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<document ID-DOI="http://doi.org/10.5281/zenodo.5662798" ID-GBIF-Dataset="78c8f10e-2034-47dd-b518-7ae7a74df5ec" ID-GBIF-Taxon="137281650" ID-Zenodo-Dep="5662798" LSID="urn:lsid:zoobank.org:act:4A7AFFEE-D9E7-4F74-A586-129B5D401F60" checkinTime="1510613567129" checkinUser="plazi" docAuthor="Kevin W. Conway, Glenn I. Moore &amp; Adam P. Summers" docDate="2017" docId="AC2B87D71D61CE72FFC70F673159EA8E" docLanguage="en" docName="copeia.105.1.128.pdf" docOrigin="Copeia 105 (1)" docStyle="DocumentStyle{}" docTitle="Nettorhamphos radula Kevin W. Conway &amp; Glenn I. Moore &amp; Adam P. Summers 2017, new species" docType="treatment" docUuid="4A7AFFEE-D9E7-4F74-A586-129B5D401F60" docUuidSource="ZooBank" docVersion="5" lastPageId="3" lastPageNumber="131" masterDocId="5012FFAF1D63CE71FFA00A593407EC45" masterDocTitle="A New Genus and Species of Clingfish (Teleostei: Gobiesocidae) from Western Australia" masterLastPageNumber="140" masterPageNumber="128" pageId="2" pageNumber="130" updateTime="1636502482336" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A New Genus and Species of Clingfish (Teleostei: Gobiesocidae) from Western Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kevin W. Conway</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Glenn I. Moore</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Adam P. Summers</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Copeia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-04-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>105</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>128</mods:start>
<mods:end>140</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1643/CI-16-560</mods:identifier>
<mods:identifier type="GBIF-Dataset">78c8f10e-2034-47dd-b518-7ae7a74df5ec</mods:identifier>
<mods:identifier type="Zenodo-Dep">1048359</mods:identifier>
<mods:identifier type="ZooBank">75690F95-BE77-4AEF-A218-9D46B2228D9F</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5662798" ID-GBIF-Taxon="137281650" ID-Zenodo-Dep="5662798" LSID="urn:lsid:zoobank.org:act:4A7AFFEE-D9E7-4F74-A586-129B5D401F60" httpUri="http://treatment.plazi.org/id/AC2B87D71D61CE72FFC70F673159EA8E" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130">
<subSubSection box="[103,521,1342,1365]" pageId="2" pageNumber="130" type="nomenclature">
<paragraph blockId="2.[103,667,1342,1454]" box="[103,521,1342,1365]" pageId="2" pageNumber="130">
<heading bold="true" box="[103,521,1342,1365]" fontSize="9" level="2" pageId="2" pageNumber="130" reason="6">
<emphasis bold="true" box="[103,521,1342,1365]" pageId="2" pageNumber="130">
<emphasis bold="true" box="[103,370,1342,1365]" italics="true" pageId="2" pageNumber="130">
<taxonomicName authority="Kevin W. Conway &amp; Glenn I. Moore &amp; Adam P. Summers, 2017" authorityName="Kevin W. Conway &amp; Glenn I. Moore &amp; Adam P. Summers" authorityYear="2017" box="[103,364,1342,1365]" class="Actinopterygii" family="Gobiesocidae" genus="Nettorhamphos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiesociformes" pageId="2" pageNumber="130" phylum="Chordata" rank="species" species="radula" status="new species">Nettorhamphos radula</taxonomicName>
,
</emphasis>
<taxonomicNameLabel box="[377,521,1342,1365]" pageId="2" pageNumber="130" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="130" type="description">
<paragraph blockId="2.[103,667,1342,1454]" pageId="2" pageNumber="130">urn:lsid:zoobank.org:act:4A7AFFEE-D9E7-4F74-A586-129B5D401F60</paragraph>
</subSubSection>
<subSubSection box="[104,499,1431,1454]" pageId="2" pageNumber="130" type="vernacular_names">
<paragraph blockId="2.[103,667,1342,1454]" box="[104,499,1431,1454]" pageId="2" pageNumber="130">Common name: Duckbilled Clingfish</paragraph>
</subSubSection>
<subSubSection box="[104,240,1461,1484]" pageId="2" pageNumber="130" type="description">
<paragraph blockId="2.[104,240,1461,1484]" box="[104,240,1461,1484]" pageId="2" pageNumber="130">
<figureCitation box="[104,240,1461,1484]" captionStart-0="Fig. 1" captionStart-1="Fig. 2" captionStart-2="Fig. 3" captionStart-3="Fig. 4" captionStart-4="Fig. 5" captionStart-5="Fig. 6" captionStart-6="Fig. 7" captionStart-7="Fig. 8" captionStart-8="Fig. 9" captionStart-9="Fig. 10" captionStartId-0="1.[142,170,1369,1389]" captionStartId-1="2.[103,131,1068,1088]" captionStartId-2="3.[142,170,1104,1124]" captionStartId-3="5.[142,170,1817,1837]" captionStartId-4="5.[142,170,1464,1484]" captionStartId-5="6.[103,131,721,741]" captionStartId-6="6.[102,130,1923,1943]" captionStartId-7="7.[142,170,953,973]" captionStartId-8="8.[103,131,1341,1361]" captionStartId-9="9.