262 lines
30 KiB
XML
262 lines
30 KiB
XML
<document ID-CLB-Dataset="9342" ID-DOI="http://dx.doi.org/10.3897/zookeys.1003.55541" ID-GBIF-Dataset="a157aae8-b7dd-40f9-8180-8ed7ae225903" ID-PMC="PMC7752891" ID-Pensoft-Pub="1313-2970-1003-31" ID-Pensoft-UUID="279B98048C245FF5AC90D9CADA7E8C17" ID-PubMed="33384561" ID-ZooBank="D4A53AD77D9E42789277E931291AD4EC" ModsDocID="1313-2970-1003-31" checkinTime="1607975026353" checkinUser="pensoft" docAuthor="Kodada, Jan, Jaech, Manfred A., Freitag, Hendrik, Ciamporova-Zatovicova, Zuzana, Goffova, Katarina, Selnekovic, David & Jr, Fedor Ciampor" docDate="2020" docId="ACEF8DE161EA5B979C88CBF774952C80" docLanguage="en" docName="ZooKeys 1003: 31-55" docOrigin="ZooKeys 1003" docPubDate="2020-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1003.55541" docTitle="Ancyronyx pulcherrimus Kodada, Jaech & Ciampor 2014" docType="treatment" docVersion="7" id="279B98048C245FF5AC90D9CADA7E8C17" lastPageNumber="31" masterDocId="279B98048C245FF5AC90D9CADA7E8C17" masterDocTitle="Ancyronyx lianlabangorum sp. nov., a new spider riffle beetle from Sarawak, and new distribution records for A. pulcherrimus Kodada, Jaech & Ciampor based on DNA barcodes (Coleoptera, Elmidae)" masterLastPageNumber="55" masterPageNumber="31" pageNumber="31" updateTime="1732840719915" updateUser="ExternalLinkService">
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<mods:title id="048B20E6EADFA67BCBB913DA1A0B7B15">Ancyronyx lianlabangorum sp. nov., a new spider riffle beetle from Sarawak, and new distribution records for A. pulcherrimus Kodada, Jaech & Ciampor based on DNA barcodes (Coleoptera, Elmidae)</mods:title>
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<mods:namePart id="3D9E9D8537DB3A85A544B3F494FF1B13">Kodada, Jan</mods:namePart>
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<mods:nameIdentifier id="66097BE8D23152CF9EED09084113BE9C" type="ORCID">https://orcid.org/0000-0002-1355-4323</mods:nameIdentifier>
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<mods:affiliation id="DAA37FF269CE72A0298C00CB9FEE8988">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina B- 1, SK- 842 15, Bratislava, Slovakia</mods:affiliation>
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<mods:nameIdentifier id="06340414BA1D55F04B5942D7A30FB0AD" type="email">jan.kodada@uniba.sk</mods:nameIdentifier>
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<mods:namePart id="D592A5C31672558C01679DEAA2F8050F">Jaech, Manfred A.</mods:namePart>
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<mods:affiliation id="303E779A258A5D99C2DB2C777F1CC7C0">Naturhistorisches Museum Wien, Burgring 7, A- 1010, Wien, Austria</mods:affiliation>
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<mods:namePart id="7C03438D69A8A398388618701E95BDF0">Freitag, Hendrik</mods:namePart>
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<mods:nameIdentifier id="744551B7E0972124B83C23D22B593893" type="ORCID">https://orcid.org/0000-0002-1325-0979</mods:nameIdentifier>
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<mods:affiliation id="E390C03730E0B97D0FA53CB237473B92">Ateneo de Manila University, Biology Department, School of Science and Engineering, Loyola Heights, Quezon City, 1108, Philippines & Universiti Brunei Darussalam, Environmental and Life Sciences, Faculty of Science, Jalan Tungku Link, Gadong, BE 1410, Brunei & Museum fuer Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, Berlin, 10115, Germany</mods:affiliation>
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<mods:namePart id="F5E31F9368BC72CBCF879025303E9261">Ciamporova-Zatovicova, Zuzana</mods:namePart>
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<mods:affiliation id="4DF215C81E44E42750BBA6A7DE5E18AB">Zoology Lab, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dubravska cesta 9, SK- 84523, Bratislava, Slovakia</mods:affiliation>
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<mods:namePart id="EE857D37ECE532150643A99084CDA38D">Goffova, Katarina</mods:namePart>
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<mods:affiliation id="9A314381892BF98F7E16102E8542921F">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina B- 1, SK- 842 15, Bratislava, Slovakia</mods:affiliation>
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<mods:namePart id="E11135C0F3ACD252FD0229CDF2BA139C">Selnekovic, David</mods:namePart>
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<mods:nameIdentifier id="0699028CCD3E01FF0973EBAFA40B89DB" type="ORCID">https://orcid.org/0000-0002-9228-1174</mods:nameIdentifier>
