageNumber="147">Zyras russiceps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFDFE52C551384C26665FC55" box="[1334,1390,970,993]" pageId="30" pageNumber="147" rank="species">sp. n.</taxonomicNameLabel>
det.
<determinerName id="7E6DDA68FFDFE52C531A84E260A9FBB5" box="[831,930,1001,1025]" pageId="30" pageNumber="147">V. Assing</determinerName>
2016” (
<collectionCode id="ED1FAE69FFDFE52C53DB84E1675FFBB5" box="[1022,1108,1001,1025]" pageId="30" pageNumber="147">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFDFE52C544F84E26762FB95" ID-GBIF-Occurrence="3455147301" collectingDate="1988-04-18" collectingDateMax="1988-04-27" collectingDateMin="1988-04-18" collectionCode="BMNH" collectorName="M. J. D. Brendell." country="Thailand" determinerName="V. Assing" elevation="300" latitude="15.466666" location="Umphang District" longLatPrecision="1285" longitude="98.8" pageId="30" pageNumber="147" specimenCount="1" stateProvince="Tak" typeStatus="paratype">
<typeStatus id="54B5880EFFDFE52C544F84E267DFFBB6" box="[1130,1236,1002,1026]" pageId="30" pageNumber="147" type="paratype">Paratypes</typeStatus>
: 2 ♂♂: same data as holotype (
<collectionCode id="ED1FAE69FFDFE52C53EF83016715FB95" box="[970,1054,1033,1057]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="30" pageNumber="147">BMNH</collectionCode>
, cAss);
</materialsCitation>
<materialsCitation id="3B663CF1FFDFE52C5454830167BBFBD5" ID-GBIF-Occurrence="3455147311" collectingDate="1988-04-18" collectingDateMax="1988-04-27" collectingDateMin="1988-04-18" collectionCode="BMNH" collectorName="M. J. D. Brendell." country="Thailand" determinerName="V. Assing" elevation="300" latitude="15.466666" location="Umphang District" longLatPrecision="1285" longitude="98.8" pageId="30" pageNumber="147" specimenCount="1" stateProvince="Tak" typeStatus="paratype">
1 ♂: “
<collectingCountry id="F319763CFFDFE52C54898301663AFB95" box="[1196,1329,1033,1057]" name="Malaysia" pageId="30" pageNumber="147">MALAYSIA</collectingCountry>
:
<collectingRegion id="49CAF84EFFDFE52C551E83016693FB95" box="[1339,1432,1033,1057]" country="Malaysia" name="Selangor" pageId="30" pageNumber="147">Selangor</collectingRegion>
,
<location id="8ED16077FFDFE52C531A832167ACFBF5" LSID="urn:lsid:plazi:treatment:03A787BAFFDFE52D534085DB606CFDD5:8ED16077FFDFE52C531A832167ACFBF5" box="[831,1191,1065,1089]" country="Malaysia" name="Ulu Gombak Field Studies Centre" pageId="30" pageNumber="147" stateProvince="Selangor">Ulu Gombak Field Studies Centre</location>
(
<quantity id="4CF69B49FFDFE52C5491832167F1FBF5" box="[1204,1274,1065,1089]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="147" unit="m" value="250.0">
<elevation id="0023D19FFFDFE52C5491832167F1FBF5" box="[1204,1274,1065,1089]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="147" unit="m" value="250.0">250 m</elevation>
</quantity>
),
<date id="FFB0106CFFDFE52C552883216696FBF5" box="[1293,1437,1065,1089]" pageId="30" pageNumber="147" value="2004-03-02" valueMax="2004-03-18" valueMin="2004-03-02">
<collectingDate id="EFF4E984FFDFE52C552883216696FBF5" box="[1293,1437,1065,1089]" pageId="30" pageNumber="147" value="2004-03-02" valueMax="2004-03-18" valueMin="2004-03-02">2–18.III.2004</collectingDate>
</date>
(
<collectionCode id="ED1FAE69FFDFE52C536283416078FBD5" box="[839,883,1097,1121]" pageId="30" pageNumber="147">FIT</collectionCode>
), Maruyama M. et al.” (cMar)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFDFE52C531A838160C0FB55" pageId="30" pageNumber="147" type="etymology">
<paragraph id="8BB136ACFFDFE52C531A838160C0FB55" blockId="30.[831,1436,1161,1249]" pageId="30" pageNumber="147">
<emphasis id="B97AEABEFFDFE52C531A838160BDFB15" bold="true" box="[831,950,1161,1185]" pageId="30" pageNumber="147">Etymology</emphasis>
: The specific epithet is a noun in apposition and alludes to the distinctive coloration of the head (russus: red).
