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<document id="D1AB5C2D78D9C48C2AD948118B18DF6E" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625105362570" checkinUser="felipe" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel-Angel" docDate="2005" docId="03C487D0C45FFFBB174FFA742033A594" docLanguage="en" docName="zt01012p037.pdf" docOrigin="Zootaxa 1012 (1012)" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrodactylus costulatus Bates 1887" docType="treatment" docVersion="3" lastPageNumber="26" masterDocId="FFFDFFA8C45DFFB81647FFEA2418A64A" masterDocTitle="New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua" masterLastPageNumber="37" masterPageNumber="23" pageNumber="25" updateTime="1698980141271" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="838A67B7C50479A5CFB5C562F0675851" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8335B51D282A62095D04D8B7778F12B7">
<mods:title id="9144F935DFC09C152668B4980C760125">New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua</mods:title>
</mods:titleInfo>
<mods:name id="8517D55A8F32B088A12D236D8C4A5D20" type="personal">
<mods:role id="4C2DB28056E5B6BFE97159BFD5FC917D">
<mods:roleTerm id="11E7BBD996466A4D9B6F5574D865B6B3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5F7926E81573C3611E1399DB201BE92">Arce-Pérez, Roberto</mods:namePart>
</mods:name>
<mods:name id="F7144185A34DA2788D8CA8D2BB84E920" type="personal">
<mods:role id="5EB171CA4119E8996586BA1C83BDE087">
<mods:roleTerm id="89BFD7834FF48C5E22B3C6ED5884E648">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1D85E4C5D4EC1F7548CFAB4F5B0D022">Morón, Miguel-Angel</mods:namePart>
</mods:name>
<mods:typeOfResource id="B93EA32156B4132D003D3EFF58422F5F">text</mods:typeOfResource>
<mods:relatedItem id="8397E3B6998B2D71C695D9398155E185" type="host">
<mods:titleInfo id="145AA85F13B3DACD4063D17C29747586">
<mods:title id="D33F140F877DD21D9C42040A026644F4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D641F4243672830B3BDE91532A98DCEA">
<mods:date id="F7F0600F4FE642B5EA383AAAE2B05690">2005</mods:date>
<mods:detail id="5029F7138153460347821C116AF38B72" type="pubDate">
<mods:number id="8C78909EEFEAA702328A1DB32B83A625">2005-06-28</mods:number>
</mods:detail>
<mods:detail id="44A98204B3EF4AD54ACC69F3EFAA6BDE" type="volume">
<mods:number id="AF64728AA7B2DE1221AB2F3809B0BFF3">1012</mods:number>
</mods:detail>
<mods:detail id="09C41525FD4A45965465F6CEA9EC4E7C" type="issue">
<mods:number id="9C4A054D9764D22B93C1E485911CFF7D">1012</mods:number>
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<mods:extent id="86AA72BA92CA3D88919144EA8B2CD057" unit="page">
<mods:start id="633F3E09C76CF787E779EA43CCEBF560">23</mods:start>
<mods:end id="622DA6844914929A4DA1FAAF6475B1EC">37</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B386AF9C191B2ED5E348FC0269362925">journal article</mods:classification>
<mods:identifier id="F6E8F2CA4BB19B24DE4A1C4B7DD38666" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="EE75A8DEDC9D8476B223F02583634020" type="ZooBank">E84D2D81-6B26-4311-B56F-6D3C4E672C78</mods:identifier>
</mods:mods>
<treatment id="03C487D0C45FFFBB174FFA742033A594" ID-DOI="http://doi.org/10.5281/zenodo.5049174" ID-Zenodo-Dep="5049174" LSID="urn:lsid:plazi:treatment:03C487D0C45FFFBB174FFA742033A594" httpUri="http://treatment.plazi.org/id/03C487D0C45FFFBB174FFA742033A594" lastPageId="3" lastPageNumber="26" pageId="2" pageNumber="25">