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<mods:affiliation id="17224C8B2D04AF80AB2E833474241C32">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina B- 1, SK- 842 15, Bratislava, Slovakia</mods:affiliation>
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<mods:namePart id="FE3ED049F71321B8D8438394996FB549">Jr, Fedor Ciampor</mods:namePart>
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<mods:affiliation id="EB695B282D894C9BA0CADF5F26EE64F6">Zoology Lab, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dubravska cesta 9, SK- 84523, Bratislava, Slovakia</mods:affiliation>
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<mods:date id="71519270964DA69AA2581876CFADA04F">2020</mods:date>
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<mods:number id="0AC1C313D4735649868BB1E8E1A9C881">2020-12-14</mods:number>
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<mods:number id="B43C3E1136D3F2963BB0D5DC0FABB235">1003</mods:number>
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<mods:start id="77F904C02CD48295EFFD4BF25B67F291">31</mods:start>
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<mods:url id="8CE3199E177021A01EB331C14BDEEF53">http://dx.doi.org/10.3897/zookeys.1003.55541</mods:url>
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<mods:classification id="70C50394EF893F3E1129FE1EE13537A6">journal article</mods:classification>
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<mods:identifier id="B5700EBFF86C2CB98201576331A612B5" type="DOI">http://dx.doi.org/10.3897/zookeys.1003.55541</mods:identifier>
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<mods:identifier id="CC4FFDE8340AD10C11FBD366E3702D6D" type="Pensoft-Pub">1313-2970-1003-31</mods:identifier>
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<subSection id="5189810B5799DFCB22D052318BC7C76F" pageId="0" pageNumber="31" type="results">
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<treatment id="ACEF8DE161EA5B979C88CBF774952C80" ID-GBIF-Taxon="174795567" LSID="urn:lsid:plazi:treatment:ACEF8DE161EA5B979C88CBF774952C80" httpUri="http://treatment.plazi.org/id/ACEF8DE161EA5B979C88CBF774952C80" lastPageNumber="31" pageId="0" pageNumber="31" scope_family="Elmidae" scope_order="Coleoptera">
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<subSubSection id="A27390D4D463DF66B0DC77A5FDAB5E98" pageId="0" pageNumber="31" type="nomenclature">
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<paragraph id="2DA6EBFCB4FB9C67999A8CCE69EA2EBF" pageId="0" pageNumber="31">
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<taxonomicName id="D6C104495FEBB64913B9283B617D7B5A" ID-CoL="DPSY" LSID="ACEF8DE1-61EA-5B97-9C88-CBF774952C80" authority="Kodada, Jaech & Ciampor, 2014" authorityName="Kodada, Jaech & Ciampor" authorityYear="2014" class="Insecta" family="Elmidae" genus="Ancyronyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancyronyx pulcherrimus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="pulcherrimus">
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Ancyronyx pulcherrimus Kodada,
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<normalizedToken id="C537FE7C244D2729440A4224217AA769" originalValue="Jäch">Jaech</normalizedToken>
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&
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<normalizedToken id="4AAB754196AD6AB094E5B4E65DFB6ED4" originalValue="Čiampor">Ciampor</normalizedToken>
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, 2014
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="5F2E642664163D67A7AD33E794B0A96E" pageId="0" pageNumber="31" type="type locality">
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<paragraph id="E25AF46E0DB2B066EDAA02553C01E3BF" pageId="0" pageNumber="31">Type locality.</paragraph>
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<paragraph id="D09EF64F970A8A8101F45C9FDB3010BA" pageId="0" pageNumber="31">Ingei River, forest pool,, Belait District, Brunei.</paragraph>
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</subSubSection>
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<subSubSection id="72040597302BB59D281E8B1025A14D5E" pageId="0" pageNumber="31" type="materials_examined">
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<paragraph id="F2D46150F944DF4FAC25D9DDBCB570FF" pageId="0" pageNumber="31">Material examined.</paragraph>
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<paragraph id="CEE2476D7171A127BE7F4AC1B3961A44" pageId="0" pageNumber="31">
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<emphasis id="DD9BC423DC50FA2BD6B7F37CCFE142DE" bold="true" pageId="0" pageNumber="31">Type material</emphasis>
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.