[142,170,1016,1036]" captionTargetBox-0="[142,1530,130,1339]" captionTargetBox-1="[103,1490,130,1038]" captionTargetBox-2="[143,812,131,1075]" captionTargetBox-3="[103,1491,130,1926]" captionTargetBox-4="[142,1530,130,1434]" captionTargetBox-5="[103,1491,130,692]" captionTargetBox-6="[102,1490,1023,1894]" captionTargetBox-7="[0,1529,53,923]" captionTargetBox-8="[103,1491,130,1312]" captionTargetBox-9="[0,1530,53,986]" captionTargetId-0="figure@1.[142,1530,130,1339]" captionTargetId-1="figure@2.[103,1490,130,1038]" captionTargetId-2="figure@3.[142,812,130,1075]" captionTargetId-3="figure@4.[103,1491,130,1926]" captionTargetId-4="figure@5.[142,1530,130,1434]" captionTargetId-5="figure@6.[103,1491,130,692]" captionTargetId-6="figure@6.[102,1490,1023,1894]" captionTargetId-7="figure@7.[142,1529,130,923]" captionTargetId-8="figure@8.[103,1491,130,1312]" captionTargetId-9="figure@9.[142,1530,130,986]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionTargetPageId-6="6" captionTargetPageId-7="7" captionTargetPageId-8="8" captionTargetPageId-9="9" captionText-0="Fig. 1. Nettorhamphos radula, WAM P. 26052 - 004, holotype, 40.2 mm SL, Indian Ocean, Western Australia." captionText-1="Fig. 2. Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL, Indian Ocean, Western Australia." captionText-2="Fig. 3. Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. Line drawing of head in dorsal, lateral, and ventral view showing position of cephalic lateral line canal pores. Abbreviations: AN, anterior nostril; FL, fleshy lobe; LC, lacrimal canal pores; MC, mandibular canal pores; NC, nasal canal pores; PN, posterior nostril; PO, postorbital canal pores; PR, preopercular canal pores." captionText-3="Fig. 4. Computed tomography scanned anterior skeleton, including cranium, paired-fin girdles, and abdominal region of vertebral column, of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Lateral view. (C) Ventral view. Single scale bar shared by A C. Abbreviations: ACh, anterior ceratohyal; Ana, anguloarticular; Apa, autopalatine; Asph, autosphenotic; Bh, basihyal; Boc, basioccipital; Bp, basipterygium; Br, branchiostegal rays; Cb 5, ceratobranchial five; Cl, cleithrum; De, dentary; DPcl, dorsal postcleithrum; E, ethmoid; Ec, epicentral; Epoc, epioccipital; Fr, frontal; Hy, hyomandibular; Ihc, interhyal; Iop, interopercle; La, lacrimal; LE, lateral ethmoid; Mx, maxilla; Na, nasal; NS, neural spine; Op, opercle; Pa, parietal; PCh, posterior ceratohyal; Pmx, premaxilla; Pop, preopercle; Pro, prootic; Psph, parasphenoid; Pt, posttemporal; Pte, pterotic; Q, quadrate; R, fin ray; Ra, retroarticular; Ri, rib; Scl, supracleithrum; Soc, supraoccipital; Sop, subopercle; Sy, symplectic; Ur, urohyal; V, vertebral centrum; VPcl, ventral postcleithrum." captionText-4="Fig. 5. Computed tomography scanned jaws, hyopalatine arch, and opercular series of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Lateral view. (B) Medial view. (C) Anterolateral view. (D) Ventral view of isolated premaxillae. (E) Dorsal (upper) and ventral (lower) views of isolated dentaries. Scale bar shared between A and B. Abbreviations: Ana, anguloarticular; Apa, autopalatine; De, dentary; Ecpt, ectopterygoid; Hy, hyomandibular; Iop, interopercle; La, lacrimal; Mx, maxilla; Op, opercle; Pmx, premaxilla; Pop, preopercle; Q, quadrate; Ra, retroarticular; Sop, subopercle; Sy, symplectic." captionText-5="Fig. 6. Computed tomography scanned hyoid bar (A, B) and pharyngeal jaws (C E) of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Lateroventral view. (B) Medial view. (C) Dorsal view. (D) Ventral view. (E) Anterior view. Single scale bar shared between A and B and C E. Abbreviations: ACh, anterior ceratohyal; Bh, basihyal; Br, branchiostegal rays; Cb 5, ceratobranchial 5; DHh, dorsal hypohal; Ih, interhyal; Pb 3 TP, pharyngobranchial 3 toothplate; PCh, posterior ceratohyal; Uh, urohyal." captionText-6="Fig. 7. Computed tomography scanned skeleton of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Lateral view. (C) Ventral view. (D) Close up of box in B showing posterior region of vertebral column, dorsal, anal, and caudal fin skeletons. Single scale bar shared between A C. Scale bar not provided for D. Abbreviations: Ep, epural; HS 1, hemal spine on vertebral centrum 17; LHP, lower hypural plate; NS 1, neural spine of the first vertebral centrum; PU 2, second preural centrum; R, fin ray; UC, ural centrum; UHP, upper hypural plate; V 17, vertebral centrum 17." captionText-7="Fig. 8. Adhesive disc of Nettorhamphos radula, WAM P. 26052 - 004, holotype, 40.2 mm SL. (A) Adhesive disc stained with cyanine blue, in ventral view. (B) Schematic illustration of adhesive disc with intact papillae in disc regions A D shown in gray. Anterior margin of disc covered by gill membrane along midline. Scars of dislodged papillae