treatments-xml/data/03/A7/87/03A787A4FFFDFFD13584F44B1867FF5D.xml

<document id="8F5C57EF4C6F8D27307731DDE2D10F12" ID-CLB-Dataset="7383" ID-DOI="10.11646/zootaxa.4951.3.7" ID-GBIF-Dataset="a6ee038f-4cda-4f38-8fde-8ad0efec8f0f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4668453" ID-ZooBank="EB876346-4282-4B4E-8585-ED50F1A616E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617782094822" checkinUser="plazi" docAuthor="Martens, Jochen, Maghradze, Eter &amp; Barjadze, Shalva" docDate="2021" docId="03A787A4FFFDFFD13584F44B1867FF5D" docLanguage="en" docName="zootaxa.4951.3.7.pdf" docOrigin="Zootaxa 4951 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nemaspela melouri Martens &amp; Maghradze &amp; Barjadze 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="548" masterDocId="FF9EFFDCFFFFFFD63513F30F1D1AFFA4" masterDocTitle="Two new species of the genus Nemaspela Šilhavý from caves in Georgia (Opiliones Nemastomatidae)" masterLastPageNumber="558" masterPageNumber="541" pageNumber="543" updateTime="1698936134096" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="36FEDC903E01187598A2BDCA7AEEA545">Two new species of the genus Nemaspela Šilhavý from caves in Georgia (Opiliones Nemastomatidae)</mods:title>
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<mods:namePart id="B4BE4413E9AE4DE0225B8D7013B89025">Martens, Jochen</mods:namePart>
<mods:affiliation id="0330F08BCCB90AA2CC9461A64A4E2F2E">Johannes Gutenberg-Universität, Institut für Organismische und Molekulare Evolutionsbiologie (iomE), D- 55099 Mainz, Germany; Senckenberg Research Institute, Arachnology, D- 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:namePart id="4613C9D89DA07A414D7569F4BD0E5257">Maghradze, Eter</mods:namePart>
<mods:affiliation id="39006B6E7F60366DC15DCED56871669D">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia</mods:affiliation>
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</mods:role>
<mods:namePart id="6C5A54DF0981A5ACB9C9BBDC4754F12B">Barjadze, Shalva</mods:namePart>
<mods:affiliation id="1322B28378E3E40A323E6F9C92388704">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia</mods:affiliation>
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<mods:title id="D218BE36B213C336D1DB91949926FF3D">Zootaxa</mods:title>
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<mods:date id="775EF4335BDB3B5C9249305936B2CA2F">2021</mods:date>
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<mods:number id="2A0B2F8F27E05450B255B37461B757CA">2021-04-07</mods:number>
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<mods:number id="9D3A7F73008AA6D413CF90508B4CC237">4951</mods:number>
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<treatment id="03A787A4FFFDFFD13584F44B1867FF5D" ID-DOI="http://doi.org/10.5281/zenodo.4685057" ID-GBIF-Taxon="181003514" ID-Zenodo-Dep="4685057" LSID="urn:lsid:plazi:treatment:03A787A4FFFDFFD13584F44B1867FF5D" httpUri="http://treatment.plazi.org/id/03A787A4FFFDFFD13584F44B1867FF5D" lastPageId="7" lastPageNumber="548" pageId="2" pageNumber="543">
<subSubSection id="C3146539FFFDFFD43584F44B1CF2F8FB" box="[151,488,1860,1887]" pageId="2" pageNumber="543" type="nomenclature">
<paragraph id="8BB136B2FFFDFFD43584F44B1CF2F8FB" blockId="2.[151,488,1860,1923]" box="[151,488,1860,1887]" pageId="2" pageNumber="543">
<heading id="D0F981DEFFFDFFD43584F44B1CF2F8FB" bold="true" box="[151,488,1860,1887]" fontSize="11" level="1" pageId="2" pageNumber="543" reason="1">
<emphasis id="B97AEAA0FFFDFFD43584F44B1CF2F8FB" bold="true" box="[151,488,1860,1887]" pageId="2" pageNumber="543">
<taxonomicName id="4C0E4D31FFFDFFD43584F44B1C99F8FB" ID-CoL="9X5J9" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[151,387,1860,1887]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="melouri" status="sp. nov.">
<emphasis id="B97AEAA0FFFDFFD43584F44B1C99F8FB" bold="true" box="[151,387,1860,1887]" italics="true" pageId="2" pageNumber="543">Nemaspela melouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957DBFFFDFFD43499F44A1CF2F8FB" box="[394,488,1861,1887]" pageId="2" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFFDFFD43584F4671C63F827" box="[151,377,1896,1923]" pageId="2" pageNumber="543" type="description">
<paragraph id="8BB136B2FFFDFFD43584F4671C63F827" blockId="2.[151,488,1860,1923]" box="[151,377,1896,1923]" pageId="2" pageNumber="543">
(
<figureCitation id="13352A37FFFDFFD4358CF4671C08F827" box="[159,274,1896,1923]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–12" captionStart-2="FIGURES 13–18" captionStartId-0="4.[151,264,1924,1948]" captionStartId-1="6.[151,264,1943,1967]" captionStartId-2="7.[151,264,1195,1219]" captionTargetBox-0="[152,1436,345,1900]" captionTargetBox-1="[194,1389,197,1903]" captionTargetBox-2="[166,1421,285,1171]" captionTargetId-0="figure-105@4.[151,1436,345,1901]" captionTargetId-1="figure-23@6.[180,1408,181,1919]" captionTargetId-2="figure-53@7.[151,1436,273,1172]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 1–4. Nemaspela melouri sp. nov., Habitus of paratype specimens in lateral view. 1. Male (bar: 2 mm); 2. Male (bar: 1.5 mm); 3. Female CJM 8705 (bar: 1.5 mm); 4. Female CJM 8705 (bar: 1 mm). Arrow indicates male cheliceral apophysis. Photographs by J. Severin." captionText-1="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." captionText-2="FIGURES 13–18. Nemaspela melouri sp. nov. Chelicerae in retro-lateral view. 13. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Cheliceral apophyses in retro-lateral view. 15. The first male (bar: 0.05 mm); 16. The second male (bar: 0.05 mm). Cheliceral apophyses in pro-lateral view.17. The first male (bar: 0.05 mm); 18. The second male (bar: 0.05 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.4668457" figureDoi-1="http://doi.org/10.5281/zenodo.4668461" figureDoi-2="http://doi.org/10.5281/zenodo.4668465" httpUri-0="https://zenodo.org/record/4668457/files/figure.png" httpUri-1="https://zenodo.org/record/4668461/files/figure.png" httpUri-2="https://zenodo.org/record/4668465/files/figure.png" pageId="2" pageNumber="543">Figs 1–18</figureCitation>
, 
<tableCitation id="C68C0309FFFDFFD4340EF4671C74F827" box="[285,366,1896,1923]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Measurements of Nemaspela melouri sp. nov. Measurements are given in millimeters. Note: body length is measured in all available specimens, while length of appendages was measured only in holotype male and single female." httpUri="http://table.plazi.org/id/DF71663AFFFAFFD33584F3981DC8FF53" pageId="2" pageNumber="543" tableUuid="DF71663AFFFAFFD33584F3981DC8FF53">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFFDFFD53584F4BE1947FF71" lastPageId="3" lastPageNumber="544" pageId="2" pageNumber="543" type="materials_examined">
<paragraph id="8BB136B2FFFDFFD43584F4BE1984F84B" blockId="2.[151,1436,1968,2031]" pageId="2" pageNumber="543">
<emphasis id="B97AEAA0FFFDFFD43584F4BE1C52F86E" bold="true" box="[151,328,1968,1995]" pageId="2" pageNumber="543">Type material:</emphasis>
<materialsCitation id="3B663CEFFFFDFFD43443F4BF1984F84B" ID-GBIF-Occurrence="3071286301" collectingDate="2020-02-01" collectionCode="IZISU" country="Georgia" county="West" elevation="415" latitude="42.38753" location="Melouri Cave" longLatPrecision="1" longitude="42.628193" municipality="Sataplia-Tskaltubo" pageId="2" pageNumber="543" specimenCount="1" specimenCount-male="1" stateProvince="Imereti" typeStatus="holotype">
<typeStatus id="54B58810FFFDFFD43443F4BF1CA7F86E" box="[336,445,1968,1994]" pageId="2" pageNumber="543" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="9D08FD3BFFFDFFD434DBF4BE1CC5F86F" box="[456,479,1969,1995]" pageId="2" pageNumber="543" type="male">♂</specimenCount>
, 
<date id="FFB01072FFFDFFD434FFF4BF1F7AF86E" box="[492,608,1968,1994]" pageId="2" pageNumber="543" value="2020-02-01">
<collectingDate id="EFF4E99AFFFDFFD434FFF4BF1F7AF86E" box="[492,608,1968,1994]" pageId="2" pageNumber="543" value="2020-02-01">01.ii.2020</collectingDate>
</date>
, dark zone, 
<location id="8ED16069FFFDFFD437F9F4BF1E9CF86F" LSID="urn:lsid:plazi:treatment:03A787A4FFFDFFD13584F44B1867FF5D:8ED16069FFFDFFD437F9F4BF1E9CF86F" box="[746,902,1968,1995]" country="Georgia" county="West" latitude="42.38753" longLatPrecision="1" longitude="42.628193" municipality="Sataplia-Tskaltubo" name="Melouri Cave" pageId="2" pageNumber="543" stateProvince="Imereti">Melouri Cave</location>
(
<geoCoordinate id="EE3A5075FFFDFFD43685F4BF192DF86F" box="[918,1079,1968,1995]" degrees="42" direction="north" minutes="23" orientation="latitude" pageId="2" pageNumber="543" precision="1" seconds="15.1" value="42.38753">42°23’15.1”N</geoCoordinate>