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<emphasis id="C57AAD18C42646D6FA48D9F4E93AE0C0" bold="true" italics="true" pageId="0" pageNumber="31">Holotype</emphasis>
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♂ (BM): "Brunei: Belait Distr. Sg. Ingei, forest pool near Base Camp, 14.VI.2010 leg. Mayyer Ling (34)".
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<emphasis id="CCF2FE5C8DC72E6DC1DDF87AE9DED1E2" bold="true" italics="true" pageId="0" pageNumber="31">Paratypes</emphasis>
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(NMW, UB): 1 ♂ and 2 ♀♀, same label data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="12D499FFFE4EF10495C7D64E15027568" pageId="0" pageNumber="31" type="materials_examined">
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<paragraph id="D8DBF010D9AEB42CA2CE5F9E5D10494B" pageId="0" pageNumber="31">Additional material</paragraph>
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<paragraph id="3813CA15C4A8FB848F7199ABE9B88808" pageId="0" pageNumber="31">(CCB, CFDS, CKB, NMW, UBDM).</paragraph>
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<paragraph id="534FCCE9BB82C2C4DF75655055B0323F" pageId="0" pageNumber="31">
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<emphasis id="98520F4521202331CA68D755C229390B" bold="true" pageId="0" pageNumber="31">Lineage 1. Sarawak</emphasis>
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9 ♂♂ [incl. JK100, JK213, JK214, JK215], 15 ♀♀ [incl. JK101, JK102, JK208, FZ1632]: "Malaysia, Sarawak, Marudi distr., Gunung Mulu NP, 16.10.2018, (40)
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<geoCoordinate id="16DF2B15EB5B8EC28B54F915DB7D456D" degrees="04.0267" direction="north" orientation="latitude" precision="5" value="4.0267">04.0267°N</geoCoordinate>
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,
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<geoCoordinate id="A8065F304EAC5E1C2D15A08618C7C132" degrees="114.8234" direction="east" orientation="longitude" precision="5" value="114.8234">114.8234°E</geoCoordinate>
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, 55 m a.s.l., small stream in primary forest, Kodada &
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<normalizedToken id="F42071FFA598CF2E4211F53B45096ECE" originalValue="Selnekovič">Selnekovic</normalizedToken>
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lgt.".
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<emphasis id="3473080696A3B952E124F219B838443B" bold="true" pageId="0" pageNumber="31">Brunei</emphasis>
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2 ♂♂, 1 ♀ [FR324]: "Brunei: Temburong; small Sungai Peradayan; rivulet, subm. wood; dist. primary forest; c. 25m a.s.l. c. 04.11.1997 leg. Mendoza (6f)".
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</paragraph>
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<paragraph id="6588D3C1012D11D8061132E12A8ABF3B" pageId="0" pageNumber="31">
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<emphasis id="9962D8C9BBE80AE2664F967F4453A526" bold="true" pageId="0" pageNumber="31">Lineages 2 and 3. Sarawak</emphasis>
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5 ♂♂ [incl. JK105, JK106], 1 ♀ [JK12]: "Malaysia, Sarawak, Kuching distr., Bayur riv. near Kampung Bayur, 20.10.2018,
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<geoCoordinate id="5E14E04ECDF583F9325C6B4E00BBA6F6" degrees="1" direction="north" minutes="14" orientation="latitude" precision="1" seconds="42.3" value="1.2450833">1°14'42.3"N</geoCoordinate>
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,
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<geoCoordinate id="D136BC0BC02C1D9FBD5A91A23F56797D" degrees="110" direction="east" minutes="17" orientation="longitude" precision="1" seconds="35.3" value="110.29314">110°17'35.3"E</geoCoordinate>
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40 m a.s.l., J. Kodada & D.
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<normalizedToken id="89FD65D5056BDEABF5974F28E026EF6B" originalValue="Selnekovič">Selnekovic</normalizedToken>
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lgt."; 2 ♂♂ [JK197, JK199], 4 ♀♀ [incl. JK193, JK195]: same locality and collectors, but 22.02.2019, (No.43); 1 ♂ [FR344]: "Malaysia: Sarawak, Kg. Bayur 35km SSE Kuching, small hill river Bayur R.; run, subm. wood; second. veget./farmland; c. 50m a.s.l., c. 12.11.1993 leg. Mendoza (19f)".