visible on the surface of the disc in A not indicated in B." captionText-8="Fig. 9. Computed tomography scanned paired-fin girdles of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Anterior view. Single scale bar shared between A D. Abbreviations: Bp, basipterygium; Cl, cleithrum; Cor, coracoid; DPcl, dorsal postcleithrum; PR, pectoral radial; Pt, posttemporal; R, fin ray; Sc, scapula; Scl, supracleithrum; SR, spinous pelvic-fin ray; VPcl, ventral postcleithrum." captionText-9="Fig. 10. Skin surface of Nettorhamphos radula, WAM P. 26052 - 004, holotype, 40.2 mm SL. (A) Overview of dorsal surface of head, right side, anterior to top of page. (B) Close up of box in A. (C) Close up of box in B. (D) Skin surface after cyanine blue staining." httpUri-0="https://zenodo.org/record/1048361/files/figure.png" httpUri-1="https://zenodo.org/record/1048363/files/figure.png" httpUri-2="https://zenodo.org/record/1048365/files/figure.png" httpUri-3="https://zenodo.org/record/1048369/files/figure.png" httpUri-4="https://zenodo.org/record/1048367/files/figure.png" httpUri-5="https://zenodo.org/record/1048371/files/figure.png" httpUri-6="https://zenodo.org/record/1048373/files/figure.png" httpUri-7="https://zenodo.org/record/1048375/files/figure.png" httpUri-8="https://zenodo.org/record/1048377/files/figure.png" httpUri-9="https://zenodo.org/record/1048379/files/figure.png" pageId="2" pageNumber="130">Figures 110</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="130" type="materials_examined">
<paragraph blockId="2.[103,773,1514,1625]" pageId="2" pageNumber="130">
<materialsCitation ID-GBIF-Occurrence="1699979718" collectingDate="1977-06-30" collectionCode="WAM" collectorName="B. R. Wilson &amp; R. Buick &amp; FRV Flinders" country="Australia" elevation="0" latitude="-32.183334" location="Indian Ocean, west of Five Fathom" longLatPrecision="1212" longitude="115.566666" pageId="2" pageNumber="130" specimenCode="P.26052-004" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis bold="true" box="[103,228,1514,1537]" italics="true" pageId="2" pageNumber="130">
<typeStatus box="[103,200,1514,1537]" pageId="2" pageNumber="130">Holotype</typeStatus>
.—
</emphasis>
<collectionCode box="[227,289,1515,1536]" collectionName="Australia, Western Australia, Perth, Western Australian Museum" pageId="2" pageNumber="130">WAM</collectionCode>
<specimenCode box="[298,436,1514,1537]" pageId="2" pageNumber="130">P.26052-004</specimenCode>
,
<quantity box="[448,550,1515,1536]" metricMagnitude="-2" metricUnit="m" metricValue="4.02" pageId="2" pageNumber="130" unit="mm" value="40.2">40.2 mm</quantity>
SL,
<collectingCountry box="[599,698,1514,1537]" name="Australia" pageId="2" pageNumber="130">Australia</collectingCountry>
,
<collectingRegion country="Australia" name="Western Australia" pageId="2" pageNumber="130">Western Australia</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:AC2B87D71D61CE72FFC70F673159EA8E:215D601A1D61CE73FEA70C513690EA5A" box="[263,663,1544,1567]" country="Australia" latitude="-32.183334" longLatPrecision="1212" longitude="115.566666" name="Indian Ocean, west of Five Fathom" pageId="2" pageNumber="130" stateProvince="Western Australia">Indian Ocean, west of Five Fathom</location>
Bank,
<elevation box="[745,772,1545,1566]" pageId="2" pageNumber="130">33</elevation>
meters in depth,
<geoCoordinate box="[290,374,1570,1596]" direction="south" orientation="latitude" pageId="2" pageNumber="130" precision="925" value="-32.183334">32°11'S</geoCoordinate>
,
<geoCoordinate box="[385,484,1570,1597]" direction="east" orientation="longitude" pageId="2" pageNumber="130" precision="925" value="115.566666">115°34'E</geoCoordinate>
,
<collectorName box="[495,629,1574,1597]" pageId="2" pageNumber="130">B. R. Wilson</collectorName>
and
<collectorName box="[684,773,1574,1597]" pageId="2" pageNumber="130">R. Buick</collectorName>
(
<emphasis box="[111,244,1603,1625]" italics="true" pageId="2" pageNumber="130">
<collectorName box="[111,244,1603,1625]" pageId="2" pageNumber="130">FRV Flinders</collectorName>
</emphasis>
),
<date box="[267,418,1604,1625]" pageId="2" pageNumber="130" value="1977-06-30">
<collectingDate box="[267,418,1604,1625]" pageId="2" pageNumber="130" value="1977-06-30">30 June 1977</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="2.[103,773,1657,1768]" pageId="2" pageNumber="130">
<emphasis bold="true" box="[103,228,1657,1680]" italics="true" pageId="2" pageNumber="130">Paratype.—</emphasis>
WAM P.25805-004, 1, 20.3 mm SL, Australia, Western Australia, Indian Ocean, west of Five Fathom Bank, 37 meters in depth, 32°17'S, 115°31'E, N. Sarti and R. George (
<emphasis box="[111,244,1746,1768]" italics="true" pageId="2" pageNumber="130">FRV Flinders</emphasis>
), 29 June 1977.