, 
<geoCoordinate id="EE3A5075FFFDFFD43157F4BF19F8F86F" box="[1092,1250,1968,1995]" degrees="42" direction="east" minutes="37" orientation="longitude" pageId="2" pageNumber="543" precision="1" seconds="41.5" value="42.628193">42°37’41.5”E</geoCoordinate>
; 
<quantity id="4CF69B57FFFDFFD431FCF4BF182CF86F" box="[1263,1334,1968,1995]" metricMagnitude="2" metricUnit="m" metricValue="4.15" pageId="2" pageNumber="543" unit="m" value="415.0">
<elevation id="0023D181FFFDFFD431FCF4BF182CF86F" box="[1263,1334,1968,1995]" metricMagnitude="2" metricUnit="m" metricValue="4.15" pageId="2" pageNumber="543" unit="m" value="415.0">415 m</elevation>
</quantity>
alt.), Ts- kaltubo 
<location id="8ED16069FFFDFFD435E1F4DB1C9EF84A" LSID="urn:lsid:plazi:treatment:03A787A4FFFDFFD13584F44B1867FF5D:8ED16069FFFDFFD435E1F4DB1C9EF84A" box="[242,388,2004,2030]" country="Georgia" county="West" latitude="42.38753" longLatPrecision="1" longitude="42.628193" municipality="Sataplia-Tskaltubo" name="Municipality" pageId="2" pageNumber="543" stateProvince="Imereti">Municipality</location>
, 
<collectingMunicipality id="6BD5ACC8FFFDFFD4349CF4DB1F7BF84A" box="[399,609,2004,2030]" pageId="2" pageNumber="543">Sataplia-Tskaltubo</collectingMunicipality>
karst massif, 
<collectingRegion id="49CAF850FFFDFFD437EFF4DB1E84F84A" box="[764,926,2004,2030]" country="Georgia" name="Imereti" pageId="2" pageNumber="543">Imereti region</collectingRegion>
, 
<collectingCounty id="62D04E3EFFFDFFD436BAF4DA1EFAF84B" box="[937,992,2005,2031]" pageId="2" pageNumber="543">West</collectingCounty>
Georgia, 
<collectionCode id="ED1FAE77FFFDFFD4315CF4DA1980F84B" box="[1103,1178,2005,2031]" pageId="2" pageNumber="543">IZISU</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB136B2FFFCFFD535D4F3981947FF71" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
<materialsCitation id="3B663CEFFFFCFFD535D4F3981F81FF15" ID-GBIF-Occurrence="3071286308" box="[199,667,151,177]" collectingDate="2020-02-01" collectionCode="IZISU" country="Georgia" county="West" elevation="415" latitude="42.38753" location="The" longLatPrecision="1" longitude="42.628193" municipality="Sataplia-Tskaltubo" pageId="3" pageNumber="544" specimenCount="1" stateProvince="Imereti" typeStatus="paratype">
<emphasis id="B97AEAA0FFFCFFD535D4F3981C25FF15" bold="true" box="[199,319,151,177]" pageId="3" pageNumber="544">
<typeStatus id="54B58810FFFCFFD535D4F3981C25FF15" box="[199,319,151,177]" pageId="3" pageNumber="544" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<location id="8ED16069FFFCFFD5345FF3981C62FF15" LSID="urn:lsid:plazi:treatment:03A787A4FFFDFFD13584F44B1867FF5D:8ED16069FFFCFFD5345FF3981C62FF15" box="[332,376,151,177]" name="The" pageId="3" pageNumber="544">The</location>
same locality as holotype
</materialsCitation>
; 
<materialsCitation id="3B663CEFFFFCFFD537B6F3981EBEFF15" ID-GBIF-Occurrence="3071286303" box="[677,932,151,177]" collectingDate="2019-02-21" collectionCode="IZISU" country="Georgia" pageId="3" pageNumber="544" specimenCount="2" specimenCount-female="2" stateProvince="Imereti" typeStatus="paratype">
<specimenCount id="9D08FD3BFFFCFFD537B6F3981FD4FF15" box="[677,718,151,177]" pageId="3" pageNumber="544" type="female">2 ♀</specimenCount>
, 
<date id="FFB01072FFFCFFD537C8F3981E54FF15" box="[731,846,151,177]" pageId="3" pageNumber="544" value="2019-02-21">
<collectingDate id="EFF4E99AFFFCFFD537C8F3981E54FF15" box="[731,846,151,177]" pageId="3" pageNumber="544" value="2019-02-21">21.ii.2019</collectingDate>
</date>
, 
<collectionCode id="ED1FAE77FFFCFFD5364AF3981EBEFF15" box="[857,932,151,177]" pageId="3" pageNumber="544">IZISU</collectionCode>
</materialsCitation>
; 
<specimenCount id="9D08FD3BFFFCFFD536BCF3981EC2FF15" box="[943,984,151,177]" pageId="3" pageNumber="544" type="female">2 ♀</specimenCount>
, 
<date id="FFB01072FFFCFFD536F6F3981943FF15" box="[997,1113,151,177]" pageId="3" pageNumber="544" value="2020-02-01">01.ii.2020</date>
, CJM 8705; 
<specimenCount id="9D08FD3BFFFCFFD531F5F398180BFF15" box="[1254,1297,151,177]" pageId="3" pageNumber="544" type="male">1 ♂</specimenCount>
, 
<date id="FFB01072FFFCFFD5300FF398188DFF15" box="[1308,1431,151,177]" pageId="3" pageNumber="544" value="2020-03-03">03.iii.2020</date>
, CJM 8706; 
<materialsCitation id="3B663CEFFFFCFFD53408F3B41947FF71" ID-GBIF-Occurrence="3071286307" box="[283,1117,187,213]" collectingDate="2020-03-03" collectionCode="IZISU" collectorName="All &amp; Eter Maghradze." country="Georgia" pageId="3" pageNumber="544" specimenCount="2" specimenCount-male="2" stateProvince="Imereti" typeStatus="paratype">
<specimenCount id="9D08FD3BFFFCFFD53408F3B41C5DFF71" box="[283,327,187,213]" pageId="3" pageNumber="544" type="male">2 ♂</specimenCount>
, 
<date id="FFB01072FFFCFFD53440F3B41CD5FF71" box="[339,463,187,213]" pageId="3" pageNumber="544" value="2020-03-03">
<collectingDate id="EFF4E99AFFFCFFD53440F3B41CD5FF71" box="[339,463,187,213]" pageId="3" pageNumber="544" value="2020-03-03">03.iii.2020</collectingDate>
</date>
, 
<collectionCode id="ED1FAE77FFFCFFD534C8F3B41F3CFF71" box="[475,550,187,213]" pageId="3" pageNumber="544">IZISU</collectionCode>
. 
<collectorName id="26FB5364FFFCFFD5373CF3B41F49FF71" box="[559,595,187,213]" pageId="3" pageNumber="544">All</collectorName>
specimens were collected by 
<collectorName id="26FB5364FFFCFFD536B1F3B41947FF71" box="[930,1117,187,213]" pageId="3" pageNumber="544">Eter Maghradze.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFFCFFD535D4F3D119F7FE76" pageId="3" pageNumber="544" type="diagnosis">
<paragraph id="8BB136B2FFFCFFD535D4F3D119F7FE76" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
<emphasis id="B97AEAA0FFFCFFD535D4F3D11C58FF5C" bold="true" box="[199,322,222,248]" pageId="3" pageNumber="544">Diagnosis.</emphasis>
Generally similar in size and habitus to the other 
<taxonomicName id="4C0E4D31FFFCFFD53680F3D01915FF5D" authorityName="Silhavy" authorityYear="1966" box="[915,1039,223,249]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="544" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEAA0FFFCFFD53680F3D01915FF5D" box="[915,1039,223,249]" italics="true" pageId="3" pageNumber="544">Nemaspela</emphasis>
</taxonomicName>
species from Georgian caves, but considerably smaller and with less sclerotized cuticle, and most similar in external characters to the two species from localities in the close vicinity (
<taxonomicName id="4C0E4D31FFFCFFD5373BF2281FF0FEE5" authorityName="Martens" authorityYear="2006" box="[552,746,295,321]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="femorecurvata">
<emphasis id="B97AEAA0FFFCFFD5373BF2281FF0FEE5" box="[552,746,295,321]" italics="true" pageId="3" pageNumber="544">N. femorecurvata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0E4D31FFFCFFD53633F2281EDBFEE5" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[800,961,295,321]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B97AEAA0FFFCFFD53633F2281EDBFEE5" box="[800,961,295,321]" italics="true" pageId="3" pageNumber="544">N. prometheus</emphasis>
</taxonomicName>
<emphasis id="B97AEAA0FFFCFFD536D4F228193AFEE5" bold="true" box="[967,1056,295,321]" pageId="3" pageNumber="544">
<taxonomicNameLabel id="A24957DBFFFCFFD536D4F228193AFEE5" box="[967,1056,295,321]" pageId="3" pageNumber="544" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), characterized by small size (less than 
<quantity id="4CF69B57FFFCFFD535DCF2441C08FEC1" box="[207,274,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="544" unit="mm" value="2.0">2 mm</quantity>
, slightly smaller than 
<taxonomicName id="4C0E4D31FFFCFFD5371FF2441FB5FEC1" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[524,687,331,357]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B97AEAA0FFFCFFD5371FF2441FB5FEC1" box="[524,687,331,357]" italics="true" pageId="3" pageNumber="544">N. prometheus</emphasis>
</taxonomicName>
<emphasis id="B97AEAA0FFFCFFD537A5F2441E0AFEC1" bold="true" box="[694,784,331,357]" pageId="3" pageNumber="544">
<taxonomicNameLabel id="A24957DBFFFCFFD537A5F2441E0AFEC1" box="[694,784,331,357]" pageId="3" pageNumber="544" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
including slightly shorter appendages), extremely weakly sclerotized exoskeleton, specific genital morphology (slender form of truncus and small size of basal inflated part of truncus) and armament of male chelicera (with small frontad-directed apophysis on basal article of male chelicera). No sexual dimorphism of appendages (pedipalp and legs in males and in females of the same size).