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFDFE52D531A8201608AFE95" lastPageId="31" lastPageNumber="148" pageId="30" pageNumber="147" type="description">
<paragraph id="8BB136ACFFDFE52C531A820160A7F955" blockId="30.[831,1437,1289,2081]" pageId="30" pageNumber="147">
<emphasis id="B97AEABEFFDFE52C531A820160CFFA95" bold="true" box="[831,964,1289,1313]" pageId="30" pageNumber="147">Description</emphasis>
: Rather large species; body length
<quantity id="4CF69B49FFDFE52C531A822160C8FAF5" box="[831,963,1321,1345]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" metricValueMax="8.3" metricValueMin="7.5" pageId="30" pageNumber="147" unit="mm" value="7.9" valueMax="8.3" valueMin="7.5">7.5–8.3 mm</quantity>
; length of forebody
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. Coloration (
<figureCitation id="13352A29FFDFE52C5359824160C6FAD5" box="[892,973,1353,1377]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="30" pageNumber="147">Figs 14</figureCitation>
,
<figureCitation id="13352A29FFDFE52C53FE824160FCFAD5" box="[987,1015,1353,1377]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="30" pageNumber="147">61</figureCitation>
,
<figureCitation id="13352A29FFDFE52C542182416726FAD5" box="[1028,1069,1353,1377]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="30" pageNumber="147">114</figureCitation>
) distinctive: head and pronotum reddish; elytra blackish-brown to black, with only the anterior margin (visible in antero-dorsal view) narrowly reddish; abdomen bicoloured: tergites II–V reddish, VI reddish with the median portion extensively black, and VII–VIII blackish except for the reddish antero-lateral portions, segments IX–X brownish; legs pale-yellowish; antennae black with the basal two antennomeres reddish to reddish-brown and the apical 2–4 antennomeres pale-yellow to dark-yellow, more or less sharply contrasting with the black preceding antennomeres; maxillary palpi reddish with the terminal palpomere yellowish.
</paragraph>
<paragraph id="8BB136ACFFDFE52C531A81E16700F795" blockId="30.[831,1437,1289,2081]" pageId="30" pageNumber="147">
Head (
<figureCitation id="13352A29FFDFE52C53A281E160D9F8B5" box="[903,978,1769,1793]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="30" pageNumber="147">Fig. 61</figureCitation>
) distinctly transverse, middle and anterior portion extensively impunctate; punctation in lateral and posterior dorsal portions sparse and rather coarse. Eyes very large, nearly twice as long as postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFDFE52C54138061678BF835" box="[1078,1152,1897,1921]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="30" pageNumber="147">Fig. 14</figureCitation>
)
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long; antennomeres IV–V weakly oblong, VI approximately as broad as long, VII–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long, and XI approximately as long as the combined length of IX and X.
</paragraph>
<paragraph id="8BB136ACFFDEE52D50B287E26135FDB5" blockId="31.[151,757,233,1729]" pageId="31" pageNumber="148">
Pronotum (
<figureCitation id="13352A29FFDEE52D513687E16256FEB5" box="[275,349,233,257]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="31" pageNumber="148">Fig. 61</figureCitation>
) approximately 1.15 times as broad as long and 1.3 times as broad as head, broadest near anterior angles, distinctly tapering posteriad; lateral margins straight in posterior two-thirds (dorsal view); punctation coarse and nearly regularly distributed, with or without impunctate patches in antero-lateral portion; impunctate median band narrow or indistinct; lateral margins each with five long black setae, antero-lateral portion of disc with several additional long black setae.
