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<document id="55DB41FD066B368200D89D89F2FEC2E9" ID="10.11646/zootaxa.4231.4.2SLASH10150" ID-CLB-Dataset="36559" ID-DOI="10.11646/zootaxa.4231.4.2" ID-GBIF-Dataset="fb0a7416-8d2e-4793-9dc3-525ffccb9e62" ID-ISSN="1175-5326" ID-Zenodo-Dep="292674" ID-ZooBank="99D539A1-D4BF-48C4-AEE1-0CA8F198C631" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487060700847" checkinUser="plazi" docAuthor="Cruz, Paulo Vilela, Salles, Frederico Falcão &amp; Hamada, Neusa" docDate="2017" docId="039E3278FFAFFFE1DDDA25AC0B7891EA" docLanguage="en" docName="zootaxa.4231.4.2SLASH10150.pdf" docOrigin="Zootaxa 4231 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Callibaetis nigrivenosus Banks 1918" docType="treatment" docVersion="12" lastPageNumber="513" masterDocId="FFA74A00FFA3FFECDD4D253B09589112" masterDocTitle="Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America" masterLastPageNumber="534" masterPageNumber="500" pageNumber="512" updateTime="1698436872380" updateUser="ExternalLinkService">
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<mods:title id="8FBFD73839FF5346971D1E70919C7D6D">Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America</mods:title>
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<mods:namePart id="B34F05CC30F7F287572F375F1002A136">Cruz, Paulo Vilela</mods:namePart>
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<mods:namePart id="4BF0F973142329EDE3CCCE579EEA7E59">Salles, Frederico Falcão</mods:namePart>
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<mods:namePart id="F4E2262C73DD47A74E972C45CDDEF417">Hamada, Neusa</mods:namePart>
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<paragraph id="8B88836EFFAFFFE0DDDA25AC0A4891A3" blockId="12.[151,784,151,211]" box="[151,784,151,178]" pageId="12" pageNumber="512">
<heading id="D0C03402FFAFFFE0DDDA25AC0A4891A3" bold="true" box="[151,784,151,178]" fontSize="11" level="1" pageId="12" pageNumber="512" reason="1">
<emphasis id="B9435F7CFFAFFFE0DDDA25AC0A4891A3" bold="true" box="[151,784,151,178]" pageId="12" pageNumber="512">
<taxonomicName id="4C37F8EDFFAFFFE0DDDA25AC0B0991A0" ID-CoL="PM94" authority="Banks 1918" authorityName="Banks" authorityYear="1918" box="[151,593,151,178]" class="Insecta" family="Baetidae" genus="Callibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="512" phylum="Arthropoda" rank="species" species="nigrivenosus">
<emphasis id="B9435F7CFFAFFFE0DDDA25AC08E291A3" bold="true" box="[151,442,151,177]" italics="true" pageId="12" pageNumber="512">Callibaetis nigrivenosus</emphasis>
<bibRefCitation id="EFA6FE9FFFAFFFE0DC8C25A30B0991A0" author="Banks" box="[449,593,152,178]" pageId="12" pageNumber="512" refString="Banks, N. (1918) New neuropteroid insects. Bulletin of the Museum of Comparative Zoology, 62 (1), 1 - 22. https: // doi. org / 10.5962 / bhl. title. 28704" type="journal article" year="1918">Banks 1918</bibRefCitation>
</taxonomicName>
<emphasis id="B9435F7CFFAFFFE0DF1525AC0A4891A3" bold="true" box="[600,784,151,177]" italics="true" pageId="12" pageNumber="512">nomen dubium</emphasis>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B88836EFFAFFFE0DDDA2581086F91C1" blockId="12.[151,784,151,211]" box="[151,311,186,211]" pageId="12" pageNumber="512">
(
<figureCitation id="130C9FEBFFAFFFE0DDD2258109B291C1" box="[159,234,186,211]" captionStart="FIGURE 8" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[152,1435,757,1925]" captionTargetId="figure@12.[152,1435,757,1925]" captionTargetPageId="12" captionText="FIGURE 8. Callibaetis nigrivenosus nomen dubium female imago (holotype). A. labels (Harvard); B. lateral view of head and thorax; C. lateral view of abdomen; D. left forewing; E. right forewing." httpUri="https://zenodo.org/record/292682/files/figure.png" pageId="12" pageNumber="512">Figs. 8</figureCitation>