</paragraph>
</subSubSection>
<subSubSection box="[103,476,1800,1823]" pageId="2" pageNumber="130" type="diagnosis">
<paragraph blockId="2.[103,476,1800,1823]" box="[103,476,1800,1823]" pageId="2" pageNumber="130">
<emphasis bold="true" box="[103,236,1800,1823]" italics="true" pageId="2" pageNumber="130">Diagnosis.—</emphasis>
See generic diagnosis.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130" type="description">
<paragraph blockId="2.[103,773,1853,2024]" lastBlockId="2.[820,1491,1132,2024]" pageId="2" pageNumber="130">
<emphasis bold="true" box="[103,252,1853,1876]" italics="true" pageId="2" pageNumber="130">Description.—</emphasis>
General body shape as in
<figureCitation box="[575,681,1853,1876]" captionStart="Fig. 1" captionStartId="1.[142,170,1369,1389]" captionTargetBox="[142,1530,130,1339]" captionTargetId="figure@1.[142,1530,130,1339]" captionTargetPageId="1" captionText="Fig. 1. Nettorhamphos radula, WAM P. 26052 - 004, holotype, 40.2 mm SL, Indian Ocean, Western Australia." httpUri="https://zenodo.org/record/1048361/files/figure.png" pageId="2" pageNumber="130">Figures 1</figureCitation>
and
<figureCitation box="[752,768,1854,1875]" captionStart="Fig. 2" captionStartId="2.[103,131,1068,1088]" captionTargetBox="[103,1490,130,1038]" captionTargetId="figure@2.[103,1490,130,1038]" captionTargetPageId="2" captionText="Fig. 2. Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL, Indian Ocean, Western Australia." httpUri="https://zenodo.org/record/1048363/files/figure.png" pageId="2" pageNumber="130">2</figureCitation>
. Morphometric characters listed in
<tableCitation box="[496,579,1883,1906]" captionStart="Table 1" captionStartId="9.[859,908,1295,1315]" captionTargetBox="[859,1529,1360,2007]" captionText="Table 1. Morphometric characters obtained from holotype and paratype of Nettorhamphos radula." httpUri="http://table.plazi.org/id/70FD66491D6ACE78FCFB0F5630ABE978" pageId="2" pageNumber="130" tableUuid="70FD66491D6ACE78FCFB0F5630ABE978">Table 1</tableCitation>
. Head wide, flat, strongly dorsoventrally compressed. Body moderately dorsoventrally compressed anteriorly, becoming increasingly laterally compressed posteriorly. Widest point of head wider than widest point of body (immediately behind head). Body width and depth tapering gradually posteriorly. Eye large, positioned on dorso-lateral surface of head; orbit visible in ventral view. Center of eye closer to tip of snout than to posterior margin of operculum. Snout long, broad, anterior margin rounded. Anterior nostril tubular, with long, fleshy cirrus extending from posterior margin. Posterior nostril surrounded by a low fleshy rim; situated posterior to imaginary vertical line through anterior margin of orbit. Gill membranes united and free from isthmus (gill membranes damaged in paratype). Subopercular spine well developed. Deep pocket present in branchiostegal membrane between posteriormost branchiostegal ray and medial face of subopercular spine.
</paragraph>
<paragraph blockId="2.[820,1491,1132,2024]" lastBlockId="3.[859,1530,131,1073]" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130">
Mouth terminal, large, posterior edge of upper jaw reaching imaginary vertical line through center of orbit when mouth closed. Upper jaw longer and wider than lower jaw, creating a large gap between margin of upper and lower jaws when closed (
<figureCitation box="[1028,1090,1642,1665]" captionStart="Fig. 3" captionStartId="3.[142,170,1104,1124]" captionTargetBox="[143,812,131,1075]" captionTargetId="figure@3.[142,812,130,1075]" captionTargetPageId="3" captionText="Fig. 3. Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. Line drawing of head in dorsal, lateral, and ventral view showing position of cephalic lateral line canal pores. Abbreviations: AN, anterior nostril; FL, fleshy lobe; LC, lacrimal canal pores; MC, mandibular canal pores; NC, nasal canal pores; PN, posterior nostril; PO, postorbital canal pores; PR, preopercular canal pores." httpUri="https://zenodo.org/record/1048365/files/figure.png" pageId="2" pageNumber="130">Fig. 3</figureCitation>
). Upper lip fleshy, widest anteriorly, separated from snout by shallow groove. Lower lip with pair of shallow fleshy lobes below symphysis. Premaxilla with
<emphasis bold="true" box="[820,838,1732,1754]" pageId="2" pageNumber="130">~</emphasis>
15 rows of
<emphasis bold="true" box="[962,980,1732,1754]" pageId="2" pageNumber="130">~</emphasis>
4050 tiny, dagger-like conical teeth (
<figureCitation box="[1395,1478,1732,1755]" captionStart="Fig. 5" captionStartId="5.[142,170,1464,1484]" captionTargetBox="[142,1530,130,1434]" captionTargetId="figure@5.[142,1530,130,1434]" captionTargetPageId="5" captionText="Fig. 5. Computed tomography scanned jaws, hyopalatine arch, and opercular series of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Lateral view. (B) Medial view. (C) Anterolateral view. (D) Ventral view of isolated premaxillae. (E) Dorsal (upper) and ventral (lower) views of isolated dentaries. Scale bar shared between A and B. Abbreviations: Ana, anguloarticular; Apa, autopalatine; De, dentary; Ecpt, ectopterygoid; Hy, hyomandibular; Iop, interopercle; La, lacrimal; Mx, maxilla; Op, opercle; Pmx, premaxilla; Pop, preopercle; Q, quadrate; Ra, retroarticular; Sop, subopercle; Sy, symplectic." httpUri="https://zenodo.org/record/1048367/files/figure.png" pageId="2" pageNumber="130">Fig. 5D</figureCitation>