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFFCFFD535D4F2D41C6CFDBD" pageId="3" pageNumber="544" type="etymology">
<paragraph id="8BB136B2FFFCFFD535D4F2D41C6CFDBD" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
<emphasis id="B97AEAA0FFFCFFD535D4F2D41C54FE51" bold="true" box="[199,334,475,501]" pageId="3" pageNumber="544">Etymology.</emphasis>
The name is given in accordance with the 
<typeStatus id="54B58810FFFCFFD53625F2D41E7CFE51" box="[822,870,475,501]" pageId="3" pageNumber="544">type</typeStatus>
locality and single known locality of this species. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFFCFFD135D4F12D1F33FF15" lastPageId="7" lastPageNumber="548" pageId="3" pageNumber="544" type="description">
<paragraph id="8BB136B2FFFCFFD535D4F12D184DFD0C" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
<emphasis id="B97AEAA0FFFCFFD535D4F12D1CFDFD98" bold="true" box="[199,487,546,573]" pageId="3" pageNumber="544">Description. Male (n=4).</emphasis>
<emphasis id="B97AEAA0FFFCFFD534FDF12C1FB7FD99" box="[494,685,547,573]" italics="true" pageId="3" pageNumber="544">Body, dorsal side</emphasis>
(
<figureCitation id="13352A37FFFCFFD537A9F12C1E04FD99" box="[698,798,547,573]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1924,1948]" captionTargetBox="[152,1436,345,1900]" captionTargetId="figure-105@4.[151,1436,345,1901]" captionTargetPageId="4" captionText="FIGURES 1–4. Nemaspela melouri sp. nov., Habitus of paratype specimens in lateral view. 1. Male (bar: 2 mm); 2. Male (bar: 1.5 mm); 3. Female CJM 8705 (bar: 1.5 mm); 4. Female CJM 8705 (bar: 1 mm). Arrow indicates male cheliceral apophysis. Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668457" httpUri="https://zenodo.org/record/4668457/files/figure.png" pageId="3" pageNumber="544">Figs 1–2</figureCitation>
, 
<figureCitation id="13352A37FFFCFFD5363AF12C1E5DFD99" box="[809,839,547,573]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">11</figureCitation>
): Prosoma and opisthosoma forming two independent strongly globular entities with a deep depression between them; dorsal scutum uniformly light brownish to yellowish, smooth, difficult to distinguish from neighbouring exoskeletal parts, without any darker spots or markings, no setae present on dorsal side except for a few minute ones on rear end of opisthosoma, Tu oc and eyes absent.
</paragraph>
<paragraph id="8BB136B2FFFCFFD535D4F1BC1913FCB1" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
<emphasis id="B97AEAA0FFFCFFD535D4F1BC1C95FD69" box="[199,399,691,717]" italics="true" pageId="3" pageNumber="544">Body, ventral side</emphasis>
(
<figureCitation id="13352A37FFFCFFD5348EF1BC1CFFFD69" box="[413,485,691,717]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1924,1948]" captionTargetBox="[152,1436,345,1900]" captionTargetId="figure-105@4.[151,1436,345,1901]" captionTargetPageId="4" captionText="FIGURES 1–4. Nemaspela melouri sp. nov., Habitus of paratype specimens in lateral view. 1. Male (bar: 2 mm); 2. Male (bar: 1.5 mm); 3. Female CJM 8705 (bar: 1.5 mm); 4. Female CJM 8705 (bar: 1 mm). Arrow indicates male cheliceral apophysis. Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668457" httpUri="https://zenodo.org/record/4668457/files/figure.png" pageId="3" pageNumber="544">Figs 2</figureCitation>
, 
<figureCitation id="13352A37FFFCFFD534E2F1BC1F15FD69" box="[497,527,691,717]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">11</figureCitation>
): Cx I–IV with a row of strong blackish setae, light brown to yellowish, similar to all parts of the ventral side, surface of Cx coarse, a few irregularly brown to black spots on Cx I–IV and on Op gen, no Tu present. Op gen and free sternites with some black setae at rear margins.
</paragraph>
<paragraph id="8BB136B2FFFCFFD535D4F0101ED4FB91" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
<emphasis id="B97AEAA0FFFCFFD535D4F0101C8EFC9D" box="[199,404,799,825]" italics="true" pageId="3" pageNumber="544">Chelicera of male</emphasis>
(
<figureCitation id="13352A37FFFCFFD534B6F0101CF4FC9D" box="[421,494,799,825]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1924,1948]" captionTargetBox="[152,1436,345,1900]" captionTargetId="figure-105@4.[151,1436,345,1901]" captionTargetPageId="4" captionText="FIGURES 1–4. Nemaspela melouri sp. nov., Habitus of paratype specimens in lateral view. 1. Male (bar: 2 mm); 2. Male (bar: 1.5 mm); 3. Female CJM 8705 (bar: 1.5 mm); 4. Female CJM 8705 (bar: 1 mm). Arrow indicates male cheliceral apophysis. Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668457" httpUri="https://zenodo.org/record/4668457/files/figure.png" pageId="3" pageNumber="544">Figs 1</figureCitation>
, 
<figureCitation id="13352A37FFFCFFD534EDF0101F6FFC9D" box="[510,629,798,825]" captionStart="FIGURES 13–18" captionStartId="7.[151,264,1195,1219]" captionTargetBox="[166,1421,285,1171]" captionTargetId="figure-53@7.[151,1436,273,1172]" captionTargetPageId="7" captionText="FIGURES 13–18. Nemaspela melouri sp. nov. Chelicerae in retro-lateral view. 13. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Cheliceral apophyses in retro-lateral view. 15. The first male (bar: 0.05 mm); 16. The second male (bar: 0.05 mm). Cheliceral apophyses in pro-lateral view.17. The first male (bar: 0.05 mm); 18. The second male (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668465" httpUri="https://zenodo.org/record/4668465/files/figure.png" pageId="3" pageNumber="544">13, 15–18</figureCitation>
): Rather slender; basal article slightly constricted close to its proximal end, from there slightly widened distally, scanty bristle armament, one bristle dorsally, two bristles retro-laterally, dorsally a sub-distal small and slender frontad-directed Apo slightly surpassing the front margin of proximal article; Apo (
<figureCitation id="13352A37FFFCFFD535C5F0841C4FFC01" box="[214,341,906,933]" captionStart="FIGURES 13–18" captionStartId="7.[151,264,1195,1219]" captionTargetBox="[166,1421,285,1171]" captionTargetId="figure-53@7.[151,1436,273,1172]" captionTargetPageId="7" captionText="FIGURES 13–18. Nemaspela melouri sp. nov. Chelicerae in retro-lateral view. 13. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Cheliceral apophyses in retro-lateral view. 15. The first male (bar: 0.05 mm); 16. The second male (bar: 0.05 mm). Cheliceral apophyses in pro-lateral view.17. The first male (bar: 0.05 mm); 18. The second male (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668465" httpUri="https://zenodo.org/record/4668465/files/figure.png" pageId="3" pageNumber="544">Figs 15–18</figureCitation>
) smallest at basis, longer than broad, slightly widening distally, upper side smoothly rounded, less so lower side, and tapering to the rounded tip (in lateral view). Neither a pro-lateral excavation nor a pro-lateral tuft of setae present, but distinct sparse setation present consisting of three distal and one dorsal setae in lower half of Apo, the last one shortest. Second cheliceral article (
<figureCitation id="13352A37FFFCFFD53610F0F81E49FBB5" box="[771,851,1015,1041]" captionStart="FIGURES 13–18" captionStartId="7.[151,264,1195,1219]" captionTargetBox="[166,1421,285,1171]" captionTargetId="figure-53@7.[151,1436,273,1172]" captionTargetPageId="7" captionText="FIGURES 13–18. Nemaspela melouri sp. nov. Chelicerae in retro-lateral view. 13. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Cheliceral apophyses in retro-lateral view. 15. The first male (bar: 0.05 mm); 16. The second male (bar: 0.05 mm). Cheliceral apophyses in pro-lateral view.17. The first male (bar: 0.05 mm); 18. The second male (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668465" httpUri="https://zenodo.org/record/4668465/files/figure.png" pageId="3" pageNumber="544">Fig. 13</figureCitation>
) slender, moderately inflated, with several long and strong bristles mostly on frontal surface¸ few shorter ones ventro-distally.
