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<document id="A75FD4A090BA21AEFB132174B514546B" ID-CLB-Dataset="120098" ID-DOI="10.11646/zootaxa.5174.3.3" ID-GBIF-Dataset="8ba8a760-9e80-400d-9287-123084a40ed7" ID-ISSN="1175-5326" ID-Zenodo-Dep="6986267" ID-ZooBank="E98F1B0F-702D-4CD4-B68E-055AE2024F5F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660325729645" checkinUser="plazi" docAuthor="Deltshev, Christo, Naumova, Maria, Matevski, Dragan &amp; Indzhov, Simeon" docDate="2022" docId="039B8783FFFFFFEAC38FBFC6FC04FE9B" docLanguage="en" docName="zootaxa.5174.3.3.pdf" docOrigin="Zootaxa 5174 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zodarion scutatum Wunderlich 1980" docType="treatment" docVersion="7" lastPageNumber="256" masterDocId="FFA2FFFBFFF7FFE3C318BF50FFE6FF86" masterDocTitle="New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species" masterLastPageNumber="261" masterPageNumber="247" pageNumber="255" updateTime="1699366682012" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E81152B3529D1D9DCA065007CCB72FED" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2F339B4E351397F6ECE271B8CB99A63A">New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species</mods:title>
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<mods:namePart id="0DC21198559E198BB84B897ADD8BD6EA">Deltshev, Christo</mods:namePart>
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<mods:affiliation id="580691CA6BB071D9DDDD214410B80BB5">deltshev @ gmail. com https: // orcid. org / 0000 - 0003 - 0539 - 5673</mods:affiliation>
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<mods:name id="C119AD99A9C5AE71514EFF7A964C2EB1" type="personal">
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<mods:namePart id="A978C8E687B79B98D8D2C449AB432874">Naumova, Maria</mods:namePart>
<mods:nameIdentifier id="119B8FBCD1C030A84CEFCEE5609DCE84" type="ORCID">0000-0003-0060-048X</mods:nameIdentifier>
<mods:affiliation id="034B61A01CFD550B3C8C034F952C00A3">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1, Tsar Osvoboditel blvd., 1000 Sofia, Bulgaria munnyto @ gmail. com; munny @ abv. bg; https: // orcid. org / 0000 - 0003 - 0060 - 048 X</mods:affiliation>
<mods:nameIdentifier id="0D081EE1B15AFD8F54B1653E74B6207B" type="email">munnyto@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C8F8AD650416150F9CB89EF20F64EA3C">Matevski, Dragan</mods:namePart>
<mods:nameIdentifier id="57B0DD91853402E10656D08B05D18405" type="ORCID">0000-0002-0973-8824</mods:nameIdentifier>
<mods:affiliation id="7FF6E7096E03E25BBFD1EF4B0C8C5C31">Institute of Biology, Faculty of Natural Sciences and Mathematics, Ss. Cyril and Methodius University, Arhimedova 3, P. O. Box 162, 1000 Skopje, North Macedonia. dragan. matevski @ hotmail. com; https: // orcid. org / 0000 - 0002 - 0973 - 8824</mods:affiliation>
<mods:nameIdentifier id="AD22A6E10A2DEBCE8BD738FA9EA227E5" type="email">dragan.matevski@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="37C0FEF17FB8D6BEAF0BE7CD14238CFE">Indzhov, Simeon</mods:namePart>
<mods:nameIdentifier id="C12139D8CBA0D2CBB5AD564A5D11DA47" type="ORCID">0000-0002-8937-9373</mods:nameIdentifier>