). Lower jaw of holotype with
<emphasis bold="true" box="[1126,1144,1762,1784]" pageId="2" pageNumber="130">~</emphasis>
10 rows of
<emphasis bold="true" box="[1263,1281,1762,1784]" pageId="2" pageNumber="130">~</emphasis>
3040 tiny, daggerlike conical teeth; lower jaw of paratype with single row of 8 large conical teeth anteriorly, followed by
<emphasis bold="true" box="[1284,1302,1822,1844]" pageId="2" pageNumber="130">~</emphasis>
10 rows of 3040 tiny, dagger-like conical teeth (
<figureCitation box="[1154,1230,1851,1874]" captionStart="Fig. 5" captionStartId="5.[142,170,1464,1484]" captionTargetBox="[142,1530,130,1434]" captionTargetId="figure@5.[142,1530,130,1434]" captionTargetPageId="5" captionText="Fig. 5. Computed tomography scanned jaws, hyopalatine arch, and opercular series of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Lateral view. (B) Medial view. (C) Anterolateral view. (D) Ventral view of isolated premaxillae. (E) Dorsal (upper) and ventral (lower) views of isolated dentaries. Scale bar shared between A and B. Abbreviations: Ana, anguloarticular; Apa, autopalatine; De, dentary; Ecpt, ectopterygoid; Hy, hyomandibular; Iop, interopercle; La, lacrimal; Mx, maxilla; Op, opercle; Pmx, premaxilla; Pop, preopercle; Q, quadrate; Ra, retroarticular; Sop, subopercle; Sy, symplectic." httpUri="https://zenodo.org/record/1048367/files/figure.png" pageId="2" pageNumber="130">Fig. 5E</figureCitation>
). Tiny, dagger-like teeth of lower jaw supported by thin, poorly ossified bony shelf on dorsal surface of dentary. Pharyngeal jaws comprising patch of 1215 small conical teeth on pharyngobranchial toothplate 3 and two rows of 510 small conical teeth along ceratobranchial 5 (
<figureCitation box="[1054,1170,2001,2024]" captionStart="Fig. 6" captionStartId="6.[103,131,721,741]" captionTargetBox="[103,1491,130,692]" captionTargetId="figure@6.[103,1491,130,692]" captionTargetPageId="6" captionText="Fig. 6. Computed tomography scanned hyoid bar (A, B) and pharyngeal jaws (C E) of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Lateroventral view. (B) Medial view. (C) Dorsal view. (D) Ventral view. (E) Anterior view. Single scale bar shared between A and B and C E. Abbreviations: ACh, anterior ceratohyal; Bh, basihyal; Br, branchiostegal rays; Cb 5, ceratobranchial 5; DHh, dorsal hypohal; Ih, interhyal; Pb 3 TP, pharyngobranchial 3 toothplate; PCh, posterior ceratohyal; Uh, urohyal." httpUri="https://zenodo.org/record/1048371/files/figure.png" pageId="2" pageNumber="130">Fig. 6CE</figureCitation>
). Poorly ossified basihyal with vertebrae 314. Epicentrals 12* or 13, associated with vertebrae 314* or 315.
</paragraph>
<paragraph blockId="3.[859,1530,131,1073]" pageId="3" pageNumber="131">
Adhesive disc large, double (
<figureCitation box="[1196,1257,190,213]" captionStart="Fig. 8" captionStartId="7.[142,170,953,973]" captionTargetBox="[0,1529,53,923]" captionTargetId="figure@7.[142,1529,130,923]" captionTargetPageId="7" captionText="Fig. 8. Adhesive disc of Nettorhamphos radula, WAM P. 26052 - 004, holotype, 40.2 mm SL. (A) Adhesive disc stained with cyanine blue, in ventral view. (B) Schematic illustration of adhesive disc with intact papillae in disc regions A D shown in gray. Anterior margin of disc covered by gill membrane along midline. Scars of dislodged papillae visible on the surface of the disc in A not indicated in B." httpUri="https://zenodo.org/record/1048375/files/figure.png" pageId="3" pageNumber="131">Fig. 8</figureCitation>
); disc papillae and outer margin of disc degraded in both specimens. Disc papillae remaining in holotype as follows: disc region A with 12 incomplete transverse rows of papillae along anteromedian edge and smaller, sporadic clusters of papillae laterally over ventral surface of pelvic-fin rays, including a large patch comprised of 3 incomplete transverse rows beneath first soft pelvic-fin ray on left side; disc region B with 12 incomplete transverse rows of papillae along posterior edge and sporadic clusters of papillae anterolaterally; disc region C and D with incomplete median clusters of papillae. Basipterygium widest anteriorly, tapering posteriorly into thin, elongate processus ischiaticus (
<figureCitation box="[1006,1133,546,569]" captionStart="Fig. 9" captionStartId="8.[103,131,1341,1361]" captionTargetBox="[103,1491,130,1312]" captionTargetId="figure@8.[103,1491,130,1312]" captionTargetPageId="8" captionText="Fig. 9. Computed tomography scanned paired-fin girdles of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Anterior view. Single scale bar shared between A D. Abbreviations: Bp, basipterygium; Cl, cleithrum; Cor, coracoid; DPcl, dorsal postcleithrum; PR, pectoral radial; Pt, posttemporal; R, fin ray; Sc, scapula; Scl, supracleithrum; SR, spinous pelvic-fin ray; VPcl, ventral postcleithrum." httpUri="https://zenodo.org/record/1048377/files/figure.png" pageId="3" pageNumber="131">Fig. 9A, B</figureCitation>
); a large foramen at center of basipterygium. Anterior margin of pelvic girdle concave, tightly sutured to anteromedial face of pectoral girdle (