</paragraph>
<paragraph id="8BB136B2FFFCFFD535D4F7301F62FB4D" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
<emphasis id="B97AEAA0FFFCFFD535D4F7301C36FBFD" box="[199,300,1087,1113]" italics="true" pageId="3" pageNumber="544">Pedipalp</emphasis>
(
<figureCitation id="13352A37FFFCFFD53429F7301C84FBFD" box="[314,414,1087,1113]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1924,1948]" captionTargetBox="[152,1436,345,1900]" captionTargetId="figure-105@4.[151,1436,345,1901]" captionTargetPageId="4" captionText="FIGURES 1–4. Nemaspela melouri sp. nov., Habitus of paratype specimens in lateral view. 1. Male (bar: 2 mm); 2. Male (bar: 1.5 mm); 3. Female CJM 8705 (bar: 1.5 mm); 4. Female CJM 8705 (bar: 1 mm). Arrow indicates male cheliceral apophysis. Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668457" httpUri="https://zenodo.org/record/4668457/files/figure.png" pageId="3" pageNumber="544">Figs 1–2</figureCitation>
, 
<figureCitation id="13352A37FFFCFFD534B9F7311CA0FBFD" box="[426,442,1086,1113]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">5</figureCitation>
): Male: Very slender and relatively long (in terms of 
<taxonomicName id="4C0E4D31FFFCFFD53114F7301999FBFD" authorityName="Silhavy" authorityYear="1966" box="[1031,1155,1087,1113]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="544" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEAA0FFFCFFD53114F7301999FBFD" box="[1031,1155,1087,1113]" italics="true" pageId="3" pageNumber="544">Nemaspela</emphasis>
</taxonomicName>
morphology), all articles parallel-sided, no article except Cx noticeably inflated; all articles except Tr bearing clavate setae, these most conspicuous on Fe ventral, on Pt mostly ventral, a few dorsal, on Ti a few at distal end ventrally and on Ta on all sides. Setae on Fe but less on Pt, on ventral side longer than breadth of article at the position. No article with special armament except setation in males and females.
</paragraph>
<paragraph id="8BB136B2FFFCFFD535D4F7FC1FC4FADD" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
<emphasis id="B97AEAA0FFFCFFD535D4F7FC1DE6FAA8" box="[199,252,1267,1292]" italics="true" pageId="3" pageNumber="544">Legs</emphasis>
(
<figureCitation id="13352A37FFFCFFD5341FF7FC1C68FAA9" box="[268,370,1267,1293]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1924,1948]" captionTargetBox="[152,1436,345,1900]" captionTargetId="figure-105@4.[151,1436,345,1901]" captionTargetPageId="4" captionText="FIGURES 1–4. Nemaspela melouri sp. nov., Habitus of paratype specimens in lateral view. 1. Male (bar: 2 mm); 2. Male (bar: 1.5 mm); 3. Female CJM 8705 (bar: 1.5 mm); 4. Female CJM 8705 (bar: 1 mm). Arrow indicates male cheliceral apophysis. Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668457" httpUri="https://zenodo.org/record/4668457/files/figure.png" pageId="3" pageNumber="544">Figs 1–2</figureCitation>
): Light brownish including Tr, extremely long and slender, thread-like, round in cross-section, armed with scattered and relatively long black setae. Many pseudo-articulations on Fe I–IV, densely packed near basis of articles, toward distal end of article distances between them increase consecutively. Number of pseudo-articulations of Fe: Fe I 7, Fe II 13, Fe III 9, Fe IV 10.
</paragraph>
<paragraph id="8BB136B2FFFCFFD535D4F68C1C61F9D1" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
<emphasis id="B97AEAA0FFFCFFD535D4F68C1CFBFA39" box="[199,481,1411,1437]" italics="true" pageId="3" pageNumber="544">Male genital morphology</emphasis>
(
<figureCitation id="13352A37FFFCFFD534FCF68C1F7EFA39" box="[495,612,1411,1437]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">Figs 7–10</figureCitation>
): Truncus penis (
<figureCitation id="13352A37FFFCFFD5360CF68C1E98FA39" box="[799,898,1411,1437]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">Figs 7–8</figureCitation>
) extremely slender; basis forming a small inflat- ed part (occupying slightly less than one fifth of whole penis length) and only inconspicuously separated from the remainder of the truncus; inflated part slender and deeply split medially, completely filled by two penial muscles, their tendons spanning the whole truncus length up to the glans. Truncus parallel-sided (dorsal/ventral view, 
<figureCitation id="13352A37FFFCFFD53063F6E01DBCF989" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">Fig. 7</figureCitation>
) along its length, slightly enlarged just below transition to the glans; in lateral view (
<figureCitation id="13352A37FFFCFFD53161F51C19ACF989" box="[1138,1206,1555,1581]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">Fig. 8</figureCitation>
) broadest above the inflated basal part, from there slightly tapering toward central section, then slightly widening toward glans, narrowest just below glans.
</paragraph>
<paragraph id="8BB136B2FFFCFFD535D4F5701999F8D5" blockId="3.[151,1437,150,2014]" pageId="3" pageNumber="544">
Glans (
<figureCitation id="13352A37FFFCFFD5340AF5701C95F93D" box="[281,399,1663,1689]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">Figs 9–10</figureCitation>
): Only inconspicuously separated from distal part of truncus outlined, short, starting where two tendons are attached to inner truncus wall; from there slightly enlarged to proximal third in dorsal/ventral view (
<figureCitation id="13352A37FFFCFFD535B3F5C81DEBF945" box="[160,241,1735,1761]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">Fig. 10</figureCitation>
) of glans, markedly broader than truncus, then continuously tapering toward stylus, stylus short, a continuation of the glans, tapering to distal rounded tip, in lateral view (
<figureCitation id="13352A37FFFCFFD5364CF5E41EB8F8A1" box="[863,930,1771,1797]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">Fig. 9</figureCitation>
) forming a continuation of distal part of truncus, but tapering slightly to the stylus. A few short, stiff and unspecialized spicules forming the armament of glans, their arrangement symmetrical in ventral, dorsal and lateral view; even smaller spicules concentrated at the basis of stylus and more densely packed than on glans proper. Measurements are given in 
<tableCitation id="C68C0309FFFCFFD5313AF4581967F8D5" box="[1065,1149,1879,1905]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Measurements of Nemaspela melouri sp. nov. Measurements are given in millimeters. Note: body length is measured in all available specimens, while length of appendages was measured only in holotype male and single female." httpUri="http://table.plazi.org/id/DF71663AFFFAFFD33584F3981DC8FF53" pageId="3" pageNumber="544" tableUuid="DF71663AFFFAFFD33584F3981DC8FF53">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BB136B2FFFCFFD235D4F4751F8CFF71" blockId="3.[151,1437,150,2014]" lastBlockId="4.[151,1437,151,322]" lastPageId="4" lastPageNumber="545" pageId="3" pageNumber="544">
<emphasis id="B97AEAA0FFFCFFD535D4F4751F19F830" bold="true" box="[199,515,1914,1941]" pageId="3" pageNumber="544">Description. Female (n=1).</emphasis>
In external appearance and size very similar to male (
<figureCitation id="13352A37FFFCFFD53141F47419ACF831" box="[1106,1206,1915,1941]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1924,1948]" captionTargetBox="[152,1436,345,1900]" captionTargetId="figure-105@4.[151,1436,345,1901]" captionTargetPageId="4" captionText="FIGURES 1–4. Nemaspela melouri sp. nov., Habitus of paratype specimens in lateral view. 1. Male (bar: 2 mm); 2. Male (bar: 1.5 mm); 3. Female CJM 8705 (bar: 1.5 mm); 4. Female CJM 8705 (bar: 1 mm). Arrow indicates male cheliceral apophysis. Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668457" httpUri="https://zenodo.org/record/4668457/files/figure.png" pageId="3" pageNumber="544">Figs 3–4</figureCitation>
, 
<figureCitation id="13352A37FFFCFFD531D3F47419C5F831" box="[1216,1247,1915,1941]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">12</figureCitation>
). 
<emphasis id="B97AEAA0FFFCFFD531E1F474187AF831" box="[1266,1376,1915,1941]" italics="true" pageId="3" pageNumber="544">Chelicera</emphasis>
(
<figureCitation id="13352A37FFFCFFD5307CF4741DAEF81D" captionStart="FIGURES 13–18" captionStartId="7.[151,264,1195,1219]" captionTargetBox="[166,1421,285,1171]" captionTargetId="figure-53@7.[151,1436,273,1172]" captionTargetPageId="7" captionText="FIGURES 13–18. Nemaspela melouri sp. nov. Chelicerae in retro-lateral view. 13. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Cheliceral apophyses in retro-lateral view. 15. The first male (bar: 0.05 mm); 16. The second male (bar: 0.05 mm). Cheliceral apophyses in pro-lateral view.17. The first male (bar: 0.05 mm); 18. The second male (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668465" httpUri="https://zenodo.org/record/4668465/files/figure.png" pageId="3" pageNumber="544">Fig. 14</figureCitation>
) similar to male, but dorso-distal Apo on basal article absent, 2 
<superScript id="7C7B9BFAFFFCFFD53664F4911E9DF808" attach="left" box="[887,903,1950,1964]" fontSize="6" pageId="3" pageNumber="544">nd</superScript>
article more stout and bristles markedly longer. 