<mods:affiliation id="DBE1F63A68EFD1E8B264ED5CA433B9DB">Independent researcher, Sofia, Bulgaria. saindzhov @ gmail. com; https: // orcid. org / 0000 - 0002 - 8937 - 9373</mods:affiliation>
<mods:nameIdentifier id="ED6C4A202EA3759E9A866D17BB906EB6" type="email">saindzhov@gmail.com</mods:nameIdentifier>
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<mods:title id="A659529837484BAD13162EC93C1E1AAA">Zootaxa</mods:title>
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<mods:date id="430894FCDB67698BF28887C0A32ADFD1">2022</mods:date>
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<treatment id="039B8783FFFFFFEAC38FBFC6FC04FE9B" ID-DOI="http://doi.org/10.5281/zenodo.6989285" ID-GBIF-Taxon="199014920" ID-Zenodo-Dep="6989285" LSID="urn:lsid:plazi:treatment:039B8783FFFFFFEAC38FBFC6FC04FE9B" httpUri="http://treatment.plazi.org/id/039B8783FFFFFFEAC38FBFC6FC04FE9B" lastPageId="9" lastPageNumber="256" pageId="8" pageNumber="255">
<subSubSection id="C328651EFFFFFFEBC38FBFC6FDA8FF36" box="[151,590,150,177]" pageId="8" pageNumber="255" type="nomenclature">
<paragraph id="8B8D3695FFFFFFEBC38FBFC6FDA8FF36" blockId="8.[151,590,150,213]" box="[151,590,150,177]" pageId="8" pageNumber="255">
<heading id="D0C581F9FFFFFFEBC38FBFC6FDA8FF36" bold="true" box="[151,590,150,177]" fontSize="11" level="1" pageId="8" pageNumber="255" reason="1">
<taxonomicName id="4C324D16FFFFFFEBC38FBFC6FDA8FF36" ID-CoL="5D6D7" authority="Wunderlich, 1980" authorityName="Wunderlich" authorityYear="1980" box="[151,590,150,177]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="255" phylum="Arthropoda" rank="species" species="scutatum">
<emphasis id="B946EA87FFFFFFEBC38FBFC6FDA8FF36" bold="true" box="[151,590,150,177]" pageId="8" pageNumber="255">
<emphasis id="B946EA87FFFFFFEBC38FBFC6FE92FF37" bold="true" box="[151,372,150,177]" italics="true" pageId="8" pageNumber="255">Zodarion scutatum</emphasis>
<bibRefCitation id="EFA34B64FFFFFFEBC262BFC6FDA8FF36" author="Wunderlich, J." box="[378,590,150,176]" pageId="8" pageNumber="255" pagination="229 - 240" refId="ref7970" refString="Wunderlich, J. (1980) Zur Gattung Zodarion Walckenaer 1847 mit Neubeschreibungen (Arachnida: Araneae: Zodariidae). Senckenbergiana Biologica, 60, 229 - 240." type="journal article" year="1980">Wunderlich, 1980</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C328651EFFFFFFEBC38FBFEAFEFAFF53" box="[151,284,186,213]" pageId="8" pageNumber="255" type="description">
<paragraph id="8B8D3695FFFFFFEBC38FBFEAFEFAFF53" blockId="8.[151,590,150,213]" box="[151,284,186,213]" pageId="8" pageNumber="255">
<figureCitation id="13092A10FFFFFFEBC38FBFEAFEFEFF53" box="[151,280,186,213]" captionStart="FIGURES 57–60" captionStartId="8.[151,264,1967,1991]" captionTargetBox="[154,1417,445,1924]" captionTargetId="figure-79@8.[151,1435,442,1944]" captionTargetPageId="8" captionText="FIGURES 57–60. Z. scutatum, male: habitus (57, 58), and palp (59, 60); dorsal (57), ventral (58, 59) and retrolateral (60) views. Scales: 0.6 mm (57, 58), 0.2 mm (59, 60)." figureDoi="http://doi.org/10.5281/zenodo.6986279" httpUri="https://zenodo.org/record/6986279/files/figure.png" pageId="8" pageNumber="255">Figs 57–60</figureCitation>
.