<figureCitation captionStart="Fig. 9" captionStartId="8.[103,131,1341,1361]" captionTargetBox="[103,1491,130,1312]" captionTargetId="figure@8.[103,1491,130,1312]" captionTargetPageId="8" captionText="Fig. 9. Computed tomography scanned paired-fin girdles of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Anterior view. Single scale bar shared between A D. Abbreviations: Bp, basipterygium; Cl, cleithrum; Cor, coracoid; DPcl, dorsal postcleithrum; PR, pectoral radial; Pt, posttemporal; R, fin ray; Sc, scapula; Scl, supracleithrum; SR, spinous pelvic-fin ray; VPcl, ventral postcleithrum." httpUri="https://zenodo.org/record/1048377/files/figure.png" pageId="3" pageNumber="131">Fig. 9A, B</figureCitation>
). Dorsal postcleithrum small, roughly semi-circular in shape; without contact to coracoid or ventral postcleithrum. Ventral postcleithrum larger than dorsal postcleithrum; irregular in shape, wider laterally then medially with a concave anterior margin and raised medial edge developed into a prominent anterodorsal process. Anterodorsal processes of ventral postcleithra in contact across midline and extended anterodorsally towards processus ischiatici of basipterygia (
<figureCitation box="[1007,1091,872,895]" captionStart="Fig. 9" captionStartId="8.[103,131,1341,1361]" captionTargetBox="[103,1491,130,1312]" captionTargetId="figure@8.[103,1491,130,1312]" captionTargetPageId="8" captionText="Fig. 9. Computed tomography scanned paired-fin girdles of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Anterior view. Single scale bar shared between A D. Abbreviations: Bp, basipterygium; Cl, cleithrum; Cor, coracoid; DPcl, dorsal postcleithrum; PR, pectoral radial; Pt, posttemporal; R, fin ray; Sc, scapula; Scl, supracleithrum; SR, spinous pelvic-fin ray; VPcl, ventral postcleithrum." httpUri="https://zenodo.org/record/1048377/files/figure.png" pageId="3" pageNumber="131">Fig. 9D</figureCitation>
). Elongate fimbriae not observed along posterior margin of ventral postcleithra. Four poorly ossified pectoral radials; tightly sutured via small, fingerlike projections of bone (
<figureCitation box="[1017,1097,961,984]" captionStart="Fig. 9" captionStartId="8.[103,131,1341,1361]" captionTargetBox="[103,1491,130,1312]" captionTargetId="figure@8.[103,1491,130,1312]" captionTargetPageId="8" captionText="Fig. 9. Computed tomography scanned paired-fin girdles of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Ventral view. (C) Lateral view. (D) Anterior view. Single scale bar shared between A D. Abbreviations: Bp, basipterygium; Cl, cleithrum; Cor, coracoid; DPcl, dorsal postcleithrum; PR, pectoral radial; Pt, posttemporal; R, fin ray; Sc, scapula; Scl, supracleithrum; SR, spinous pelvic-fin ray; VPcl, ventral postcleithrum." httpUri="https://zenodo.org/record/1048377/files/figure.png" pageId="3" pageNumber="131">Fig. 9C</figureCitation>
). Skin associated with last pelvic-fin ray attaching to base of pectoral fin opposite 6th lowermost pectoral-fin ray. No fleshy pad on lateral surface of pectoral base.
</paragraph>
<caption httpUri="https://zenodo.org/record/1048365/files/figure.png" pageId="3" pageNumber="131" targetBox="[143,812,131,1075]" targetPageId="3">
<paragraph blockId="3.[142,812,1104,1257]" pageId="3" pageNumber="131">
<emphasis bold="true" box="[142,195,1104,1124]" pageId="3" pageNumber="131">Fig. 3.</emphasis>
<emphasis box="[216,437,1104,1124]" italics="true" pageId="3" pageNumber="131">
<taxonomicName authorityName="Kevin W. Conway &amp; Glenn I. Moore &amp; Adam P. Summers" authorityYear="2017" box="[216,433,1104,1124]" class="Actinopterygii" family="Gobiesocidae" genus="Nettorhamphos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="131" phylum="Chordata" rank="species" species="radula">Nettorhamphos radula</taxonomicName>
,
</emphasis>
WAM P.25805-004, paratype, 20.3 mm SL. Line drawing of head in dorsal, lateral, and ventral view showing position of cephalic lateral line canal pores. Abbreviations: AN, anterior nostril; FL, fleshy lobe; LC, lacrimal canal pores; MC, mandibular canal pores; NC, nasal canal pores; PN, posterior nostril; PO, postorbital canal pores; PR, preopercular canal pores.
</paragraph>
</caption>
<paragraph blockId="3.[142,812,1303,2024]" pageId="3" pageNumber="131">
spatula-shaped. Other aspects of gill arches not observed. Branchiostegal rays 6; 2 anteriormost rays articulating medially with hyoid bar along anterior ceratohyal; posterior rays articulating with hyoid bar laterally, including 3 along anterior ceratohyal and 1 along posterior ceratohyal (
<figureCitation captionStart="Fig. 6" captionStartId="6.[103,131,721,741]" captionTargetBox="[103,1491,130,692]" captionTargetId="figure@6.[103,1491,130,692]" captionTargetPageId="6" captionText="Fig. 6. Computed tomography scanned hyoid bar (A, B) and pharyngeal jaws (C E) of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Lateroventral view. (B) Medial view. (C) Dorsal view. (D) Ventral view. (E) Anterior view. Single scale bar shared between A and B and C E. Abbreviations: ACh, anterior ceratohyal; Bh, basihyal; Br, branchiostegal rays; Cb 5, ceratobranchial 5; DHh, dorsal hypohal; Ih, interhyal; Pb 3 TP, pharyngobranchial 3 toothplate; PCh, posterior ceratohyal; Uh, urohyal." httpUri="https://zenodo.org/record/1048371/files/figure.png" pageId="3" pageNumber="131">Fig. 6A, B</figureCitation>
).