<emphasis id="B97AEAA0FFFCFFD53584F4CC1DE6F879" box="[151,252,1987,2013]" italics="true" pageId="3" pageNumber="544">Pedipalp</emphasis>
(
<figureCitation id="13352A37FFFCFFD53418F4CC1C6BF879" box="[267,369,1987,2013]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1924,1948]" captionTargetBox="[152,1436,345,1900]" captionTargetId="figure-105@4.[151,1436,345,1901]" captionTargetPageId="4" captionText="FIGURES 1–4. Nemaspela melouri sp. nov., Habitus of paratype specimens in lateral view. 1. Male (bar: 2 mm); 2. Male (bar: 1.5 mm); 3. Female CJM 8705 (bar: 1.5 mm); 4. Female CJM 8705 (bar: 1 mm). Arrow indicates male cheliceral apophysis. Photographs by J. Severin." figureDoi="http://doi.org/10.5281/zenodo.4668457" httpUri="https://zenodo.org/record/4668457/files/figure.png" pageId="3" pageNumber="544">Figs 3–4</figureCitation>
, 
<figureCitation id="13352A37FFFCFFD5346EF4CC1C96F87A" box="[381,396,1987,2014]" captionStart="FIGURES 5–12" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[194,1389,197,1903]" captionTargetId="figure-23@6.[180,1408,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–12. Nemaspela melouri sp. nov. Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="3" pageNumber="544">6</figureCitation>
) very similar to male including setation, size nearly identical, only palpal Fe slightly shorter than in male. Number of pseudo-articulations of Fe of legs slightly differing from male: Fe I 10, Fe II 14, Fe III 8, Fe IV 11. Measurements are given in 
<tableCitation id="C68C0309FFFBFFD2372FF3B41F8AFF71" box="[572,656,187,213]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Measurements of Nemaspela melouri sp. nov. Measurements are given in millimeters. Note: body length is measured in all available specimens, while length of appendages was measured only in holotype male and single female." httpUri="http://table.plazi.org/id/DF71663AFFFAFFD33584F3981DC8FF53" pageId="4" pageNumber="545" tableUuid="DF71663AFFFAFFD33584F3981DC8FF53">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BB136B2FFFBFFD235D4F3D11955FEE5" blockId="4.[151,1437,151,322]" pageId="4" pageNumber="545">
<emphasis id="B97AEAA0FFFBFFD235D4F3D11C25FF5C" bold="true" box="[199,319,222,248]" pageId="4" pageNumber="545">Variation.</emphasis>
Apparently very little. Outline of the Apo of the male basal cheliceral article in the two specimens examined is nearly identical, even the setation, with two distal setae retro-laterally, one dorsally, and two (a longer one distally and a shorter one more basally) at nearly identical position (
<figureCitation id="13352A37FFFBFFD236D3F2281958FEE5" box="[960,1090,294,321]" captionStart="FIGURES 13–18" captionStartId="7.[151,264,1195,1219]" captionTargetBox="[166,1421,285,1171]" captionTargetId="figure-53@7.[151,1436,273,1172]" captionTargetPageId="7" captionText="FIGURES 13–18. Nemaspela melouri sp. nov. Chelicerae in retro-lateral view. 13. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Cheliceral apophyses in retro-lateral view. 15. The first male (bar: 0.05 mm); 16. The second male (bar: 0.05 mm). Cheliceral apophyses in pro-lateral view.17. The first male (bar: 0.05 mm); 18. The second male (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.4668465" httpUri="https://zenodo.org/record/4668465/files/figure.png" pageId="4" pageNumber="545">Figs 15–18</figureCitation>
).
</paragraph>
<caption id="DF71663AFFFBFFD23584F48B1CBDF840" ID-DOI="http://doi.org/10.5281/zenodo.4668457" ID-Zenodo-Dep="4668457" httpUri="https://zenodo.org/record/4668457/files/figure.png" pageId="4" pageNumber="545" startId="4.[151,264,1924,1948]" targetBox="[152,1436,345,1900]" targetPageId="4">
<paragraph id="8BB136B2FFFBFFD23584F48B1CBDF840" blockId="4.[151,1437,1924,2020]" pageId="4" pageNumber="545">
<emphasis id="B97AEAA0FFFBFFD23584F48B1C20F838" bold="true" box="[151,314,1924,1948]" pageId="4" pageNumber="545">FIGURES 1–4.</emphasis>
<taxonomicName id="4C0E4D31FFFBFFD23453F48B1F1CF838" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[320,518,1924,1948]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="545" phylum="Arthropoda" rank="species" species="melouri" status="sp. nov.">
<emphasis id="B97AEAA0FFFBFFD23453F48B1F1CF838" box="[320,518,1924,1948]" italics="true" pageId="4" pageNumber="545">Nemaspela melouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957DBFFFBFFD23718F48B1F47F838" box="[523,605,1924,1948]" pageId="4" pageNumber="545" rank="species">
<emphasis id="B97AEAA0FFFBFFD23718F48B1F4DF838" bold="true" box="[523,599,1924,1948]" pageId="4" pageNumber="545">sp. nov</emphasis>
.
</taxonomicNameLabel>
, Habitus of paratype specimens in lateral view. 1. Male (bar: 2 mm); 2. Male (bar: 1.5 mm); 3. Female CJM 8705 (bar: 1.5 mm); 4. Female CJM 8705 (bar: 1 mm). Arrow indicates male cheliceral apophysis. Photographs by J. Severin.
</paragraph>
</caption>
<caption id="DF71663AFFFAFFD33584F3981DC8FF53" ID-Table-UUID="DF71663AFFFAFFD33584F3981DC8FF53" httpUri="http://table.plazi.org/id/DF71663AFFFAFFD33584F3981DC8FF53" pageId="5" pageNumber="546" startId="5.[151,239,151,175]" targetBox="[166,1291,264,1936]" targetIsTable="true" targetPageId="5">
<paragraph id="8BB136B2FFFAFFD33584F3981DC8FF53" blockId="5.[151,1437,151,247]" pageId="5" pageNumber="546">
<emphasis id="B97AEAA0FFFAFFD33584F3981C17FF0B" bold="true" box="[151,269,151,175]" pageId="5" pageNumber="546">TABLE 1.</emphasis>
Measurements of 
<taxonomicName id="4C0E4D31FFFAFFD334CFF3981FB5FF0B" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[476,687,151,175]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="546" phylum="Arthropoda" rank="species" species="melouri" status="sp. nov.">
<emphasis id="B97AEAA0FFFAFFD334CFF3981FB5FF0B" box="[476,687,151,175]" italics="true" pageId="5" pageNumber="546">Nemaspela melouri</emphasis>
</taxonomicName>
<emphasis id="B97AEAA0FFFAFFD337ABF3971E0BFF14" bold="true" box="[696,785,152,176]" pageId="5" pageNumber="546">
<taxonomicNameLabel id="A24957DBFFFAFFD337ABF3971E0BFF14" box="[696,785,152,176]" pageId="5" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Measurements are given in millimeters. Note: body length is measured in all available specimens, while length of appendages was measured only in holotype male and single female.
</paragraph>
</caption>
<paragraph id="8BB136B2FFFAFFD335B5F20719D7F82B" pageId="5" pageNumber="546">
<table id="F90EC412FFFA002935B5F2071811F834" box="[166,1291,264,1936]" gridcols="3" gridrows="42" pageId="5" pageNumber="546">
<tr id="353E34F0FFFA002935B5F2071811FE84" box="[166,1291,264,288]" gridrow="0" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F2071C8BFE84" box="[166,401,264,288]" gridcol="0" gridrow="0" pageId="5" pageNumber="546">Character</th>
<th id="76EF5D8CFFFA002937CFF2071E42FE84" box="[732,856,264,288]" gridcol="1" gridrow="0" pageId="5" pageNumber="546">Males (n=4)</th>
<th id="76EF5D8CFFFA00293192F2071811FE84" box="[1153,1291,264,288]" gridcol="2" gridrow="0" pageId="5" pageNumber="546">Female (n=1)</th>
</tr>
<tr id="353E34F0FFFA002935B5F23F1811FEEC" box="[166,1291,304,328]" gridrow="1" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F23F1C8BFEEC" box="[166,401,304,328]" gridcol="0" gridrow="1" pageId="5" pageNumber="546">Body length</th>
<td id="76EF5D8CFFFA002937CFF23F1E42FEEC" box="[732,856,304,328]" gridcol="1" gridrow="1" pageId="5" pageNumber="546">1.2–1.6</td>
<td id="76EF5D8CFFFA00293192F23F1811FEEC" box="[1153,1291,304,328]" gridcol="2" gridrow="1" pageId="5" pageNumber="546">1.5</td>
</tr>
<tr id="353E34F0FFFA002935B5F2571811FED4" box="[166,1291,344,368]" gridrow="2" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F2571811FED4" box="[166,1291,344,368]" colspan="3" colspanRight="2" gridcol="0" gridrow="2" pageId="5" pageNumber="546">