</paragraph>
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<subSubSection id="C328651EFFFFFFEBC38FBFAFFD1FFEB3" pageId="8" pageNumber="255" type="reference_group">
<paragraph id="8B8D3695FFFFFFEBC38FBFAFFD05FE91" blockId="8.[151,761,255,309]" box="[151,739,255,279]" pageId="8" pageNumber="255">
<treatmentCitationGroup id="AB2211BBFFFFFFEBC38FBFAFFD05FE91" box="[151,739,255,279]" pageId="8" pageNumber="255">
<treatmentCitation id="0A931084FFFFFFEBC38FBFAFFDA1FE91" author="Wunderlich, J." box="[151,583,255,279]" page="113" pageId="8" pageNumber="255" year="1980">
<taxonomicName id="4C324D16FFFFFFEBC38FBFAFFDA1FE91" ID-CoL="5D6D7" authority="Wunderlich, 1980: 113" authorityName="Wunderlich" authorityPageNumber="113" authorityYear="1980" box="[151,583,255,279]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="255" phylum="Arthropoda" rank="species" species="scutatum">
<emphasis id="B946EA87FFFFFFEBC38FBFAFFEB1FE90" box="[151,343,255,279]" italics="true" pageId="8" pageNumber="255">Zodarion scutatum</emphasis>
<bibRefCitation id="EFA34B64FFFFFFEBC245BFAFFDA1FE91" author="Wunderlich, J." box="[349,583,255,279]" pageId="8" pageNumber="255" pagination="229 - 240" refId="ref7970" refString="Wunderlich, J. (1980) Zur Gattung Zodarion Walckenaer 1847 mit Neubeschreibungen (Arachnida: Araneae: Zodariidae). Senckenbergiana Biologica, 60, 229 - 240." type="journal article" year="1980">Wunderlich, 1980: 113</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, f. 1–6 (D
<specimenCount id="9D34FD1CFFFFFFEBC1ABBFAFFD2EFE91" box="[691,712,255,279]" pageId="8" pageNumber="255" type="male">♂</specimenCount>
<specimenCount id="9D34FD1CFFFFFFEBC1D0BFAFFD31FE91" box="[712,727,255,279]" pageId="8" pageNumber="255" type="female">♀</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8D3695FFFFFFEBC38FBE4DFD1FFEB3" blockId="8.[151,761,255,309]" box="[151,761,285,309]" pageId="8" pageNumber="255">
<treatmentCitationGroup id="AB2211BBFFFFFFEBC38FBE4DFD1FFEB3" box="[151,761,285,309]" pageId="8" pageNumber="255">
<treatmentCitation id="0A931084FFFFFFEBC38FBE4DFDD6FEB3" author="Bosmans, R." box="[151,560,285,309]" page="226" pageId="8" pageNumber="255" year="2009">
<taxonomicName id="4C324D16FFFFFFEBC38FBE4DFDD6FEB3" ID-CoL="5D6D7" authority=": Bosmans 2009: 226" authorityName=": Bosmans" authorityPageNumber="226" authorityYear="2009" box="[151,560,285,309]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="255" phylum="Arthropoda" rank="species" species="scutatum">
<emphasis id="B946EA87FFFFFFEBC38FBE4DFEB1FEB3" box="[151,343,285,309]" italics="true" pageId="8" pageNumber="255">Zodarion scutatum</emphasis>
:
<bibRefCitation id="EFA34B64FFFFFFEBC27DBE4EFDD6FEB3" author="Bosmans, R." box="[357,560,285,309]" pageId="8" pageNumber="255" pagination="211 - 295" refId="ref6323" refString="Bosmans, R. (2009) Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History, 12, 211 - 295." type="journal article" year="2009">Bosmans 2009: 226</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, f. 18–19, 53 (