<subSubSection id="C377654DC45FFFBA174FFA7426C9A3F2" box="[264,721,1438,1464]" pageId="2" pageNumber="25" type="nomenclature">
<paragraph id="8BD236C6C45FFFBA174FFA7426C9A3F2" blockId="2.[264,721,1438,1464]" box="[264,721,1438,1464]" pageId="2" pageNumber="25">
<heading id="D09A81AAC45FFFBA174FFA7426C9A3F2" bold="true" box="[264,721,1438,1464]" fontSize="11" level="1" pageId="2" pageNumber="25" reason="1">
<taxonomicName id="4C6D4D45C45FFFBA174FFA7426C9A3F2" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[264,721,1438,1464]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="B919EAD4C45FFFBA174FFA7426C9A3F2" bold="true" box="[264,721,1438,1464]" pageId="2" pageNumber="25">
<emphasis id="B919EAD4C45FFFBA174FFA742624A3F2" bold="true" box="[264,572,1438,1464]" italics="true" pageId="2" pageNumber="25">Macrodactylus costulatus</emphasis>
<bibRefCitation id="EFFC4B37C45FFFBA1403FA7426C9A3F2" author="Bates, H. W." box="[580,721,1438,1464]" pageId="2" pageNumber="25" pagination="2" refId="ref6913" refString="Bates, H. W. (1887) Pectinicornia and Lamellicornia. In: Salvin, O. &amp; Godma, F. (Eds.), Biologia Centrali- Americana, Insecta, Coleoptera, Vol. II, parte 2. Taylor and Francis, London." type="book chapter" year="1887">Bates, 1887</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45FFFBA174FFA032740A0B9" pageId="2" pageNumber="25" type="diagnosis">
<paragraph id="8BD236C6C45FFFBA174FFA032740A0B9" blockId="2.[264,1323,1513,1899]" pageId="2" pageNumber="25">
<emphasis id="B919EAD4C45FFFBA174FFA032598A049" bold="true" box="[264,384,1513,1539]" pageId="2" pageNumber="25">Diagnosis</emphasis>
. This species is distinguished by the following combination of characters: antennal club reddish­brown with the apex darkened; pronotum black with scattered scalelike yellow setae; elytra with two well­defined longitudinal stripes of scale­like setae on the interstriae 1 and 3, interstriae 2 and 4 costate, glabrous (
<figureCitation id="13562A43C45FFFBA159DF98B2038A031" box="[986,1056,1633,1659]" captionStart="FIGURES 1–6" captionStartId="6.[264,383,1501,1525]" captionTargetBox="[275,1312,313,1474]" captionTargetId="figure-83@6.[275,1312,309,1474]" captionTargetPageId="6" captionText="FIGURES 1–6. 1­­­ Macrodactylus costulatus Bates, male; 2­­­ M. sericeicollis Bates, male; 3­­­ M. sericinus Bates, male; 4­­­ M. suavis Bates, male; 5­­­ M. dimidiatus Guérin­Méneville, male; 6­ ­­ M. sylphis Bates, male." pageId="2" pageNumber="25">Fig. 1</figureCitation>
); pygidium with scattered, scale­like, light yellow setae; abdomen black; femora and tibiae reddish­brown; tarsi without white setae; apex of the parameres lanceolate, without setae on the external borders; vestiture and color of female similar to male.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45FFFBB177FF8EB260DA7E4" lastPageId="3" lastPageNumber="26" pageId="2" pageNumber="25" type="materials_examined">
<paragraph id="8BD236C6C45FFFBB177FF8EB260DA7E4" blockId="2.[264,1323,1513,1899]" lastBlockId="3.[264,1324,284,990]" lastPageId="3" lastPageNumber="26" pageId="2" pageNumber="25">
<emphasis id="B919EAD4C45FFFBA177FF8EB2635A151" bold="true" box="[312,557,1793,1819]" pageId="2" pageNumber="25">Material examined</emphasis>
(
<specimenCount id="9D6BFD4FC45FFFBA1400F8EB26D3A151" box="[583,715,1793,1819]" pageId="2" pageNumber="25" type="male">143 males</specimenCount>
,
<specimenCount id="9D6BFD4FC45FFFBA14A7F8EB2764A151" box="[736,892,1793,1819]" pageId="2" pageNumber="25" type="female">108 females</specimenCount>
).