A8E)
</paragraph>
</subSubSection>
<subSubSection id="C32DD0E5FFAFFFE0DDDA24390829906E" pageId="12" pageNumber="512" type="reference_group">
<paragraph id="8B88836EFFAFFFE0DDDA24390AFC9025" blockId="12.[151,940,258,311]" pageId="12" pageNumber="512">
<treatmentCitationGroup id="AB27A440FFAFFFE0DDDA24390AFC9025" pageId="12" pageNumber="512">
<taxonomicName id="4C37F8EDFFAFFFE0DDDA24390B70900A" authority="Banks 1918: 11" authorityName="Banks" authorityPageNumber="11" authorityYear="1918" box="[151,552,258,280]" class="Insecta" family="Baetidae" genus="Callibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="512" phylum="Arthropoda" rank="species" species="nigrivenosa">
<emphasis id="B9435F7CFFAFFFE0DDDA243908DC900A" box="[151,388,258,280]" italics="true" pageId="12" pageNumber="512">Callibaetis nigrivenosa</emphasis>
<treatmentCitation id="0A96A57FFFAFFFE0DCC724390B70900A" author="Banks" box="[394,552,258,280]" page="11" pageId="12" pageNumber="512" year="1918">
<bibRefCitation id="EFA6FE9FFFAFFFE0DCC724390B5E900A" author="Banks" box="[394,518,258,280]" pageId="12" pageNumber="512" refString="Banks, N. (1918) New neuropteroid insects. Bulletin of the Museum of Comparative Zoology, 62 (1), 1 - 22. https: // doi. org / 10.5962 / bhl. title. 28704" type="journal article" year="1918">Banks 1918</bibRefCitation>
: 11
</treatmentCitation>
</taxonomicName>
(name emended by
<bibRefCitation id="EFA6FE9FFFAFFFE0DFB824390AFD900A" author="McCafferty" box="[757,933,258,280]" pageId="12" pageNumber="512" refString="McCafferty, W. P. (1996) Emendations to the Callibaetis (Ephemeroptera: Baetidae) of South America. Entomological News, 107 (4), 230 - 232." type="journal article" year="1996">McCafferty 1996</bibRefCitation>
)
<taxonomicName id="4C37F8EDFFAFFFE0DDDA241A0B219025" ID-CoL="PM94" authority="McCafferty 1996: 231" authorityName="McCafferty" authorityPageNumber="231" authorityYear="1996" box="[151,633,289,311]" class="Insecta" family="Baetidae" genus="Callibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="512" phylum="Arthropoda" rank="species" species="nigrivenosus">
<emphasis id="B9435F7CFFAFFFE0DDDA241A08D69025" box="[151,398,289,311]" italics="true" pageId="12" pageNumber="512">Callibaetis nigrivenosus</emphasis>
<treatmentCitation id="0A96A57FFFAFFFE0DCD9241A0B219025" author="McCafferty" box="[404,633,289,311]" page="231" pageId="12" pageNumber="512" year="1996">
<bibRefCitation id="EFA6FE9FFFAFFFE0DCD9241A0B1E9025" author="McCafferty" box="[404,582,289,311]" pageId="12" pageNumber="512" refString="McCafferty, W. P. (1996) Emendations to the Callibaetis (Ephemeroptera: Baetidae) of South America. Entomological News, 107 (4), 230 - 232." type="journal article" year="1996">McCafferty 1996</bibRefCitation>
: 231
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A96A57FFFAFFFE0DFCE241A0AF99025" author="Dominguez" box="[643,929,289,311]" page="115" pageId="12" pageNumber="512" year="2006">
<bibRefCitation id="EFA6FE9FFFAFFFE0DFCE241A0A379025" author="Dominguez" box="[643,879,289,311]" pageId="12" pageNumber="512" refString="Dominguez, E., Molineri, C., Pescador, M. L., Hubbard, M. D. &amp; Nieto, C. (2006) Aquatic Biodiversity of Latin America, Ephemeroptera of South America. Uol. 2. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">
Domínguez
<emphasis id="B9435F7CFFAFFFE0DE4D24190A6B9025" box="[768,819,289,311]" italics="true" pageId="12" pageNumber="512">et al.</emphasis>
2006
</bibRefCitation>
: 115
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B88836EFFAFFFE0DDDA245F0829906E" blockId="12.[151,1436,356,704]" box="[151,369,356,381]" pageId="12" pageNumber="512">
<emphasis id="B9435F7CFFAFFFE0DDDA245F080D906F" bold="true" box="[151,341,356,381]" pageId="12" pageNumber="512">Known stages. I</emphasis>
♀.