</paragraph>
<paragraph blockId="3.[142,812,1303,2024]" pageId="3" pageNumber="131">
Cephalic lateral-line system with 2 pores in nasal canal; 2 pores in postorbital canal; 2 pores in lacrimal canal; 3 pores in preopercular canal; and 2 pores in mandibular canal. Autosphenotic with large spine-like process laterally (
<figureCitation box="[735,799,1589,1612]" captionStart="Fig. 4" captionStartId="5.[142,170,1817,1837]" captionTargetId="figure@4.[103,1491,130,1926]" captionTargetPageId="4" captionText="Fig. 4. Computed tomography scanned anterior skeleton, including cranium, paired-fin girdles, and abdominal region of vertebral column, of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Lateral view. (C) Ventral view. Single scale bar shared by A C. Abbreviations: ACh, anterior ceratohyal; Ana, anguloarticular; Apa, autopalatine; Asph, autosphenotic; Bh, basihyal; Boc, basioccipital; Bp, basipterygium; Br, branchiostegal rays; Cb 5, ceratobranchial five; Cl, cleithrum; De, dentary; DPcl, dorsal postcleithrum; E, ethmoid; Ec, epicentral; Epoc, epioccipital; Fr, frontal; Hy, hyomandibular; Ihc, interhyal; Iop, interopercle; La, lacrimal; LE, lateral ethmoid; Mx, maxilla; Na, nasal; NS, neural spine; Op, opercle; Pa, parietal; PCh, posterior ceratohyal; Pmx, premaxilla; Pop, preopercle; Pro, prootic; Psph, parasphenoid; Pt, posttemporal; Pte, pterotic; Q, quadrate; R, fin ray; Ra, retroarticular; Ri, rib; Scl, supracleithrum; Soc, supraoccipital; Sop, subopercle; Sy, symplectic; Ur, urohyal; V, vertebral centrum; VPcl, ventral postcleithrum." httpUri="https://zenodo.org/record/1048369/files/figure.png" pageId="3" pageNumber="131">Fig. 4</figureCitation>
). Lacrimal a tiny tube-like ossification (
<figureCitation box="[561,674,1621,1644]" captionStart="Fig. 5" captionStartId="5.[142,170,1464,1484]" captionTargetBox="[142,1530,130,1434]" captionTargetId="figure@5.[142,1530,130,1434]" captionTargetPageId="5" captionText="Fig. 5. Computed tomography scanned jaws, hyopalatine arch, and opercular series of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Lateral view. (B) Medial view. (C) Anterolateral view. (D) Ventral view of isolated premaxillae. (E) Dorsal (upper) and ventral (lower) views of isolated dentaries. Scale bar shared between A and B. Abbreviations: Ana, anguloarticular; Apa, autopalatine; De, dentary; Ecpt, ectopterygoid; Hy, hyomandibular; Iop, interopercle; La, lacrimal; Mx, maxilla; Op, opercle; Pmx, premaxilla; Pop, preopercle; Q, quadrate; Ra, retroarticular; Sop, subopercle; Sy, symplectic." httpUri="https://zenodo.org/record/1048367/files/figure.png" pageId="3" pageNumber="131">Fig. 5A, C</figureCitation>
).
</paragraph>
<paragraph blockId="3.[142,812,1303,2024]" lastBlockId="3.[859,1529,1104,1275]" pageId="3" pageNumber="131">
Dorsal-fin rays 5 or 6*. Anal-fin rays 7 or 8*. Principal caudal-fin rays 5þ4, dorsal procurrent rays 3*, ventral procurrent rays 2*. Pectoral-fin rays 21* or 22. Pelvic-fin rays I.4. All fin rays unbranched and segmented. Pterygiophores not observed. Caudal fin truncate. Caudal-fin skeleton comprised of upper and lower hypural plates (
<figureCitation box="[709,800,1812,1835]" captionStart="Fig. 7" captionStartId="6.[102,130,1923,1943]" captionTargetBox="[102,1490,1023,1894]" captionTargetId="figure@6.[102,1490,1023,1894]" captionTargetPageId="6" captionText="Fig. 7. Computed tomography scanned skeleton of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Lateral view. (C) Ventral view. (D) Close up of box in B showing posterior region of vertebral column, dorsal, anal, and caudal fin skeletons. Single scale bar shared between A C. Scale bar not provided for D. Abbreviations: Ep, epural; HS 1, hemal spine on vertebral centrum 17; LHP, lower hypural plate; NS 1, neural spine of the first vertebral centrum; PU 2, second preural centrum; R, fin ray; UC, ural centrum; UHP, upper hypural plate; V 17, vertebral centrum 17." httpUri="https://zenodo.org/record/1048373/files/figure.png" pageId="3" pageNumber="131">Fig. 7D</figureCitation>
). Epural and parhypural not observed. Dorsal-fin origin situated slightly posterior to imaginary vertical line through anal-fin origin. Dorsal and anal fins collapsed and tightly adhered to body in both specimens; aspects of fin membranes not observed. Total number of vertebrae 33* or 34, consisting of 15þ18* or 16þ18 (
<figureCitation box="[509,574,2001,2024]" captionStart="Fig. 7" captionStartId="6.[102,130,1923,1943]" captionTargetBox="[102,1490,1023,1894]" captionTargetId="figure@6.[102,1490,1023,1894]" captionTargetPageId="6" captionText="Fig. 7. Computed tomography scanned skeleton of Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL. (A) Dorsal view. (B) Lateral view. (C) Ventral view. (D) Close up of box in B showing posterior region of vertebral column, dorsal, anal, and caudal fin skeletons. Single scale bar shared between A C. Scale bar not provided for D. Abbreviations: Ep, epural; HS 1, hemal spine on vertebral centrum 17; LHP, lower hypural plate; NS 1, neural spine of the first vertebral centrum; PU 2, second preural centrum; R, fin ray; UC, ural centrum; UHP, upper hypural plate; V 17, vertebral centrum 17." httpUri="https://zenodo.org/record/1048373/files/figure.png" pageId="3" pageNumber="131">Fig. 7</figureCitation>
). Ribs 11, associated
<emphasis bold="true" box="[859,1002,1104,1127]" italics="true" pageId="3" pageNumber="131">Coloration.—</emphasis>
In alcohol, head and body background color pale cream (holotype;