<materialsCitation id="3B053C9BC45FFFBA15DEF8EB2659A121" collectingDate="1994-06-11" collectionCode="NEW, COUNTRY, RECORDS" collectorName="S. Zamorano &amp; E. Galeras" country="Honduras" elevation="1700" location="El Paraiso" pageId="2" pageNumber="25" specimenCount="1" stateProvince="El Paraiso">
<emphasis id="B919EAD4C45FFFBA15DEF8EB213AA151" bold="true" box="[921,1314,1793,1819]" pageId="2" pageNumber="25">
<collectionCode id="ED7CAE03C45FFFBA15DEF8EB27C7A151" box="[921,991,1793,1819]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="2" pageNumber="25">NEW</collectionCode>
<collectionCode id="ED7CAE03C45FFFBA15B7F8EB213AA151" box="[1008,1314,1793,1819]" pageId="2" pageNumber="25">COUNTRY RECORDS</collectionCode>
</emphasis>
:
<collectingCountry id="F37A7656C45FFFBA174FF8C325A8A109" box="[264,432,1833,1859]" name="Honduras" pageId="2" pageNumber="25">HONDURAS</collectingCountry>
:
<collectingRegion id="49A9F824C45FFFBA17FCF8C3262BA109" box="[443,563,1833,1859]" country="Honduras" name="El Paraiso" pageId="2" pageNumber="25">El Paraiso</collectingRegion>
<quantity id="4C959B23C45FFFBA147CF8C32682A109" box="[571,666,1833,1859]" metricMagnitude="4" metricUnit="m" metricValue="1.1099999999999999" pageId="2" pageNumber="25" unit="km" value="11.1">11.1 km</quantity>
<collectorName id="26985310C45FFFBA14E4F8C32723A109" box="[675,827,1833,1859]" pageId="2" pageNumber="25">S. Zamorano</collectorName>
&amp;
<quantity id="4C959B23C45FFFBA1525F8C327B4A109" box="[866,940,1833,1859]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="25" unit="km" value="10.0">10 km</quantity>
<collectorName id="26985310C45FFFBA15F2F8C3202AA109" box="[949,1074,1833,1859]" pageId="2" pageNumber="25">E. Galeras</collectorName>
,
<quantity id="4C959B23C45FFFBA1278F8C32082A109" box="[1087,1178,1833,1859]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="25" unit="m" value="1700.0">
<elevation id="0040D1F5C45FFFBA1278F8C32082A109" box="[1087,1178,1833,1859]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="25" unit="m" value="1700.0">1700 m</elevation>
</quantity>
, oak / cloud forest,
<date id="FFD31006C45FFFBA1723F8BB261DA121" box="[356,517,1873,1899]" pageId="2" pageNumber="25" value="1994-06-11">
<collectingDate id="EF97E9EEC45FFFBA1723F8BB261DA121" box="[356,517,1873,1899]" pageId="2" pageNumber="25" value="1994-06-11">11 June 1994</collectingDate>
</date>
, Col
</materialsCitation>
.
<materialsCitation id="3B053C9BC45FFFBB140AF8BB2609A7E4" collectingDate="1994-05-15" collectingDateMax="1994-06-08" collectingDateMin="1994-05-15" collectionCode="R, CMNC, BDGC, EAPZ, HAHC, IEXA, MXAL" collectorName="S. Anderson &amp; Cols. H" country="Honduras" elevation="1030" lastPageId="3" lastPageNumber="26" location="Paraiso Yuscaran" pageId="2" pageNumber="25" specimenCount="251" specimenCount-female="108" specimenCount-male="143" stateProvince="El Paraiso">
<collectionCode id="ED7CAE03C45FFFBA140AF8BB267CA121" box="[589,612,1873,1899]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="25">R</collectionCode>
.