</paragraph>
</subSubSection>
<subSubSection id="C32DD0E5FFAFFFE0DD8A24BC09AA93AD" pageId="12" pageNumber="512" type="discussion">
<paragraph id="8B88836EFFAFFFE0DD8A24BC09AA93AD" blockId="12.[151,1436,356,704]" pageId="12" pageNumber="512">
<emphasis id="B9435F7CFFAFFFE0DD8A24BC081790B2" bold="true" box="[199,335,391,416]" pageId="12" pageNumber="512">Comments.</emphasis>
We studied the
<typeStatus id="548C3DCCFFAFFFE0DF4A24B30B6F90B2" box="[519,567,392,416]" pageId="12" pageNumber="512">type</typeStatus>
specimen from photographs (
<figureCitation id="130C9FEBFFAFFFE0DEC624BC0A8190B2" box="[907,985,391,416]" captionStart="FIGURE 8" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[152,1435,757,1925]" captionTargetId="figure@12.[152,1435,757,1925]" captionTargetPageId="12" captionText="FIGURE 8. Callibaetis nigrivenosus nomen dubium female imago (holotype). A. labels (Harvard); B. lateral view of head and thorax; C. lateral view of abdomen; D. left forewing; E. right forewing." httpUri="https://zenodo.org/record/292682/files/figure.png" pageId="12" pageNumber="512">Figs. 8</figureCitation>
A8E). It is fragile, badly damaged and lacks the hind wings. The only reliable structure is the forewing, which allowed us to identify the specimen to the family level (
<taxonomicName id="4C37F8EDFFAFFFE0DC6024F408CD90FA" box="[301,405,463,488]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="512" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
). The absence of hind wings in the
<typeStatus id="548C3DCCFFAFFFE0DE6B24EB0A0E90FA" box="[806,854,464,488]" pageId="12" pageNumber="512">type</typeStatus>
specimen and its description lead us to believe that these structures were always absent. The manuscript is very brief and is not illustrated. The
<typeStatus id="548C3DCCFFAFFFE0D9D324CE0D96931F" box="[1182,1230,501,525]" pageId="12" pageNumber="512">type</typeStatus>
locality presented in the manuscript and shown on the label is not precise, referring to a broad geographical area. Even if additional specimens of this species were collected, there would be little possibility of a reliable identification using the
<typeStatus id="548C3DCCFFAFFFE0D82C27060CC49347" box="[1377,1436,573,597]" pageId="12" pageNumber="512">types</typeStatus>
or the original description. Based on the lack of information in the description, the damaged
<typeStatus id="548C3DCCFFAFFFE0D84B275B0C6E936A" box="[1286,1334,608,632]" pageId="12" pageNumber="512">type</typeStatus>
and the improbability of assigning specimens to this species, we propose
<taxonomicName id="4C37F8EDFFAFFFE0DEEC27BF0C51938F" authority="Banks" authorityName="Banks" box="[929,1289,644,669]" class="Insecta" family="Baetidae" genus="Callibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="512" phylum="Arthropoda" rank="species" species="nigrivenosus">
<emphasis id="B9435F7CFFAFFFE0DEEC27BF0DEF938E" box="[929,1207,644,668]" italics="true" pageId="12" pageNumber="512">Callibaetis nigrivenosus</emphasis>
Banks
</taxonomicName>
as a
<emphasis id="B9435F7CFFAFFFE0D81C27BE09B393D2" italics="true" pageId="12" pageNumber="512">nomen dubium</emphasis>
.