<emphasis id="B97AEAA0FFFAFFD335B5F2571C14FED4" bold="true" box="[166,270,344,368]" pageId="5" pageNumber="546">Pedipalp:</emphasis>
</th>
</tr>
<tr id="353E34F0FFFA002935B5F28F1811FE3C" box="[166,1291,384,408]" gridrow="3" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F28F1C8BFE3C" box="[166,401,384,408]" gridcol="0" gridrow="3" pageId="5" pageNumber="546">Trochanter</th>
<td id="76EF5D8CFFFA002937CFF28F1E42FE3C" box="[732,856,384,408]" gridcol="1" gridrow="3" pageId="5" pageNumber="546">0.5</td>
<td id="76EF5D8CFFFA00293192F28F1811FE3C" box="[1153,1291,384,408]" gridcol="2" gridrow="3" pageId="5" pageNumber="546">0.4</td>
</tr>
<tr id="353E34F0FFFA002935B5F2A71811FE64" box="[166,1291,424,448]" gridrow="4" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F2A71C8BFE64" box="[166,401,424,448]" gridcol="0" gridrow="4" pageId="5" pageNumber="546">Femur</th>
<td id="76EF5D8CFFFA002937CFF2A71E42FE64" box="[732,856,424,448]" gridcol="1" gridrow="4" pageId="5" pageNumber="546">1.4</td>
<td id="76EF5D8CFFFA00293192F2A71811FE64" box="[1153,1291,424,448]" gridcol="2" gridrow="4" pageId="5" pageNumber="546">1.35</td>
</tr>
<tr id="353E34F0FFFA002935B5F2DF1811FE4D" box="[166,1291,464,489]" gridrow="5" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F2DF1C8BFE4D" box="[166,401,464,489]" gridcol="0" gridrow="5" pageId="5" pageNumber="546">Patella</th>
<td id="76EF5D8CFFFA002937CFF2DF1E42FE4D" box="[732,856,464,489]" gridcol="1" gridrow="5" pageId="5" pageNumber="546">1.5</td>
<td id="76EF5D8CFFFA00293192F2DF1811FE4D" box="[1153,1291,464,489]" gridcol="2" gridrow="5" pageId="5" pageNumber="546">1.4</td>
</tr>
<tr id="353E34F0FFFA002935B5F2F61811FDB5" box="[166,1291,505,529]" gridrow="6" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F2F61C8BFDB5" box="[166,401,505,529]" gridcol="0" gridrow="6" pageId="5" pageNumber="546">Tibia</th>
<td id="76EF5D8CFFFA002937CFF2F61E42FDB5" box="[732,856,505,529]" gridcol="1" gridrow="6" pageId="5" pageNumber="546">0.95</td>
<td id="76EF5D8CFFFA00293192F2F61811FDB5" box="[1153,1291,505,529]" gridcol="2" gridrow="6" pageId="5" pageNumber="546">0.9</td>
</tr>
<tr id="353E34F0FFFA002935B5F12E1811FD9D" box="[166,1291,545,569]" gridrow="7" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F12E1C8BFD9D" box="[166,401,545,569]" gridcol="0" gridrow="7" pageId="5" pageNumber="546">Tarsus</th>
<td id="76EF5D8CFFFA002937CFF12E1E42FD9D" box="[732,856,545,569]" gridcol="1" gridrow="7" pageId="5" pageNumber="546">0.5</td>
<td id="76EF5D8CFFFA00293192F12E1811FD9D" box="[1153,1291,545,569]" gridcol="2" gridrow="7" pageId="5" pageNumber="546">0.5</td>
</tr>
<tr id="353E34F0FFFA002935B5F1461811FDC5" box="[166,1291,585,609]" gridrow="8" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F1461C8BFDC5" box="[166,401,585,609]" gridcol="0" gridrow="8" pageId="5" pageNumber="546">Pedipalp length in total</th>
<td id="76EF5D8CFFFA002937CFF1461E42FDC5" box="[732,856,585,609]" gridcol="1" gridrow="8" pageId="5" pageNumber="546">4.85</td>
<td id="76EF5D8CFFFA00293192F1461811FDC5" box="[1153,1291,585,609]" gridcol="2" gridrow="8" pageId="5" pageNumber="546">4.55</td>
</tr>
<tr id="353E34F0FFFA002935B5F17E1811FD2D" box="[166,1291,625,649]" gridrow="9" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F17E1811FD2D" box="[166,1291,625,649]" colspan="3" colspanRight="2" gridcol="0" gridrow="9" pageId="5" pageNumber="546">
<emphasis id="B97AEAA0FFFAFFD335B5F17E1DC7FD2D" bold="true" box="[166,221,625,649]" pageId="5" pageNumber="546">I leg:</emphasis>
</th>
</tr>
<tr id="353E34F0FFFA002935B5F1961811FD15" box="[166,1291,665,689]" gridrow="10" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F1961C8BFD15" box="[166,401,665,689]" gridcol="0" gridrow="10" pageId="5" pageNumber="546">Trochanter</th>
<td id="76EF5D8CFFFA002937CFF1961E42FD15" box="[732,856,665,689]" gridcol="1" gridrow="10" pageId="5" pageNumber="546">0.3</td>
<td id="76EF5D8CFFFA00293192F1961811FD15" box="[1153,1291,665,689]" gridcol="2" gridrow="10" pageId="5" pageNumber="546">0.2</td>
</tr>
<tr id="353E34F0FFFA002935B5F1CD1811FD7E" box="[166,1291,706,730]" gridrow="11" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F1CD1C8BFD7E" box="[166,401,706,730]" gridcol="0" gridrow="11" pageId="5" pageNumber="546">Femur</th>
<td id="76EF5D8CFFFA002937CFF1CD1E42FD7E" box="[732,856,706,730]" gridcol="1" gridrow="11" pageId="5" pageNumber="546">3.0</td>
<td id="76EF5D8CFFFA00293192F1CD1811FD7E" box="[1153,1291,706,730]" gridcol="2" gridrow="11" pageId="5" pageNumber="546">2.7</td>
</tr>
<tr id="353E34F0FFFA002935B5F1E51811FCA6" box="[166,1291,746,770]" gridrow="12" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F1E51C8BFCA6" box="[166,401,746,770]" gridcol="0" gridrow="12" pageId="5" pageNumber="546">Patella</th>
<td id="76EF5D8CFFFA002937CFF1E51E42FCA6" box="[732,856,746,770]" gridcol="1" gridrow="12" pageId="5" pageNumber="546">0.5</td>
<td id="76EF5D8CFFFA00293192F1E51811FCA6" box="[1153,1291,746,770]" gridcol="2" gridrow="12" pageId="5" pageNumber="546">0.5</td>
</tr>
<tr id="353E34F0FFFA002935B5F01D1811FC8E" box="[166,1291,786,810]" gridrow="13" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F01D1C8BFC8E" box="[166,401,786,810]" gridcol="0" gridrow="13" pageId="5" pageNumber="546">Tibia</th>
<td id="76EF5D8CFFFA002937CFF01D1E42FC8E" box="[732,856,786,810]" gridcol="1" gridrow="13" pageId="5" pageNumber="546">2.3</td>
<td id="76EF5D8CFFFA00293192F01D1811FC8E" box="[1153,1291,786,810]" gridcol="2" gridrow="13" pageId="5" pageNumber="546">2.0</td>
</tr>
<tr id="353E34F0FFFA002935B5F0351811FCF6" box="[166,1291,826,850]" gridrow="14" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F0351C8BFCF6" box="[166,401,826,850]" gridcol="0" gridrow="14" pageId="5" pageNumber="546">Metatarsus</th>
<td id="76EF5D8CFFFA002937CFF0351E42FCF6" box="[732,856,826,850]" gridcol="1" gridrow="14" pageId="5" pageNumber="546">4.2</td>
<td id="76EF5D8CFFFA00293192F0351811FCF6" box="[1153,1291,826,850]" gridcol="2" gridrow="14" pageId="5" pageNumber="546">4.0</td>
</tr>
<tr id="353E34F0FFFA002935B5F06C1811FCDF" box="[166,1291,867,891]" gridrow="15" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F06C1C8BFCDF" box="[166,401,867,891]" gridcol="0" gridrow="15" pageId="5" pageNumber="546">Tarsus</th>
<td id="76EF5D8CFFFA002937CFF06C1E42FCDF" box="[732,856,867,891]" gridcol="1" gridrow="15" pageId="5" pageNumber="546">2.2</td>
<td id="76EF5D8CFFFA00293192F06C1811FCDF" box="[1153,1291,867,891]" gridcol="2" gridrow="15" pageId="5" pageNumber="546">2.0</td>
</tr>
<tr id="353E34F0FFFA002935B5F0841811FC07" box="[166,1291,907,931]" gridrow="16" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F0841C8BFC07" box="[166,401,907,931]" gridcol="0" gridrow="16" pageId="5" pageNumber="546">I leg length in total</th>
<td id="76EF5D8CFFFA002937CFF0841E42FC07" box="[732,856,907,931]" gridcol="1" gridrow="16" pageId="5" pageNumber="546">12.5</td>
<td id="76EF5D8CFFFA00293192F0841811FC07" box="[1153,1291,907,931]" gridcol="2" gridrow="16" pageId="5" pageNumber="546">11.4</td>
</tr>
<tr id="353E34F0FFFA002935B5F0BC1811FC6F" box="[166,1291,947,971]" gridrow="17" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F0BC1811FC6F" box="[166,1291,947,971]" colspan="3" colspanRight="2" gridcol="0" gridrow="17" pageId="5" pageNumber="546">
<emphasis id="B97AEAA0FFFAFFD335B5F0BC1DFDFC6F" bold="true" box="[166,231,947,971]" pageId="5" pageNumber="546">II leg:</emphasis>
</th>
</tr>
<tr id="353E34F0FFFA002935B5F0D41811FC57" box="[166,1291,987,1011]" gridrow="18" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F0D41C8BFC57" box="[166,401,987,1011]" gridcol="0" gridrow="18" pageId="5" pageNumber="546">Trochanter</th>