<figureCitation box="[1103,1160,1134,1157]" captionStart="Fig. 1" captionStartId="1.[142,170,1369,1389]" captionTargetBox="[142,1530,130,1339]" captionTargetId="figure@1.[142,1530,130,1339]" captionTargetPageId="1" captionText="Fig. 1. Nettorhamphos radula, WAM P. 26052 - 004, holotype, 40.2 mm SL, Indian Ocean, Western Australia." httpUri="https://zenodo.org/record/1048361/files/figure.png" pageId="3" pageNumber="131">Fig. 1</figureCitation>
) or light brown (paratype;
<figureCitation box="[1456,1516,1134,1157]" captionStart="Fig. 2" captionStartId="2.[103,131,1068,1088]" captionTargetBox="[103,1490,130,1038]" captionTargetId="figure@2.[103,1490,130,1038]" captionTargetPageId="2" captionText="Fig. 2. Nettorhamphos radula, WAM P. 25805 - 004, paratype, 20.3 mm SL, Indian Ocean, Western Australia." httpUri="https://zenodo.org/record/1048363/files/figure.png" pageId="3" pageNumber="131">Fig. 2</figureCitation>
). Dorsal surface of head with small, evenly spaced yellow spots tentatively identified as remnants of xanthophores in holotype only (
<figureCitation box="[1046,1131,1223,1246]" captionStart="Fig. 10" captionStartId="9.[142,170,1016,1036]" captionTargetBox="[0,1530,53,986]" captionTargetId="figure@9.[142,1530,130,986]" captionTargetPageId="9" captionText="Fig. 10. Skin surface of Nettorhamphos radula, WAM P. 26052 - 004, holotype, 40.2 mm SL. (A) Overview of dorsal surface of head, right side, anterior to top of page. (B) Close up of box in A. (C) Close up of box in B. (D) Skin surface after cyanine blue staining." httpUri="https://zenodo.org/record/1048379/files/figure.png" pageId="3" pageNumber="131">Fig. 10</figureCitation>
). Fins hyaline. Coloration in life unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="131" type="distribution">
<paragraph blockId="3.[859,1529,1306,1596]" pageId="3" pageNumber="131">
<emphasis bold="true" box="[859,1162,1306,1329]" italics="true" pageId="3" pageNumber="131">Distribution and habitat.—</emphasis>
Known to date only from two locations to the west of Five Fathom Bank (Indian Ocean), off the coast of Fremantle, Western Australia (
<figureCitation box="[1334,1411,1365,1388]" captionStart="Fig. 11" captionStartId="10.[103,131,1128,1148]" captionTargetBox="[104,773,131,1097]" captionTargetId="figure@10.[103,774,130,1098]" captionTargetPageId="10" captionText="Fig. 11. Distribution of sampling localities for Nettorhamphos radula along the coast of Western Australia. Gray square indicates type locality. Prepared by M. Allen." httpUri="https://zenodo.org/record/1048381/files/figure.png" pageId="3" pageNumber="131">Fig. 11</figureCitation>
). The new species was collected via trawls through algae/sponge reefs between 3337 meters in depth. Other species of fishes collected by the same trawls include the pipefish
<emphasis italics="true" pageId="3" pageNumber="131">
<taxonomicName box="[1394,1528,1455,1476]" class="Actinopterygii" family="Syngnathidae" genus="Stigmatopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syngnathiformes" pageId="3" pageNumber="131" phylum="Chordata" rank="genus">Stigmatopora</taxonomicName>
nigra,
</emphasis>
the labrids
<emphasis box="[1054,1299,1484,1506]" italics="true" pageId="3" pageNumber="131">
<taxonomicName authorityName="Steindachner" authorityYear="1884" box="[1054,1184,1484,1506]" class="Actinopterygii" family="Labridae" genus="Austrolabrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="131" phylum="Chordata" rank="genus">Austrolabrus</taxonomicName>
maculatus
</emphasis>
,
<emphasis italics="true" pageId="3" pageNumber="131">
<taxonomicName authorityName="Ramsay &amp; Ogilby" authorityYear="1888" box="[1314,1452,1484,1506]" class="Actinopterygii" family="Labridae" genus="Eupetrichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="131" phylum="Chordata" rank="genus">Eupetrichthys</taxonomicName>
angustipes, Siphonognathus caninus
</emphasis>
, and
<emphasis box="[1269,1412,1515,1536]" italics="true" pageId="3" pageNumber="131">S. attenuatus</emphasis>
, and the gobiesocids
<emphasis box="[997,1196,1543,1565]" italics="true" pageId="3" pageNumber="131">
<taxonomicName authorityName="Cloquet" authorityYear="1816" box="[997,1065,1543,1565]" class="Actinopterygii" family="Gobiesocidae" genus="Alabes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="131" phylum="Chordata" rank="genus">Alabes</taxonomicName>
occidentalis
</emphasis>
and Genus A sp. 2 (
<emphasis box="[1473,1529,1545,1566]" italics="true" pageId="3" pageNumber="131">sensu</emphasis>
Hutchins, 2008).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="131" type="etymology">
<paragraph blockId="3.[859,1530,1627,1739]" pageId="3" pageNumber="131">
<emphasis bold="true" box="[859,1000,1627,1650]" italics="true" pageId="3" pageNumber="131">Etymology.—</emphasis>
From the Latin
<taxonomicName authorityName="Kevin W. Conway &amp; Glenn I. Moore &amp; Adam P. Summers" authorityYear="2017" box="[1177,1243,1627,1649]" class="Actinopterygii" family="Gobiesocidae" genus="Nettorhamphos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="131" phylum="Chordata" rank="species" species="radula">
<emphasis box="[1177,1243,1627,1649]" italics="true" pageId="3" pageNumber="131">radula</emphasis>
</taxonomicName>
, a tool used for scraping. In reference to the many tiny conical teeth on the lingual surface of the premaxilla and dentary, which are reminiscent of the
<taxonomicName authorityName="Kevin W. Conway &amp; Glenn I. Moore &amp; Adam P. Summers" authorityYear="2017" box="[934,1003,1716,1739]" class="Actinopterygii" family="Gobiesocidae" genus="Nettorhamphos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="131" phylum="Chordata" rank="species" species="radula">radula</taxonomicName>
of a snail. A noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>