<collectorName id="26985310C45FFFBA1436F8BB271FA121" box="[625,775,1873,1899]" pageId="2" pageNumber="25">S. Anderson</collectorName>
,
<specimenCount id="9D6BFD4FC45FFFBA1553F8BB2770A121" box="[788,872,1873,1899]" pageId="2" pageNumber="25" type="male">1 male</specimenCount>
,
<specimenCount id="9D6BFD4FC45FFFBA1532F8BB27C5A121" box="[885,989,1873,1899]" pageId="2" pageNumber="25" type="female">1 female</specimenCount>
(
<collectionCode id="ED7CAE03C45FFFBA15A9F8BB2053A121" box="[1006,1099,1873,1899]" pageId="2" pageNumber="25">CMNC</collectionCode>
);
<location id="8EB2601DC45FFFBA1227F8BB2132A121" LSID="urn:lsid:plazi:treatment:03C487D0C45FFFBB174FFA742033A594:8EB2601DC45FFFBA1227F8BB2132A121" box="[1120,1322,1873,1899]" country="Honduras" name="Paraiso Yuscaran" pageId="2" pageNumber="25">Paraíso Yuscarán</location>
<quantity id="4C959B23C45EFFBB174FFEF6257DA77C" box="[264,357,284,310]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="26" unit="m" value="1030.0">
<elevation id="0040D1F5C45EFFBB174FFEF6257DA77C" box="[264,357,284,310]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="26" unit="m" value="1030.0">1030 m</elevation>
</quantity>
,
<date id="FFD31006C45EFFBB1734FEF6261EA77C" box="[371,518,284,310]" pageId="3" pageNumber="26" value="1994-06-08">
<collectingDate id="EF97E9EEC45EFFBB1734FEF6261EA77C" box="[371,518,284,310]" pageId="3" pageNumber="26" value="1994-06-08">8 June 1994</collectingDate>
</date>
,
<collectedFrom id="3871753BC45EFFBB1453FEF6200AA77C" box="[532,1042,284,310]" pageId="3" pageNumber="26">
on
<taxonomicName id="4C6D4D45C45EFFBB147CFEF6200AA77C" authority=", Cols. H &amp; A Howden" authorityName="Cols. H &amp; A Howden" box="[571,1042,284,310]" class="Magnoliopsida" family="Sapotaceae" genus="Pouteria" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="26" phylum="Tracheophyta" rank="species" species="sapota">
<emphasis id="B919EAD4C45EFFBB147CFEF626E0A77F" box="[571,760,284,309]" italics="true" pageId="3" pageNumber="26">Pouteria sapota</emphasis>
, Cols. H &amp; A Howden
</taxonomicName>
</collectedFrom>
,
<specimenCount id="9D6BFD4FC45EFFBB1258FEF62083A77C" box="[1055,1179,284,310]" pageId="3" pageNumber="26" type="male">131 males</specimenCount>
,
<specimenCount id="9D6BFD4FC45EFFBB12EEFEF62132A77C" box="[1193,1322,284,310]" pageId="3" pageNumber="26" type="female">96 females</specimenCount>
(
<collectionCode id="ED7CAE03C45EFFBB1757FEAE257EA714" box="[272,358,324,350]" pageId="3" pageNumber="26">BDGC</collectionCode>
,
<collectionCode id="ED7CAE03C45EFFBB1734FEAE25A7A714" box="[371,447,324,350]" country="Honduras" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33268" lsid="urn:lsid:biocol.org:col:33268" name="Escuela Agricola Panamericana" pageId="3" pageNumber="26">EAPZ</collectionCode>
,
<collectionCode id="ED7CAE03C45EFFBB178AFEAE263BA714" box="[461,547,324,350]" pageId="3" pageNumber="26">HAHC</collectionCode>
,
<collectionCode id="ED7CAE03C45EFFBB1477FEAE2661A714" box="[560,633,324,350]" pageId="3" pageNumber="26">IEXA</collectionCode>
,
<collectionCode id="ED7CAE03C45EFFBB14C1FEAE26FAA714" box="[646,738,324,350]" pageId="3" pageNumber="26">MXAL</collectionCode>
); Cerro Monserrat,
<quantity id="4C959B23C45EFFBB158CFEAE2010A714" box="[971,1032,324,350]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="26" unit="km" value="7.0">7 km</quantity>
SW, Yuscarán,
<date id="FFD31006C45EFFBB1288FEAE255DA7CC" pageId="3" pageNumber="26" value="1994-05-24">
<collectingDate id="EF97E9EEC45EFFBB1288FEAE255DA7CC" pageId="3" pageNumber="26" value="1994-05-24">24 May 1994</collectingDate>
</date>
,
<quantity id="4C959B23C45EFFBB1712FE8625ADA7CC" box="[341,437,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="26" unit="m" value="1800.0">