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<specimenCount id="9D34FD1CFFFFFFEBC1C3BE4DFD0DFEB3" box="[731,747,285,309]" pageId="8" pageNumber="255" type="female">♀</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C328651EFFFFFFEBC38FBE33FAF2FE27" pageId="8" pageNumber="255" type="materials_examined">
<paragraph id="8B8D3695FFFFFFEBC38FBE33FAF2FE27" blockId="8.[151,1437,355,418]" pageId="8" pageNumber="255">
<emphasis id="B946EA87FFFFFFEBC38FBE33FE62FEFB" bold="true" box="[151,388,355,381]" pageId="8" pageNumber="255">Material examined.</emphasis>
<materialsCitation id="3B5A3CC8FFFFFFEBC297BE34FAF2FE27" ID-GBIF-Occurrence="3887778307" collectingDate="2003-07-18" collectionCode="NMNHS" collectorName="S. Lazarov &amp; M. Langourov" country="Serbia" county="Tepca village" elevation="588" latitude="43.2184" location="Durmitor Mts" longLatPrecision="6" longitude="19.0803" municipality="Devojacki Most" pageId="8" pageNumber="255" specimenCount="10" specimenCount-male="10">
<specimenCount id="9D34FD1CFFFFFFEBC297BE34FE26FEF8" box="[399,448,356,382]" pageId="8" pageNumber="255" type="male">10♂</specimenCount>
(
<collectionCode id="ED23AE50FFFFFFEBC2CBBE34FDD8FEF8" box="[467,574,356,382]" pageId="8" pageNumber="255">NMNHS</collectionCode>
),
<collectingCountry id="F3257605FFFFFFEBC14CBE34FCFEFEF8" box="[596,792,356,382]" name="Serbia" pageId="8" pageNumber="255">MONTENEGRO</collectingCountry>
:
<collectorName id="26C75343FFFFFFEBC033BE34FC1CFEF8" box="[811,1018,355,382]" pageId="8" pageNumber="255">
NP
<location id="8EED604EFFFFFFEBC041BE33FC1CFEF8" LSID="urn:lsid:plazi:treatment:039B8783FFFFFFEAC38FBFC6FC04FE9B:8EED604EFFFFFFEBC041BE33FC1CFEF8" box="[857,1018,355,382]" country="Serbia" county="Tepca village" latitude="43.2184" longLatPrecision="6" longitude="19.0803" municipality="Devojacki Most" name="Durmitor Mts" pageId="8" pageNumber="255">Durmitor Mts</location>
</collectorName>
,
<location id="8EED604EFFFFFFEBC710BE34FB30FEF8" LSID="urn:lsid:plazi:treatment:039B8783FFFFFFEAC38FBFC6FC04FE9B:8EED604EFFFFFFEBC710BE34FB30FEF8" box="[1032,1238,355,382]" country="Serbia" county="Tepca village" latitude="43.2184" longLatPrecision="6" longitude="19.0803" municipality="Devojacki Most" name="Tara River Gorge" pageId="8" pageNumber="255">
<location id="8EED604EFFFFFFEBC710BE34FB65FEFB" LSID="urn:lsid:plazi:treatment:039B8783FFFFFFEAC38FBFC6FC04FE9B:8EED604EFFFFFFEBC710BE34FB65FEFB" box="[1032,1155,355,382]" country="Serbia" county="Tepca village" latitude="43.2184" longLatPrecision="6" longitude="19.0803" municipality="Devojacki Most" name="Tara River" pageId="8" pageNumber="255">
Tara
<location id="8EED604EFFFFFFEBC75DBE33FB65FEFB" LSID="urn:lsid:plazi:treatment:039B8783FFFFFFEAC38FBFC6FC04FE9B:8EED604EFFFFFFEBC75DBE33FB65FEFB" box="[1093,1155,355,381]" country="Serbia" county="Tepca village" latitude="43.2184" longLatPrecision="6" longitude="19.0803" municipality="Devojacki Most" name="River Gorge" pageId="8" pageNumber="255">River</location>