<td id="76EF5D8CFFFA002937CFF0D41E42FC57" box="[732,856,987,1011]" gridcol="1" gridrow="18" pageId="5" pageNumber="546">0.3</td>
<td id="76EF5D8CFFFA00293192F0D41811FC57" box="[1153,1291,987,1011]" gridcol="2" gridrow="18" pageId="5" pageNumber="546">0.3</td>
</tr>
<tr id="353E34F0FFFA002935B5F70C1811FBBF" box="[166,1291,1027,1051]" gridrow="19" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F70C1C8BFBBF" box="[166,401,1027,1051]" gridcol="0" gridrow="19" pageId="5" pageNumber="546">Femur</th>
<td id="76EF5D8CFFFA002937CFF70C1E42FBBF" box="[732,856,1027,1051]" gridcol="1" gridrow="19" pageId="5" pageNumber="546">5.8</td>
<td id="76EF5D8CFFFA00293192F70C1811FBBF" box="[1153,1291,1027,1051]" gridcol="2" gridrow="19" pageId="5" pageNumber="546">5.5</td>
</tr>
<tr id="353E34F0FFFA002935B5F7241811FBE7" box="[166,1291,1067,1091]" gridrow="20" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F7241C8BFBE7" box="[166,401,1067,1091]" gridcol="0" gridrow="20" pageId="5" pageNumber="546">Patella</th>
<td id="76EF5D8CFFFA002937CFF7241E42FBE7" box="[732,856,1067,1091]" gridcol="1" gridrow="20" pageId="5" pageNumber="546">0.6</td>
<td id="76EF5D8CFFFA00293192F7241811FBE7" box="[1153,1291,1067,1091]" gridcol="2" gridrow="20" pageId="5" pageNumber="546">0.6</td>
</tr>
<tr id="353E34F0FFFA002935B5F75C1811FBC8" box="[166,1291,1107,1132]" gridrow="21" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F75C1C8BFBC8" box="[166,401,1107,1132]" gridcol="0" gridrow="21" pageId="5" pageNumber="546">Tibia</th>
<td id="76EF5D8CFFFA002937CFF75C1E42FBC8" box="[732,856,1107,1132]" gridcol="1" gridrow="21" pageId="5" pageNumber="546">5.3</td>
<td id="76EF5D8CFFFA00293192F75C1811FBC8" box="[1153,1291,1107,1132]" gridcol="2" gridrow="21" pageId="5" pageNumber="546">4.7</td>
</tr>
<tr id="353E34F0FFFA002935B5F7731811FB30" box="[166,1291,1148,1172]" gridrow="22" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F7731C8BFB30" box="[166,401,1148,1172]" gridcol="0" gridrow="22" pageId="5" pageNumber="546">Metatarsus</th>
<td id="76EF5D8CFFFA002937CFF7731E42FB30" box="[732,856,1148,1172]" gridcol="1" gridrow="22" pageId="5" pageNumber="546">8.7</td>
<td id="76EF5D8CFFFA00293192F7731811FB30" box="[1153,1291,1148,1172]" gridcol="2" gridrow="22" pageId="5" pageNumber="546">8.0</td>
</tr>
<tr id="353E34F0FFFA002935B5F7AB1811FB18" box="[166,1291,1188,1212]" gridrow="23" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F7AB1C8BFB18" box="[166,401,1188,1212]" gridcol="0" gridrow="23" pageId="5" pageNumber="546">Tarsus</th>
<td id="76EF5D8CFFFA002937CFF7AB1E42FB18" box="[732,856,1188,1212]" gridcol="1" gridrow="23" pageId="5" pageNumber="546">4.8</td>
<td id="76EF5D8CFFFA00293192F7AB1811FB18" box="[1153,1291,1188,1212]" gridcol="2" gridrow="23" pageId="5" pageNumber="546">4.5</td>
</tr>
<tr id="353E34F0FFFA002935B5F7C31811FB40" box="[166,1291,1228,1252]" gridrow="24" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F7C31C8BFB40" box="[166,401,1228,1252]" gridcol="0" gridrow="24" pageId="5" pageNumber="546">II leg length in total</th>
<td id="76EF5D8CFFFA002937CFF7C31E42FB40" box="[732,856,1228,1252]" gridcol="1" gridrow="24" pageId="5" pageNumber="546">25.5</td>
<td id="76EF5D8CFFFA00293192F7C31811FB40" box="[1153,1291,1228,1252]" gridcol="2" gridrow="24" pageId="5" pageNumber="546">23.6</td>
</tr>
<tr id="353E34F0FFFA002935B5F7FB1811FAA8" box="[166,1291,1268,1292]" gridrow="25" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F7FB1811FAA8" box="[166,1291,1268,1292]" colspan="3" colspanRight="2" gridcol="0" gridrow="25" pageId="5" pageNumber="546">
<emphasis id="B97AEAA0FFFAFFD335B5F7FB1DEBFAA8" bold="true" box="[166,241,1268,1292]" pageId="5" pageNumber="546">III leg:</emphasis>
</th>
</tr>
<tr id="353E34F0FFFA002935B5F6131811FA91" box="[166,1291,1308,1333]" gridrow="26" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F6131C8BFA91" box="[166,401,1308,1333]" gridcol="0" gridrow="26" pageId="5" pageNumber="546">Trochanter</th>
<td id="76EF5D8CFFFA002937CFF6131E42FA91" box="[732,856,1308,1333]" gridcol="1" gridrow="26" pageId="5" pageNumber="546">0.2</td>
<td id="76EF5D8CFFFA00293192F6131811FA91" box="[1153,1291,1308,1333]" gridcol="2" gridrow="26" pageId="5" pageNumber="546">0.2</td>
</tr>
<tr id="353E34F0FFFA002935B5F64A1811FAF9" box="[166,1291,1349,1373]" gridrow="27" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F64A1C8BFAF9" box="[166,401,1349,1373]" gridcol="0" gridrow="27" pageId="5" pageNumber="546">Femur</th>
<td id="76EF5D8CFFFA002937CFF64A1E42FAF9" box="[732,856,1349,1373]" gridcol="1" gridrow="27" pageId="5" pageNumber="546">3.0</td>
<td id="76EF5D8CFFFA00293192F64A1811FAF9" box="[1153,1291,1349,1373]" gridcol="2" gridrow="27" pageId="5" pageNumber="546">2.8</td>
</tr>
<tr id="353E34F0FFFA002935B5F6621811FA21" box="[166,1291,1389,1413]" gridrow="28" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F6621C8BFA21" box="[166,401,1389,1413]" gridcol="0" gridrow="28" pageId="5" pageNumber="546">Patella</th>
<td id="76EF5D8CFFFA002937CFF6621E42FA21" box="[732,856,1389,1413]" gridcol="1" gridrow="28" pageId="5" pageNumber="546">0.5</td>
<td id="76EF5D8CFFFA00293192F6621811FA21" box="[1153,1291,1389,1413]" gridcol="2" gridrow="28" pageId="5" pageNumber="546">0.5</td>
</tr>
<tr id="353E34F0FFFA002935B5F69A1811FA09" box="[166,1291,1429,1453]" gridrow="29" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F69A1C8BFA09" box="[166,401,1429,1453]" gridcol="0" gridrow="29" pageId="5" pageNumber="546">Tibia</th>
<td id="76EF5D8CFFFA002937CFF69A1E42FA09" box="[732,856,1429,1453]" gridcol="1" gridrow="29" pageId="5" pageNumber="546">2.1</td>
<td id="76EF5D8CFFFA00293192F69A1811FA09" box="[1153,1291,1429,1453]" gridcol="2" gridrow="29" pageId="5" pageNumber="546">2.9</td>
</tr>
<tr id="353E34F0FFFA002935B5F6B21811FA72" box="[166,1291,1469,1494]" gridrow="30" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F6B21C8BFA72" box="[166,401,1469,1494]" gridcol="0" gridrow="30" pageId="5" pageNumber="546">Metatarsus</th>
<td id="76EF5D8CFFFA002937CFF6B21E42FA72" box="[732,856,1469,1494]" gridcol="1" gridrow="30" pageId="5" pageNumber="546">4.5</td>
<td id="76EF5D8CFFFA00293192F6B21811FA72" box="[1153,1291,1469,1494]" gridcol="2" gridrow="30" pageId="5" pageNumber="546">4.3</td>
</tr>
<tr id="353E34F0FFFA002935B5F6EA1811FA5A" box="[166,1291,1509,1534]" gridrow="31" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F6EA1C8BFA5A" box="[166,401,1509,1534]" gridcol="0" gridrow="31" pageId="5" pageNumber="546">Tarsus</th>
<td id="76EF5D8CFFFA002937CFF6EA1E42FA5A" box="[732,856,1509,1534]" gridcol="1" gridrow="31" pageId="5" pageNumber="546">2.5</td>
<td id="76EF5D8CFFFA00293192F6EA1811FA5A" box="[1153,1291,1509,1534]" gridcol="2" gridrow="31" pageId="5" pageNumber="546">2.1</td>
</tr>
<tr id="353E34F0FFFA002935B5F5011811F982" box="[166,1291,1550,1574]" gridrow="32" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F5011C8BF982" box="[166,401,1550,1574]" gridcol="0" gridrow="32" pageId="5" pageNumber="546">III leg length in total</th>
<td id="76EF5D8CFFFA002937CFF5011E42F982" box="[732,856,1550,1574]" gridcol="1" gridrow="32" pageId="5" pageNumber="546">12.8</td>
<td id="76EF5D8CFFFA00293192F5011811F982" box="[1153,1291,1550,1574]" gridcol="2" gridrow="32" pageId="5" pageNumber="546">12.8</td>
</tr>
<tr id="353E34F0FFFA002935B5F5391811F9EA" box="[166,1291,1590,1614]" gridrow="33" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F5391811F9EA" box="[166,1291,1590,1614]" colspan="3" colspanRight="2" gridcol="0" gridrow="33" pageId="5" pageNumber="546">
<emphasis id="B97AEAA0FFFAFFD335B5F5391DF5F9EA" bold="true" box="[166,239,1590,1614]" pageId="5" pageNumber="546">IV leg:</emphasis>
</th>
</tr>