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</paragraph>
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</subSubSection>
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<subSubSection id="5C323A1BED8579588392C7B06763E0B2" pageId="0" pageNumber="31" type="variability">
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<paragraph id="30308464612599EA8DB8CA9C73423D89" pageId="0" pageNumber="31">Variability.</paragraph>
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<paragraph id="046D91BDE1DD998D26EA328F70DA220F" pageId="0" pageNumber="31">
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<taxonomicName id="7DB7BBA95FFB05A56778BC09B1F5A7EC" authorityName="Kodada, Jaech & Ciampor" authorityYear="2014" class="Insecta" family="Elmidae" genus="Ancyronyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancyronyx pulcherrimus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="pulcherrimus">
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<emphasis id="BDEEF6B40F1FF5C9271F172F84FA6F08" italics="true" pageId="0" pageNumber="31">Ancyronyx pulcherrimus</emphasis>
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</taxonomicName>
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was represented by three barcode lineages, which are tentatively considered conspecific in the absence of further evidence. The specimens of the first lineage were collected in northern Sarawak and Brunei, while the specimens of the second and third lineages occur sympatrically in southwestern Sarawak. The first lineage is characterized by: (1) elytra predominantly black with a central yellowish, nearly X-shaped pattern and yellowish apices; (2) femora with black band between middle and proximal fourth; and (3) tibiae black on proximal three-quarters (compare Fig.
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<figureCitation id="835BDE54C3E231DD0A4149004890BA18" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ancyronyx pulcherrimus A male from Gunung Mulu NP, body length 1.82 mm B brachypterous male from Gunung Mulu NP, dorsal view (left elytron and apical segments of abdomen removed), body length 1.92 mm C male from Bayur River, dorsal view, body length 1.70 mm D male [JK 106] from Bayur River, dorsal view, body length 1.70 mm." figureDoi="10.3897/zookeys.1003.55541.figure4" httpUri="https://binary.pensoft.net/fig/486555" pageId="0" pageNumber="31">4</figureCitation>
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in
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<bibRefCitation id="EB013355EE35BDD15427B2F8A022C74E" DOI="https://doi.org/10.11646/zootaxa.3760.3.5" author="Kodada, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="31" pagination="383 - 395" refId="B18" refString="Kodada, J, Jaech, MA, Ciampor, Jr F, 2014. Ancyronyx reticulatus and A. pulcherrimus, two new riffle beetle species from Borneo, and discussion about elmid plastron structures (Coleoptera: Elmidae). Zootaxa 3760 (3): 383 - 395, DOI: https://doi.org/10.11646/zootaxa.3760.3.5" title="Ancyronyx reticulatus and A. pulcherrimus, two new riffle beetle species from Borneo, and discussion about elmid plastron structures (Coleoptera: Elmidae)." url="https://doi.org/10.11646/zootaxa.3760.3.5" volume="3760" year="2014">Kodada et al. 2014</bibRefCitation>
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). The type specimens are rather uniform in these characters, and all individuals recently collected in Gunung Mulu NP generally agree with them. However, there is some minor variation in the extent of the black coloration of the elytra, pronotum and legs (Fig.
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<figureCitation id="EB6FEB8301DE9D79FBB3355BC94E6C50" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ancyronyx pulcherrimus A male from Gunung Mulu NP, body length 1.82 mm B brachypterous male from Gunung Mulu NP, dorsal view (left elytron and apical segments of abdomen removed), body length 1.92 mm C male from Bayur River, dorsal view, body length 1.70 mm D male [JK 106] from Bayur River, dorsal view, body length 1.70 mm." figureDoi="10.3897/zookeys.1003.55541.figure4" httpUri="https://binary.pensoft.net/fig/486555" pageId="0" pageNumber="31">4A, B</figureCitation>
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). The examination of the hind wings of six specimens of the Gunung Mulu population and three specimens from Temburong (Brunei) revealed only short wings not reaching the abdominal apex (Fig.