</location>
Gorge
</location>
,
<collectingMunicipality id="6BE9ACEFFFFFFFEBC7FDBE33FA7BFEF8" box="[1253,1437,355,382]" pageId="8" pageNumber="255">Devojacki Most</collectingMunicipality>
bridge,
<collectingCounty id="62EC4E19FFFFFFEBC3F4BED8FE63FE27" box="[236,389,391,418]" pageId="8" pageNumber="255">Tepća village</collectingCounty>
,
<geoCoordinate id="EE065052FFFFFFEBC288BED7FDE3FE24" box="[400,517,391,418]" degrees="43.2184" direction="north" orientation="latitude" pageId="8" pageNumber="255" precision="5" value="43.2184">N43.2184</geoCoordinate>
˚,
<geoCoordinate id="EE065052FFFFFFEBC10EBED7FD6FFE24" box="[534,649,391,418]" degrees="19.0803" direction="east" orientation="longitude" pageId="8" pageNumber="255" precision="5" value="19.0803">E19.0803</geoCoordinate>
˚,
<quantity id="4CCA9B70FFFFFFEBC182BED7FD05FE24" box="[666,739,391,418]" metricMagnitude="2" metricUnit="m" metricValue="5.88" pageId="8" pageNumber="255" unit="m" value="588.0">
<elevation id="001FD1A6FFFFFFEBC182BED7FD05FE24" box="[666,739,391,418]" metricMagnitude="2" metricUnit="m" metricValue="5.88" pageId="8" pageNumber="255" unit="m" value="588.0">588 m</elevation>
</quantity>
, 13–
<date id="FF8C1055FFFFFFEBC00FBED8FC7CFE24" box="[791,922,392,418]" pageId="8" pageNumber="255" value="2003-07-18">
<collectingDate id="EFC8E9BDFFFFFFEBC00FBED8FC7CFE24" box="[791,922,392,418]" pageId="8" pageNumber="255" value="2003-07-18">18.07.2003</collectingDate>
</date>
,
<collectorName id="26C75343FFFFFFEBC0BDBED8FBF8FE24" box="[933,1054,392,418]" pageId="8" pageNumber="255">S. Lazarov</collectorName>
&amp;
<collectorName id="26C75343FFFFFFEBC75ABED8FB02FE24" box="[1090,1252,392,418]" pageId="8" pageNumber="255">M. Langourov</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF4D661DFFFFFFEBC38FB8FFFDA7F86D" ID-DOI="http://doi.org/10.5281/zenodo.6986279" ID-Zenodo-Dep="6986279" httpUri="https://zenodo.org/record/6986279/files/figure.png" pageId="8" pageNumber="255" startId="8.[151,264,1967,1991]" targetBox="[154,1417,445,1924]" targetPageId="8">
<paragraph id="8B8D3695FFFFFFEBC38FB8FFFDA7F86D" blockId="8.[151,1437,1967,2028]" pageId="8" pageNumber="255">
<emphasis id="B946EA87FFFFFFEBC38FB8FFFEB5F841" bold="true" box="[151,339,1967,1991]" pageId="8" pageNumber="255">FIGURES 57–60.</emphasis>
<taxonomicName id="4C324D16FFFFFFEBC240B8E0FE2AF841" authorityName=": Bosmans" authorityYear="2009" box="[344,460,1968,1991]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="255" phylum="Arthropoda" rank="species" species="scutatum">
<emphasis id="B946EA87FFFFFFEBC240B8E0FE2AF841" box="[344,460,1968,1991]" italics="true" pageId="8" pageNumber="255">Z. scutatum</emphasis>
</taxonomicName>
, male: habitus (57, 58), and palp (59, 60); dorsal (57), ventral (58, 59) and retrolateral (60) views. Scales: 0.6 mm (57, 58), 0.2 mm (59, 60).