<elevation id="0040D1F5C45EFFBB1712FE8625ADA7CC" box="[341,437,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="26" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<collectorName id="26985310C45EFFBB1782FE86263AA7CC" box="[453,546,364,390]" pageId="3" pageNumber="26">Cols. H</collectorName>
&amp; A Howden,
<specimenCount id="9D6BFD4FC45EFFBB149AFE862753A7CC" box="[733,843,364,390]" pageId="3" pageNumber="26" type="male">10 males</specimenCount>
,
<specimenCount id="9D6BFD4FC45EFFBB151CFE8627F9A7CC" box="[859,993,364,390]" pageId="3" pageNumber="26" type="female">11 females</specimenCount>
; same data except
<date id="FFD31006C45EFFBB128BFE86255DA7E4" pageId="3" pageNumber="26" value="1994-05-15">
<collectingDate id="EF97E9EEC45EFFBB128BFE86255DA7E4" pageId="3" pageNumber="26" value="1994-05-15">15 May 1994</collectingDate>
</date>
,
<specimenCount id="9D6BFD4FC45EFFBB1716FE7E25B8A7E4" box="[337,416,404,430]" pageId="3" pageNumber="26" type="male">1 male</specimenCount>
(
<collectionCode id="ED7CAE03C45EFFBB17F7FE7E2612A7E4" box="[432,522,404,430]" pageId="3" pageNumber="26">HAHC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45EFFBB177FFE512619A43C" pageId="3" pageNumber="26" type="biology_ecology">
<paragraph id="8BD236C6C45EFFBB177FFE512619A43C" blockId="3.[264,1324,284,990]" pageId="3" pageNumber="26">
<emphasis id="B919EAD4C45EFFBB177FFE512584A79F" bold="true" box="[312,412,443,469]" pageId="3" pageNumber="26">Biology.</emphasis>
This species inhabits tropical rain forest, tropical plantations, cloud forest, and oak forest located at altitudes from
<quantity id="4C959B23C45EFFBB149CFE0E2792A7B4" box="[731,906,484,510]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="18.0" metricValueMin="1.0" pageId="3" pageNumber="26" unit="m" value="950.0" valueMax="1800.0" valueMin="100.0">100 to 1800 m</quantity>
. Months of collection in
<collectingCountry id="F37A7656C45EFFBB12F6FE0E2130A7B4" box="[1201,1320,484,510]" name="Honduras" pageId="3" pageNumber="26">Honduras</collectingCountry>
: May (22), June (229). Many specimens were collected on
<taxonomicName id="4C6D4D45C45EFFBB15F7FDE62073A46F" box="[944,1131,524,549]" class="Magnoliopsida" family="Sapotaceae" genus="Pouteria" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="26" phylum="Tracheophyta" rank="species" species="sapota">
<emphasis id="B919EAD4C45EFFBB15F7FDE62073A46F" box="[944,1131,524,549]" italics="true" pageId="3" pageNumber="26">Pouteria sapota</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C6D4D45C45EFFBB12DCFDE6213AA46C" box="[1179,1314,524,550]" class="Magnoliopsida" family="Sapotaceae" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="26" phylum="Tracheophyta" rank="family">Sapotaceae</taxonomicName>
) at Yuscarán. Another species of
<taxonomicName id="4C6D4D45C45EFFBB14D0FDDE275EA407" authorityName="Dejean" authorityYear="1821" box="[663,838,564,589]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B919EAD4C45EFFBB14D0FDDE275EA407" box="[663,838,564,589]" italics="true" pageId="3" pageNumber="26">Macrodactylus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C6D4D45C45EFFBB151BFDDE27C1A407" box="[860,985,564,589]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B919EAD4C45EFFBB151BFDDE27C1A407" box="[860,985,564,589]" italics="true" pageId="3" pageNumber="26">M. tibialis</emphasis>
</taxonomicName>
) was collected in the same place, time, and host.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45EFFBB177FFD6E25D0A48C" pageId="3" pageNumber="26" type="distribution">
<paragraph id="8BD236C6C45EFFBB177FFD6E25D0A48C" blockId="3.[264,1324,284,990]" pageId="3" pageNumber="26">
<emphasis id="B919EAD4C45EFFBB177FFD6E25C9A4D4" bold="true" box="[312,465,644,670]" pageId="3" pageNumber="26">Distribution</emphasis>
. Southeastern México to southeastern
<collectingCountry id="F37A7656C45EFFBB15DFFD6E2012A4D4" box="[920,1034,644,670]" name="Honduras" pageId="3" pageNumber="26">Honduras</collectingCountry>