</paragraph>
</subSubSection>
<caption id="DF48D3E6FFAFFFE0DDDA22A10A3196DD" httpUri="https://zenodo.org/record/292682/files/figure.png" pageId="12" pageNumber="512" targetBox="[152,1435,757,1925]" targetPageId="12">
<paragraph id="8B88836EFFAFFFE0DDDA22A10A3196DD" blockId="12.[151,1435,1946,1999]" pageId="12" pageNumber="512">
<emphasis id="B9435F7CFFAFFFE0DDDA22A1084A96BD" bold="true" box="[151,274,1946,1968]" pageId="12" pageNumber="512">FIGURE 8.</emphasis>
<emphasis id="B9435F7CFFAFFFE0DC5522A10BEA96A2" box="[280,690,1946,1968]" italics="true" pageId="12" pageNumber="512">
<taxonomicName id="4C37F8EDFFAFFFE0DC5522A10B5796A2" box="[280,527,1946,1968]" class="Insecta" family="Baetidae" genus="Callibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="512" phylum="Arthropoda" rank="species" species="nigrivenosus">Callibaetis nigrivenosus</taxonomicName>
<emphasis id="B9435F7CFFAFFFE0DF5922A00BEA96A2" bold="true" box="[532,690,1946,1968]" italics="true" pageId="12" pageNumber="512">nomen dubium</emphasis>
</emphasis>
female imago (holotype). A. labels (Harvard); B. lateral view of head and thorax; C. lateral view of abdomen; D. left forewing; E. right forewing.
</paragraph>
</caption>
<subSubSection id="C32DD0E5FFAEFFE1DD8A25AC0B7891EA" pageId="13" pageNumber="513" type="materials_examined">
<paragraph id="8B88836EFFAEFFE1DD8A25AC0B7891EA" blockId="13.[151,1436,151,248]" pageId="13" pageNumber="513">
<materialsCitation id="3B5F8933FFAEFFE1DD8A25AC080391EA" ID-GBIF-Occurrence="1434647650" collectingDate="1917-06" country="Ecuador" elevation="1372" location="Material" municipality="Huigar" pageId="13" pageNumber="513" specimenCount="1" typeStatus="holotype">
<emphasis id="B9435F7CFFAEFFE1DD8A25AC08EB91A2" bold="true" box="[199,435,151,176]" pageId="13" pageNumber="513">
<location id="8EE8D5B5FFAEFFE1DD8A25AC087791A2" LSID="urn:lsid:plazi:treatment:039E3278FFAFFFE1DDDA25AC0B7891EA:8EE8D5B5FFAEFFE1DD8A25AC087791A2" box="[199,303,151,176]" country="Ecuador" municipality="Huigar" name="Material" pageId="13" pageNumber="513">Material</location>
examined.
</emphasis>
<taxonomicName id="4C37F8EDFFAEFFE1DCF025A30B9D91BD" box="[445,709,152,176]" class="Insecta" family="Baetidae" genus="Callibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="513" phylum="Arthropoda" rank="species" species="nigrivenosa">
<emphasis id="B9435F7CFFAEFFE1DCF025A30B9D91BD" box="[445,709,152,176]" italics="true" pageId="13" pageNumber="513">Callibaetis nigrivenosa</emphasis>
</taxonomicName>
, male imago (
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),
<collectingCountry id="F320C3FEFFAEFFE1DEF725AC0D4191A2" box="[954,1049,151,176]" name="Ecuador" pageId="13" pageNumber="513">Ecuador</collectingCountry>
,
<collectingMunicipality id="6BEC1914FFAEFFE1D96A25AC0D2F91A2" box="[1063,1143,151,176]" pageId="13" pageNumber="513">Huigar</collectingMunicipality>
,
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<quantity id="4CCF2E8BFFAEFFE1D9C925AC0DBB91A2" box="[1156,1251,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" pageId="13" pageNumber="513" unit="ft" value="4500.0">4,500 ft</quantity>
</elevation>
.,
<collectingDate id="EFCD5C46FFAEFFE1D9BB25A30C0A91BD" box="[1270,1362,152,176]" pageId="13" pageNumber="513" value="1917-06">
<date id="FF89A5AEFFAEFFE1D9BB25A30C0A91BD" box="[1270,1362,152,176]" pageId="13" pageNumber="513" value="1917-06">17 June</date>
</collectingDate>
, H. S. Parish coll., Museum of Comparative Zoology, Harvard University, MCZ-ENT 0 0 0 10057.
<emphasis id="B9435F7CFFAEFFE1DD8A25E4080391EA" bold="true" box="[199,347,223,248]" pageId="13" pageNumber="513">Distribution</emphasis>
</materialsCitation>
. Ecuador: Huigar
</paragraph>
</subSubSection>
</treatment>
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