<tr id="353E34F0FFFA002935B5F5511811F9D2" box="[166,1291,1630,1654]" gridrow="34" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F5511C8BF9D2" box="[166,401,1630,1654]" gridcol="0" gridrow="34" pageId="5" pageNumber="546">Trochanter</th>
<td id="76EF5D8CFFFA002937CFF5511E42F9D2" box="[732,856,1630,1654]" gridcol="1" gridrow="34" pageId="5" pageNumber="546">0.3</td>
<td id="76EF5D8CFFFA00293192F5511811F9D2" box="[1153,1291,1630,1654]" gridcol="2" gridrow="34" pageId="5" pageNumber="546">0.3</td>
</tr>
<tr id="353E34F0FFFA002935B5F5891811F93A" box="[166,1291,1670,1694]" gridrow="35" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F5891C8BF93A" box="[166,401,1670,1694]" gridcol="0" gridrow="35" pageId="5" pageNumber="546">Femur</th>
<td id="76EF5D8CFFFA002937CFF5891E42F93A" box="[732,856,1670,1694]" gridcol="1" gridrow="35" pageId="5" pageNumber="546">4.4</td>
<td id="76EF5D8CFFFA00293192F5891811F93A" box="[1153,1291,1670,1694]" gridcol="2" gridrow="35" pageId="5" pageNumber="546">4.5</td>
</tr>
<tr id="353E34F0FFFA002935B5F5A11811F962" box="[166,1291,1710,1734]" gridrow="36" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F5A11C8BF962" box="[166,401,1710,1734]" gridcol="0" gridrow="36" pageId="5" pageNumber="546">Patella</th>
<td id="76EF5D8CFFFA002937CFF5A11E42F962" box="[732,856,1710,1734]" gridcol="1" gridrow="36" pageId="5" pageNumber="546">0.6</td>
<td id="76EF5D8CFFFA00293192F5A11811F962" box="[1153,1291,1710,1734]" gridcol="2" gridrow="36" pageId="5" pageNumber="546">0.6</td>
</tr>
<tr id="353E34F0FFFA002935B5F5D81811F94B" box="[166,1291,1751,1775]" gridrow="37" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F5D81C8BF94B" box="[166,401,1751,1775]" gridcol="0" gridrow="37" pageId="5" pageNumber="546">Tibia</th>
<td id="76EF5D8CFFFA002937CFF5D81E42F94B" box="[732,856,1751,1775]" gridcol="1" gridrow="37" pageId="5" pageNumber="546">3.1</td>
<td id="76EF5D8CFFFA00293192F5D81811F94B" box="[1153,1291,1751,1775]" gridcol="2" gridrow="37" pageId="5" pageNumber="546">3.0</td>
</tr>
<tr id="353E34F0FFFA002935B5F5F01811F8B3" box="[166,1291,1791,1815]" gridrow="38" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F5F01C8BF8B3" box="[166,401,1791,1815]" gridcol="0" gridrow="38" pageId="5" pageNumber="546">Metatarsus</th>
<td id="76EF5D8CFFFA002937CFF5F01E42F8B3" box="[732,856,1791,1815]" gridcol="1" gridrow="38" pageId="5" pageNumber="546">6.4</td>
<td id="76EF5D8CFFFA00293192F5F01811F8B3" box="[1153,1291,1791,1815]" gridcol="2" gridrow="38" pageId="5" pageNumber="546">6.4</td>
</tr>
<tr id="353E34F0FFFA002935B5F4281811F89B" box="[166,1291,1831,1855]" gridrow="39" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F4281C8BF89B" box="[166,401,1831,1855]" gridcol="0" gridrow="39" pageId="5" pageNumber="546">Tarsus</th>
<td id="76EF5D8CFFFA002937CFF4281E42F89B" box="[732,856,1831,1855]" gridcol="1" gridrow="39" pageId="5" pageNumber="546">2.7</td>
<td id="76EF5D8CFFFA00293192F4281811F89B" box="[1153,1291,1831,1855]" gridcol="2" gridrow="39" pageId="5" pageNumber="546">2.7</td>
</tr>
<tr id="353E34F0FFFA002935B5F4401811F8C3" box="[166,1291,1871,1895]" gridrow="40" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F4401C8BF8C3" box="[166,401,1871,1895]" gridcol="0" gridrow="40" pageId="5" pageNumber="546">IV leg length in total</th>
<td id="76EF5D8CFFFA002937CFF4401E42F8C3" box="[732,856,1871,1895]" gridcol="1" gridrow="40" pageId="5" pageNumber="546">17.5</td>
<td id="76EF5D8CFFFA00293192F4401811F8C3" box="[1153,1291,1871,1895]" gridcol="2" gridrow="40" pageId="5" pageNumber="546">17.5</td>
</tr>
<tr id="353E34F0FFFA002935B5F4771811F834" box="[166,1291,1912,1936]" gridrow="41" pageId="5" pageNumber="546">
<th id="76EF5D8CFFFA002935B5F4771C8BF834" box="[166,401,1912,1936]" gridcol="0" gridrow="41" pageId="5" pageNumber="546">Penis length</th>
<td id="76EF5D8CFFFA002937CFF4771E42F834" box="[732,856,1912,1936]" gridcol="1" gridrow="41" pageId="5" pageNumber="546">1.0</td>
<td id="76EF5D8CFFFA00293192F4771811F834" box="[1153,1291,1912,1936]" gridcol="2" gridrow="41" pageId="5" pageNumber="546">–</td>
</tr>
</table>
</paragraph>
<caption id="DF71663AFFF9FFD03584F4981972F85C" ID-DOI="http://doi.org/10.5281/zenodo.4668461" ID-Zenodo-Dep="4668461" httpUri="https://zenodo.org/record/4668461/files/figure.png" pageId="6" pageNumber="547" startId="6.[151,264,1943,1967]" targetBox="[194,1389,197,1903]" targetPageId="6">
<paragraph id="8BB136B2FFF9FFD03584F4981972F85C" blockId="6.[151,1437,1943,2040]" pageId="6" pageNumber="547">
<emphasis id="B97AEAA0FFF9FFD03584F4981C52F80B" bold="true" box="[151,328,1943,1967]" pageId="6" pageNumber="547">FIGURES 5–12.</emphasis>
<taxonomicName id="4C0E4D31FFF9FFD0345DF4971F0EF814" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[334,532,1944,1968]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="547" phylum="Arthropoda" rank="species" species="melouri" status="sp. nov.">
<emphasis id="B97AEAA0FFF9FFD0345DF4971F0EF814" box="[334,532,1944,1968]" italics="true" pageId="6" pageNumber="547">Nemaspela melouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957DBFFF9FFD03709F4971F76F80B" box="[538,620,1944,1968]" pageId="6" pageNumber="547" rank="species">
<emphasis id="B97AEAA0FFF9FFD03709F4971F7CF814" bold="true" box="[538,614,1944,1968]" pageId="6" pageNumber="547">sp. nov</emphasis>
.
</taxonomicNameLabel>
Pedipalps in retro-lateral view. 5. Male (bar: 0.5 mm); 6. Female (bar: 0.5 mm). Penis. 7. Ventral view (bar: 0.2 mm); 8. Lateral view (bar: 0.2 mm). Glans penis. 9. Lateral view (bar: 0.05 mm); 10. Ventral view (bar: 0.05 mm). Habitus in lateral view. 11. Male (bar: 0.5 mm); 12. Female (bar: 0.5 mm).
</paragraph>
</caption>
<paragraph id="8BB136B2FFF8FFD135D4F3991F33FF15" blockId="7.[151,1436,150,249]" box="[199,553,150,177]" pageId="7" pageNumber="548">
<emphasis id="B97AEAA0FFF8FFD135D4F3991C6AFF14" bold="true" box="[199,368,150,176]" pageId="7" pageNumber="548">Relationships.</emphasis>
See Discussion.
</paragraph>
</subSubSection>
<subSubSection id="C3146539FFF8FFD135D4F3B51867FF5D" pageId="7" pageNumber="548" type="distribution">
<paragraph id="8BB136B2FFF8FFD135D4F3B51867FF5D" blockId="7.[151,1436,150,249]" pageId="7" pageNumber="548">
<emphasis id="B97AEAA0FFF8FFD135D4F3B51C7BFF70" bold="true" box="[199,353,186,212]" pageId="7" pageNumber="548">Distribution.</emphasis>
<taxonomicName id="4C0E4D31FFF8FFD13474F3B41CC4FF71" authority="Martens &amp; Maghradze &amp; Barjadze, 2021" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2021" box="[359,478,187,213]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="548" phylum="Arthropoda" rank="species" species="melouri" status="sp. nov.">
<emphasis id="B97AEAA0FFF8FFD13474F3B41CC4FF71" box="[359,478,187,213]" italics="true" pageId="7" pageNumber="548">N. melouri</emphasis>
</taxonomicName>
<emphasis id="B97AEAA0FFF8FFD134F7F3B41F27FF71" bold="true" box="[484,573,187,213]" pageId="7" pageNumber="548">
<taxonomicNameLabel id="A24957DBFFF8FFD134F7F3B41F27FF71" box="[484,573,187,213]" pageId="7" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from Melouri Cave in Sataplia-Tskaltubo karst massif, 
<collectingRegion id="49CAF850FFF8FFD13058F3B41DFBFF5D" country="Georgia" name="Imereti" pageId="7" pageNumber="548">Imereti region</collectingRegion>
, Western 
<collectingCountry id="F3197622FFF8FFD1345DF3D01CB1FF5D" box="[334,427,223,249]" name="Georgia" pageId="7" pageNumber="548">Georgia</collectingCountry>
. The distance between the entrances of Melouri and Prometheus caves is about 
<quantity id="4CF69B57FFF8FFD13034F3D11862FF5D" box="[1319,1400,222,249]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="548" unit="km" value="2.5">2.5 km</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>