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<figureCitation id="C2F6DCDD09AAC952850B24105175FF98" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ancyronyx pulcherrimus A male from Gunung Mulu NP, body length 1.82 mm B brachypterous male from Gunung Mulu NP, dorsal view (left elytron and apical segments of abdomen removed), body length 1.92 mm C male from Bayur River, dorsal view, body length 1.70 mm D male [JK 106] from Bayur River, dorsal view, body length 1.70 mm." figureDoi="10.3897/zookeys.1003.55541.figure4" httpUri="https://binary.pensoft.net/fig/486555" pageId="0" pageNumber="31">4B</figureCitation>
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). Macropterous or apterous forms were not detected, and the elytral shape of the remaining specimens suggests that they are flightless too.
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</paragraph>
|
|
<paragraph id="91B9C9FDF552F83D28B2CF3101A38754" pageId="0" pageNumber="31">
|
|
The specimens of the second and third lineages differ from the type specimens in the predominantly yellowish elytra, which are provided with several black lateral and sutural spots of variable size. The pronotal spots are usually smaller, and the dark femoral bands are smaller, paler or obsolete (Fig.
|
|
<figureCitation id="D9F941FFB78378FEBAF64612B55716B9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ancyronyx pulcherrimus A male from Gunung Mulu NP, body length 1.82 mm B brachypterous male from Gunung Mulu NP, dorsal view (left elytron and apical segments of abdomen removed), body length 1.92 mm C male from Bayur River, dorsal view, body length 1.70 mm D male [JK 106] from Bayur River, dorsal view, body length 1.70 mm." figureDoi="10.3897/zookeys.1003.55541.figure4" httpUri="https://binary.pensoft.net/fig/486555" pageId="0" pageNumber="31">4C, D</figureCitation>
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|
). The examination of their hind wings revealed one macropterous and five brachypterous specimens.
|
|
</paragraph>
|
|
<paragraph id="179D3535A1DFFFD67FDF01C470D3AA4D" pageId="0" pageNumber="31">
|
|
Lineages detected by the analysis of COI nucleotide sequences were not confirmed by PCA analyses of morphometric characters (Fig.
|
|
<figureCitation id="8C9A01C5D3CE1869105D02D0B814BD22" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Results of the PCA analysis showing ordination of Ancyronyx lianlabangorum sp. nov. and A. pulcherrimus Kodada, Jaech & Ciampor specimens along the first two principal components A males B females. Clusters of A. pulcherrimus specimens on left side of plot (yellow lines), clusters of A. lianlabangorum sp. nov. on right side (green lines). Yellow circles: A. pulcherrimus specimens from Gunung Mulu NP, first lineage; blue circles: A. pulcherrimus from Bayur River, second lineage; brown circles: A. pulcherrimus from Bayur River, third lineage; black circles: A. pulcherrimus from Bayur River, specimens not barcoded, not assignable to the first or second lineage according to morphology; red circle: A. pulcherrimus from Brunei; green squares: A. lianlabangorum sp. nov. Genetic voucher IDs are included according to Table 1." figureDoi="10.3897/zookeys.1003.55541.figure2" httpUri="https://binary.pensoft.net/fig/486553" pageId="0" pageNumber="31">2</figureCitation>
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|
, Table
|
|
<tableCitation id="C00136A88B49F969999DDD4085AE720D" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Metric characters of Ancyronyx lianlabangorum sp. nov. and A. pulcherrimus, parameters are provided as range and mean + / - standard deviation." httpUri="http://table.plazi.org/id/3ADE785948A5EE01D88FB0E6A7D7E580" pageId="0" pageNumber="31" tableUuid="3ADE785948A5EE01D88FB0E6A7D7E580">4</tableCitation>
|
|
) or corroborated by the genital morphology.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="55C3EEAF28FFECC82BB5C53B26FFB527" pageId="0" pageNumber="31" type="habitat">
|
|
<paragraph id="D5897075C3735C638127EDBAB1952B54" pageId="0" pageNumber="31">Habitat and syntopic taxa.</paragraph>
|
|
<paragraph id="AF29B36B36FED868A4DE7225BBAC1E85" pageId="0" pageNumber="31">
|
|
The stream inhabited by a large population of
|
|
<taxonomicName id="41124F7B25261946EA1E442B9C41FA45" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. pulcherrimus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="pulcherrimus">
|
|
<emphasis id="4B8C856DBAC819E5209F92E5801FD6E4" italics="true" pageId="0" pageNumber="31">A. pulcherrimus</emphasis>
|
|
</taxonomicName>
|
|
was a small, slowly flowing, very shallow, and entirely shaded meandering creek in Gunung Mulu NP (55 m a.s.l.). It included large amounts of accumulated leaves, and the bottom substrate contained sand and gravel, as well as woody debris (Fig.