</paragraph>
<paragraph id="8BB136ACFFDEE52D50B2850161ACFD75" blockId="31.[151,757,233,1729]" pageId="31" pageNumber="148">
Elytra (
<figureCitation id="13352A29FFDEE52D50C185016224FD95" box="[228,303,521,545]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="31" pageNumber="148">Fig. 61</figureCitation>
) approximately 0.8 times as long as pronotum; punctation very coarse and very dense, slightly sparser at posterior margin; interstices much narrower than diameter of punctures (except at posterior margin). Hind wings fully developed. Metatarsomere I elongate, slightly longer than the combined length of II–IV.
</paragraph>
<paragraph id="8BB136ACFFDEE52D50B285C161C6FA55" blockId="31.[151,757,233,1729]" pageId="31" pageNumber="148">
Abdomen (
<figureCitation id="13352A29FFDEE52D513085C16264FD55" box="[277,367,713,737]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="31" pageNumber="148">Fig. 114</figureCitation>
) approximately as broad as elytra, with moderately deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a row of non-setiferous punctures; tergite III with a lateral setiferous puncture on either side, with or without scattered additional setiferous punctures on disc, and with approximately twelve setiferous punctures at posterior margin; tergite IV with a transverse row of approximately eight setiferous punctures across disc and with 14 setiferous punctures at posterior margin; tergite V with a transverse row of approximately ten setiferous punctures across disc and with 12–16 setiferous punctures at posterior margin; tergite VI with a transverse band of numerous non-setiferous punctures anteriorly, with a transverse row of 10–12 setiferous punctures across disc and with 12–14 setiferous punctures at posterior margin; tergite VII with a broad transverse band of numerous non-setiferous punctures in anterior portion, and with two transverse series of setiferous punctures each composed of 10–14 punctures in posterior portion, posterior margin with palisade fringe; tergite VIII (
<figureCitation id="13352A29FFDEE52D50BA826163F2FA35" box="[159,249,1385,1409]" captionStart="Figs 174–190" captionStartId="58.[189,229,2020,2042]" captionTargetBox="[190,1430,246,1968]" captionTargetId="figure-107@58.[189,1436,237,1976]" captionText="Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888372" httpUri="https://zenodo.org/record/5888372/files/figure.png" pageId="31" pageNumber="148">Fig. 189</figureCitation>
) with numerous long black setae in posterior third, posterior margin convex or obtusely pointed in the middle, without median concavity; sternites III–VIII with numerous long dark setae in posterior portion.
</paragraph>
<paragraph id="8BB136ACFFDEE52D50B282E2629AF975" blockId="31.[151,757,233,1729]" pageId="31" pageNumber="148">
♂: sternite VIII (
<figureCitation id="13352A29FFDEE52D517382E162BBF9B5" box="[342,432,1513,1537]" captionStart="Figs 174–190" captionStartId="58.[189,229,2020,2042]" captionTargetBox="[190,1430,246,1968]" captionTargetId="figure-107@58.[189,1436,237,1976]" captionText="Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888372" httpUri="https://zenodo.org/record/5888372/files/figure.png" pageId="31" pageNumber="148">Fig. 190</figureCitation>
) with strongly convex posterior margin and with numerous long black setae in posterior half; median lobe of aedeagus (
<figureCitation id="13352A29FFDEE52D527C812161E6F9F5" box="[601,749,1577,1601]" captionStart="Figs 174–190" captionStartId="58.[189,229,2020,2042]" captionTargetBox="[190,1430,246,1968]" captionTargetId="figure-107@58.[189,1436,237,1976]" captionText="Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888372" httpUri="https://zenodo.org/record/5888372/files/figure.png" pageId="31" pageNumber="148">Figs 187–188</figureCitation>
)
<quantity id="4CF69B49FFDEE52D50B2814163FCF9D5" box="[151,247,1609,1633]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="31" pageNumber="148" unit="mm" value="0.98">0.98 mm</quantity>
long; ventral process apically acute both in lateral and in ventral view, subapically abruptly bent; paramere as long as median lobe and with conspicuously short apical lobe.