</paragraph>
</caption>
<subSubSection id="C328651EFFFEFFEAC3DFBFC6FC04FE9B" pageId="9" pageNumber="256" type="distribution">
<paragraph id="8B8D3695FFFEFFEAC3DFBFC6FBFBFF37" blockId="9.[151,1436,150,285]" box="[199,1053,150,177]" pageId="9" pageNumber="256">
<emphasis id="B946EA87FFFEFFEAC3DFBFC6FE87FF36" bold="true" box="[199,353,150,176]" pageId="9" pageNumber="256">Distribution.</emphasis>
<collectingCountry id="F3257605FFFEFFEAC27FBFC7FE5AFF37" box="[359,444,151,177]" name="Croatia" pageId="9" pageNumber="256">Croatia</collectingCountry>
,
<collectingCountry id="F3257605FFFEFFEAC2D0BFC7FDCCFF37" box="[456,554,151,177]" name="Slovenia" pageId="9" pageNumber="256">Slovenia</collectingCountry>
(WSC 2022) and
<collectingCountry id="F3257605FFFEFFEAC1EEBFC7FC99FF37" box="[758,895,151,177]" name="Serbia" pageId="9" pageNumber="256">Montenegro</collectingCountry>
(new record).
</paragraph>
<paragraph id="8B8D3695FFFEFFEAC3DFBFEBFC04FE9B" blockId="9.[151,1436,150,285]" pageId="9" pageNumber="256">
<emphasis id="B946EA87FFFEFFEAC3DFBFEBFEDDFF53" bold="true" box="[199,315,187,213]" pageId="9" pageNumber="256">Remarks.</emphasis>
This is the first report of this species from
<collectingCountry id="F3257605FFFEFFEAC023BFEBFC22FF53" box="[827,964,187,213]" name="Serbia" pageId="9" pageNumber="256">Montenegro</collectingCountry>
(
<figureCitation id="13092A10FFFEFFEAC0CEBFEBFBCBFF53" box="[982,1069,187,213]" captionStart="FIGURE 76" captionStartId="12.[151,250,1175,1199]" captionTargetBox="[160,1427,453,1152]" captionTargetId="figure-127@12.[160,1427,453,1152]" captionTargetPageId="12" captionText="FIGURE 76. Georeferenced locations of the new records of Zodarion species in the Balkans." figureDoi="http://doi.org/10.5281/zenodo.6986289" httpUri="https://zenodo.org/record/6986289/files/figure.png" pageId="9" pageNumber="256">Fig. 76</figureCitation>
). The examined specimen (
<figureCitation id="13092A10FFFEFFEAC672BFEBFF39FF7F" captionStart="FIGURES 57–60" captionStartId="8.[151,264,1967,1991]" captionTargetBox="[154,1417,445,1924]" captionTargetId="figure-79@8.[151,1435,442,1944]" captionTargetPageId="8" captionText="FIGURES 57–60. Z. scutatum, male: habitus (57, 58), and palp (59, 60); dorsal (57), ventral (58, 59) and retrolateral (60) views. Scales: 0.6 mm (57, 58), 0.2 mm (59, 60)." figureDoi="http://doi.org/10.5281/zenodo.6986279" httpUri="https://zenodo.org/record/6986279/files/figure.png" pageId="9" pageNumber="256">Figs 57–60</figureCitation>
) conforms well to the description and diagnosis of the species as provided in
<bibRefCitation id="EFA34B64FFFEFFEAC742BF8FFAD7FF7F" author="Wunderlich, J." box="[1114,1329,223,249]" pageId="9" pageNumber="256" pagination="229 - 240" refId="ref7970" refString="Wunderlich, J. (1980) Zur Gattung Zodarion Walckenaer 1847 mit Neubeschreibungen (Arachnida: Araneae: Zodariidae). Senckenbergiana Biologica, 60, 229 - 240." type="journal article" year="1980">Wunderlich (1980)</bibRefCitation>
and with photos of the
<typeStatus id="54898837FFFEFFEAC228BE53FE74FE9B" box="[304,402,259,285]" pageId="9" pageNumber="256" type="holotype">holotype</typeStatus>
kindly provided by László Mezőfi and Tamás Szűts.
</paragraph>
</subSubSection>
</treatment>
</document>
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