|
|
<figureCitation id="DB277628FC5765EF100137612CDE092C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitats of Ancyronyx lianlabangorum sp. nov. (A-C) and A. pulcherrimus (D) A type locality, Pa' Ramudu River B Pa' Masia River C Pa' Kasi River D shaded shallow creek in primary forest, Gunung Mulu NP." figureDoi="10.3897/zookeys.1003.55541.figure5" httpUri="https://binary.pensoft.net/fig/486556" pageId="0" pageNumber="31">5D</figureCitation>
|
|
). The specimens co-occurred with
|
|
<taxonomicName id="3A7F03ED490D0DEFB9A19B604F1AB7BC" authorityName="Jaech" authorityYear="1994" class="Insecta" family="Elmidae" genus="Ancyronyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancyronyx sarawacensis" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="sarawacensis">
|
|
<emphasis id="E979EF74A4833A02244934FEF6C53D0A" italics="true" pageId="0" pageNumber="31">Ancyronyx sarawacensis</emphasis>
|
|
</taxonomicName>
|
|
and two new species of
|
|
<taxonomicName id="ABBD718E4F4ABBF34A180FB9A473EF56" authorityName="Curtis" authorityYear="1830" class="Insecta" family="Elmidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="DA70C1DD1EC771B90E6C0E351D47A7CE" authorityName="Kodada & Ciampor" authorityYear="2003" class="Insecta" family="Elmidae" genus="Okalia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Okalia" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="A44D0E86E859AB29D19B3628C743B606" italics="true" pageId="0" pageNumber="31">Okalia</emphasis>
|
|
</taxonomicName>
|
|
Kodada &
|
|
<normalizedToken id="F1FABC7C643782C401A15192F67D2644" originalValue="Čiampor">Ciampor</normalizedToken>
|
|
and
|
|
<taxonomicName id="6552C1885D6BC812358AAA1AC70894D9" authorityName="Sharp" authorityYear="1888" class="Insecta" family="Elmidae" genus="Leptelmis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Leptelmis" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C41E6E647F7608F8CAB9AEABDE6ACA58" italics="true" pageId="0" pageNumber="31">Leptelmis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8DD59BE19F0F754754EDC8D217CAE913" pageId="0" pageNumber="31">
|
|
The second population was found in a moderately wide (5-8 m) lowland river near Kampung Bayur (Kuching Division); the sampled stretch was shallow and meandering, with sandy/gravelly substrate containing submerged logs, woody debris, and leaves. All specimens were collected exclusively in reaches with stronger current from submerged logs together with
|
|
<taxonomicName id="7FDB5FA0FB876DAA51E2B62F529D672D" authorityName="Jaech" authorityYear="1994" class="Insecta" family="Elmidae" genus="Ancyronyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancyronyx procerus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="procerus">
|
|
<emphasis id="A4FEE68E3C4F635DCDB9AD0585531CE1" italics="true" pageId="0" pageNumber="31">Ancyronyx procerus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C51A9FF51862E26DCECC130DB9C8093" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. acaroides" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="acaroides">
|
|
<emphasis id="1C38E553A95D7B31EA5509F145D582E4" italics="true" pageId="0" pageNumber="31">A. acaroides</emphasis>
|
|
</taxonomicName>
|
|
, and several other
|
|
<taxonomicName id="C8259B688AE6444D065A3FCC736BB586" authorityName="Curtis" authorityYear="1830" class="Insecta" family="Elmidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="A3F644C118448C343C9A30EEE8C12FC7" authorityName="Sharp" authorityYear="1888" class="Insecta" family="Elmidae" genus="Leptelmis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Leptelmis" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E59017C6C31B2E92E946C6C3251BDEBD" italics="true" pageId="0" pageNumber="31">Leptelmis</emphasis>
|
|
</taxonomicName>
|
|
sp. and
|
|
<taxonomicName id="DCB8815B958C40D4DD070EDF231B26B3" authorityName="Deleve" authorityYear="1968" class="Insecta" family="Elmidae" genus="Graphelmis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Graphelmis" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2F8EBA3229AC7D090F95F1C6512C3CD5" italics="true" pageId="0" pageNumber="31">Graphelmis</emphasis>