</paragraph>
<paragraph id="8BB136ACFFDEE52D50B281E1608AFE95" blockId="31.[151,756,1769,2049]" lastBlockId="31.[794,1399,233,289]" pageId="31" pageNumber="148">
<emphasis id="B97AEABEFFDEE52D50B281E16278F8B5" bold="true" box="[151,371,1769,1793]" pageId="31" pageNumber="148">Comparative notes</emphasis>
:
<taxonomicName id="4C0E4D2FFFDEE52D51AE81E26138F8B5" authorityName="Assing" authorityYear="2017" box="[395,563,1770,1793]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="148" phylum="Arthropoda" rank="species" species="russiceps">
<emphasis id="B97AEABEFFDEE52D51AE81E26138F8B5" box="[395,563,1770,1793]" italics="true" pageId="31" pageNumber="148">Zyras russiceps</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C0E4D2FFFDEE52D50B2800263FAF895" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[151,241,1801,1825]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="148" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B97AEABEFFDEE52D50B2800263FAF895" box="[151,241,1801,1825]" italics="true" pageId="31" pageNumber="148">Z. hirtus</emphasis>
</taxonomicName>
group, as can be inferred from the chaetotaxy of the body, the morphology of the median lobe of the aedeagus, and the short apical lobe of the paramere. It is distinguished from other species of this group recorded from
<collectingCountry id="F319763CFFDEE52D50F28081623CF815" box="[215,311,1929,1953]" name="Thailand" pageId="31" pageNumber="148">Thailand</collectingCountry>
and adjacent regions by the distinctive coloration (particularly of the head) alone. In addition, it is characterized by the conspicuously dense and coarse punctation of the elytra, by the punctation pattern of the abdomen, and by the shape of the median lobe of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFDEE52D533F8641606CFDD5" pageId="31" pageNumber="148" type="distribution">
<paragraph id="8BB136ACFFDEE52D533F8641606CFDD5" blockId="31.[794,1399,329,609]" pageId="31" pageNumber="148">
<emphasis id="B97AEABEFFDEE52D533F8641678EFED5" bold="true" box="[794,1157,329,353]" pageId="31" pageNumber="148">Distribution and natural history</emphasis>
: The known distribution is confined to the type locality in West
<collectingCountry id="F319763CFFDEE52D55318661667DFE35" box="[1300,1398,361,385]" name="Thailand" pageId="31" pageNumber="148">Thailand</collectingCountry>
and one locality in
<collectingRegion id="49CAF84EFFDEE52D53D186816758FE15" box="[1012,1107,393,417]" country="Malaysia" name="Selangor" pageId="31" pageNumber="148">Selangor</collectingRegion>
, Peninsular
<collectingCountry id="F319763CFFDEE52D54F88681664BFE15" box="[1245,1344,393,417]" name="Malaysia" pageId="31" pageNumber="148">Malaysia</collectingCountry>
. The specimens from
<collectingCountry id="F319763CFFDEE52D53F286A16732FE75" box="[983,1081,425,449]" name="Thailand" pageId="31" pageNumber="148">Thailand</collectingCountry>
were probably collected on the wing (Malaise trap?) in an evergreen rain forest at an altitude of
<quantity id="4CF69B49FFDEE52D539E86E1670BFDB5" box="[955,1024,489,513]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="148" unit="m" value="300.0">300 m</quantity>
, the
<typeStatus id="54B5880EFFDEE52D541E86E26791FDB6" box="[1083,1178,490,514]" pageId="31" pageNumber="148" type="paratype">paratype</typeStatus>
from
<collectingCountry id="F319763CFFDEE52D54C186E1664FFDB5" box="[1252,1348,489,513]" name="Malaysia" pageId="31" pageNumber="148">Malaysia</collectingCountry>
was taken with a flight interception trap at
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, in both cases together with several other species of
<taxonomicName id="4C0E4D2FFFDEE52D54D28522667CFDF5" authority="sensu" authorityName="SenSU" box="[1271,1399,554,577]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEABEFFDEE52D54D285226639FDF5" box="[1271,1330,554,577]" italics="true" pageId="31" pageNumber="148">Zyras</emphasis>
sensu
</taxonomicName>
strictu.
</paragraph>
</subSubSection>
</treatment>
</document>
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