(13 ° 30'–
<geoCoordinate id="EE595001C45EFFBB12C0FD6E20EFA4D4" box="[1159,1271,644,670]" degrees="15" direction="north" minutes="10" orientation="latitude" pageId="3" pageNumber="26" precision="925" value="15.166667">15° 10' N</geoCoordinate>
, 86° 10' –
<geoCoordinate id="EE595001C45EFFBB1705FD4625A3A48C" box="[322,443,684,710]" degrees="92" direction="west" minutes="30" orientation="longitude" pageId="3" pageNumber="26" precision="925" value="-92.5">92° 30' W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45EFFBB177FFD3E2033A594" pageId="3" pageNumber="26" type="discussion">
<paragraph id="8BD236C6C45EFFBB177FFD3E2033A594" blockId="3.[264,1324,284,990]" pageId="3" pageNumber="26">
<emphasis id="B919EAD4C45EFFBB177FFD3E25A9A4A4" bold="true" box="[312,433,724,750]" pageId="3" pageNumber="26">Remarks.</emphasis>
This species was originally described with a large series of specimens from Sarstoon River (
<collectingCountry id="F37A7656C45EFFBB1789FD162607A55C" box="[462,543,764,790]" name="Belize" pageId="3" pageNumber="26">Belize</collectingCountry>
); Dueñas, Coban (
<collectingCountry id="F37A7656C45EFFBB14B0FD162765A55C" box="[759,893,764,790]" name="Guatemala" pageId="3" pageNumber="26">Guatemala</collectingCountry>
);
<collectingRegion id="49A9F824C45EFFBB15D7FD16201DA55C" box="[912,1029,764,790]" country="Nicaragua" name="Chontales" pageId="3" pageNumber="26">Chontales</collectingRegion>
(
<collectingCountry id="F37A7656C45EFFBB1252FD162089A55C" box="[1045,1169,764,790]" name="Nicaragua" pageId="3" pageNumber="26">Nicaragua</collectingCountry>
); and Volcan de Irazu (
<collectingCountry id="F37A7656C45EFFBB1731FCCE25E5A574" box="[374,509,804,830]" name="Costa Rica" pageId="3" pageNumber="26">Costa Rica</collectingCountry>
). Arce­Pérez and
<bibRefCitation id="EFFC4B37C45EFFBB148BFCCE2769A574" author="Moron, M. A." box="[716,881,804,830]" pageId="3" pageNumber="26" pagination="123 - 239" refId="ref6864" refString="Arce-Perez R. &amp; Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) de Mexico y America del Norte. Acta Zoologica Mexicana (n. s), 79, 123 - 239." type="journal article" year="2000">Morón (2000)</bibRefCitation>
added records from Tapachula,
<collectingRegion id="49A9F824C45EFFBB12AEFCCE2529A52C" country="Mexico" name="Chiapas" pageId="3" pageNumber="26">Chiapas</collectingRegion>
, (
<collectingCountry id="F37A7656C45EFFBB170EFCA625BEA52C" box="[329,422,844,870]" name="Mexico" pageId="3" pageNumber="26">México</collectingCountry>
) and Palmar Viejo,
<collectingRegion id="49A9F824C45EFFBB14DAFCA6274DA52C" box="[669,853,844,870]" country="Guatemala" name="Quetzaltenango" pageId="3" pageNumber="26">Quetzaltenango</collectingRegion>
(
<collectingCountry id="F37A7656C45EFFBB1520FCA627F5A52C" box="[871,1005,844,870]" name="Guatemala" pageId="3" pageNumber="26">Guatemala</collectingCountry>
). The variation in the
<specimenCount id="9D6BFD4FC45EFFBB12B8FCA6259AA5C4" pageId="3" pageNumber="26" type="generic">251 specimens</specimenCount>
studied from
<collectingCountry id="F37A7656C45EFFBB1463FC9E268EA5C4" box="[548,662,884,910]" name="Honduras" pageId="3" pageNumber="26">Honduras</collectingCountry>
was within the parameters of the specimens studied previously. The new records aid to link the distribution of this species between Chontales and
<collectingCountry id="F37A7656C45EFFBB174FFC2E2592A594" box="[264,394,964,990]" name="Guatemala" pageId="3" pageNumber="26">Guatemala</collectingCountry>
, across the chain of mountains in southeastern
<collectingCountry id="F37A7656C45EFFBB15F6FC2E203FA594" box="[945,1063,964,990]" name="Honduras" pageId="3" pageNumber="26">Honduras</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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