|
|
</taxonomicName>
|
|
spp. of the
|
|
<taxonomicName id="707F587975056DBAF2F891AFA8AB5F22" lsidName="G. picta" pageId="0" pageNumber="31" rank="species" species="picta">
|
|
<emphasis id="A10137545E5278218011A579767792C3" italics="true" pageId="0" pageNumber="31">G. picta</emphasis>
|
|
</taxonomicName>
|
|
(Reitter) and
|
|
<taxonomicName id="0E578FC2FBE80FD5597BCA4414F34495" lsidName="G. marshalli" pageId="0" pageNumber="31" rank="species" species="marshalli">
|
|
<emphasis id="6CD62F149943B99D63A91313A8A4AAC0" italics="true" pageId="0" pageNumber="31">G. marshalli</emphasis>
|
|
</taxonomicName>
|
|
Hinton species groups).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C351A14B298EFB5948DA95C2F8532ED2" pageId="0" pageNumber="31" type="distribution">
|
|
<paragraph id="DEC12AC8CC79A94A9F469E4A53E590A9" pageId="0" pageNumber="31">Distribution.</paragraph>
|
|
<paragraph id="7462812473176123F15FC50A15051E18" pageId="0" pageNumber="31">
|
|
<taxonomicName id="ED6B4D2A8048049DB2E932AE53CA2152" authorityName="Kodada, Jaech & Ciampor" authorityYear="2014" class="Insecta" family="Elmidae" genus="Ancyronyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancyronyx pulcherrimus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="pulcherrimus">
|
|
<emphasis id="3BCA121FC93AF2AC9FC0F0DA022C8FF5" italics="true" pageId="0" pageNumber="31">Ancyronyx pulcherrimus</emphasis>
|
|
</taxonomicName>
|
|
was so far known only from Brunei (
|
|
<bibRefCitation id="60FB81271A71BE5B5ECBE636501C8319" DOI="https://doi.org/10.1163/9789004291775" author="Jaech, MA" journalOrPublisher="B)" pageId="0" pageNumber="31" refId="B17" refString="Jaech, MA, Kodada, J, Brojer, M, Shepard, WD, Ciampor, Jr F, 2016. Coleoptera: Elmidae and Protelmidae. World Catalogue of Insects (Vol. 14). Brill, XXI + 318 pp. https://doi.org/10.1163/9789004291775" title="Coleoptera: Elmidae and Protelmidae. World Catalogue of Insects (Vol. 14). Brill, XXI + 318 pp." url="https://doi.org/10.1163/9789004291775" year="2016">
|
|
<normalizedToken id="7043DC230EFBC49E36148C663091B762" originalValue="Jäch">Jaech</normalizedToken>
|
|
et al. 2016
|
|
</bibRefCitation>
|
|
). The species is here recorded for the first time from Malaysia, where it was collected in Gunung Mulu NP (northern Sarawak) and in the Kuching Division (southwestern Sarawak).
|
|
</paragraph>
|
|
<paragraph id="72BE4FAD98577AA74CDE22B73D7D7A34" pageId="0" pageNumber="31">
|
|
Our recent collecting activities found
|
|
<taxonomicName id="7F1A309C8DFDD2F6836AD749754446B0" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. pulcherrimus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="pulcherrimus">
|
|
<emphasis id="C566601FB9A646673B58B96196BFDD30" italics="true" pageId="0" pageNumber="31">A. pulcherrimus</emphasis>
|
|
</taxonomicName>
|
|
to be less abundant and less widely distributed than
|
|
<taxonomicName id="3ABC0F6E88BB69DFD2576E4A3F9E5BBA" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. sarawacensis" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="sarawacensis">
|
|
<emphasis id="C3D2F04DE25CD6BC52C5C8B8959AC474" italics="true" pageId="0" pageNumber="31">A. sarawacensis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="9D66DDEE42AF46EEA5AE1DA06C456AB1" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. procerus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="procerus">
|
|
<emphasis id="C5DB6DC418387BBAB2AF2B23F88DAF2F" italics="true" pageId="0" pageNumber="31">A. procerus</emphasis>
|
|
</taxonomicName>
|
|
, or
|
|
<taxonomicName id="141C71570F5B519B5AFAD01B030D0542" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. acaroides" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="acaroides">
|
|
<emphasis id="DFF2DD205AFD87FB74BE1B5ECF384E52" italics="true" pageId="0" pageNumber="31">A. acaroides</emphasis>
|
|
</taxonomicName>
|
|
, probably also due to the low dispersal ability of flightless populations.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</subSection>
|
|
</document> |