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<document id="014836C9BDC1AEA3B8F8A0B9DF5D9ECB" ID-CLB-Dataset="26202" ID-DOI="10.11646/zootaxa.4630.1.1" ID-GBIF-Dataset="db6a16a6-0fe8-4987-8ad5-42223825fcd7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3336635" ID-ZooBank="8D375836-CCBA-473C-836F-6ABD44B4F881" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1563153150055" checkinUser="plazi" docAuthor="Olmi, Massimo, Copeland, Robert S. &amp; Noort, Simon Van" docDate="2019" docId="03E987E2FE692B8FFF3ED056FA71FD2C" docLanguage="en" docName="zootaxa.4630.1.1.pdf.imf" docOrigin="Zootaxa 4630 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Gonatopus bellicosus Olmi &amp; Copeland &amp; Noort 2019, sp. nov." docType="treatment" docVersion="41" lastPageNumber="456" masterDocId="FFD0FF9AFFAF2A48FFA9D64EFFE1FF87" masterDocTitle="Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea)" masterLastPageNumber="619" masterPageNumber="1" pageNumber="455" updateTime="1721735765381" updateUser="karina">
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<mods:titleInfo id="7BF89960DD26F2F9335D8D546A9D207D">
<mods:title id="312C82B98ED4A1FA0B4592463D912EB3">Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea)</mods:title>
</mods:titleInfo>
<mods:name id="ED272F681FEABAED7EC960CA2D4A4171" type="personal">
<mods:role id="86C86F08EC27750FC6F77EFEB579E4E6">
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<mods:namePart id="C931CB9A013E3B9FD73BC430DD549A71">Olmi, Massimo</mods:namePart>
</mods:name>
<mods:name id="7EDE811D3C2157FB0851B41C371CE723" type="personal">
<mods:role id="BA17391DD30C48D5C53830E7045A03CB">
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<mods:namePart id="43CBCA3B007BADE7061CB1CAAD8F9669">Copeland, Robert S.</mods:namePart>
</mods:name>
<mods:name id="CD577B18E59FF084EF6D19387DBD7AD0" type="personal">
<mods:role id="9E74DA92C1E9F202D44E18C4440B368B">
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<mods:namePart id="5A0ACBE90BA1E41EE6A36B3DAD723935">Noort, Simon Van</mods:namePart>
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<mods:typeOfResource id="A12F369C65ACFF983D6CA9CB2F45838C">text</mods:typeOfResource>
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<mods:titleInfo id="CCCC14DEE5E5E06A1B23CB307A15B770">
<mods:title id="90CFBABAE01D5D582751F4665E9A0E9D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F8D3320DAE475F00D6C914D99A44B716">
<mods:date id="C4AD8D4CC284883D0BD8463707867EB1">2019</mods:date>
<mods:detail id="71B9855EBB9EF177528ABDD2B9A20762" type="pubDate">
<mods:number id="70A5C15CE616919271E613C74E176CDA">2019-07-10</mods:number>
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<mods:detail id="A199DD0829D06EA04BE599F20E07F553" type="volume">
<mods:number id="28D63F21D4F04ACEE5EB92CD70F62619">4630</mods:number>
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<mods:number id="B29C3FE619E18FD74BDD8278EA7C92DA">1</mods:number>
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<mods:classification id="E22291B1F2615561BBE4FD1B26E40F7E">journal article</mods:classification>
<mods:identifier id="500C957BE60E97EC0DBFAD5A8D96A857" type="CLB-Dataset">26202</mods:identifier>
<mods:identifier id="1C1C58B0F6E967DFB3129D8F9DFE1981" type="DOI">10.11646/zootaxa.4630.1.1</mods:identifier>
<mods:identifier id="73BC175835B526750EF2828E48E6AABD" type="GBIF-Dataset">db6a16a6-0fe8-4987-8ad5-42223825fcd7</mods:identifier>
<mods:identifier id="F3DE2420C47A5DB93457B5CF6AFFF1DD" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E987E2FE692B8FFF3ED056FA71FD2C" ID-GBIF-Taxon="158672642" LSID="urn:lsid:plazi:treatment:03E987E2FE692B8FFF3ED056FA71FD2C" httpUri="http://treatment.plazi.org/id/03E987E2FE692B8FFF3ED056FA71FD2C" lastPageId="455" lastPageNumber="456" pageId="454" pageNumber="455">
<subSubSection id="C35A657FFE692B8EFF3ED056FC44F9B5" box="[151,933,1559,1586]" pageId="454" pageNumber="455" type="nomenclature">
<paragraph id="8BFF36F4FE692B8EFF3ED056FC44F9B5" blockId="454.[151,933,1559,1622]" box="[151,933,1559,1586]" pageId="454" pageNumber="455">
<heading id="D0B78198FE692B8EFF3ED056FC44F9B5" bold="true" box="[151,933,1559,1586]" fontSize="11" level="1" pageId="454" pageNumber="455" reason="1">
<emphasis id="B934EAE6FE692B8EFF3ED056FC44F9B5" bold="true" box="[151,933,1559,1586]" pageId="454" pageNumber="455">
35.
<taxonomicName id="4C404D77FE692B8EFF6AD056FCDAF9B5" authority="Olmi, Copeland &amp; van Noort" authorityName="Olmi &amp; Copeland &amp; Noort" authorityYear="2019" box="[195,827,1559,1586]" class="Insecta" family="Dryinidae" genus="Gonatopus" kingdom="Animalia" order="Hymenoptera" pageId="454" pageNumber="455" phylum="Arthropoda" rank="species" species="bellicosus" status="sp. nov." superFamily="Chrysidoidea">
<emphasis id="B934EAE6FE692B8EFF6AD056FE27F9B5" bold="true" box="[195,454,1559,1586]" italics="true" pageId="454" pageNumber="455">Gonatopus bellicosus</emphasis>
Olmi, Copeland &amp; van Noort
</taxonomicName>
,
<taxonomicNameLabel id="A207579DFE692B8EFCE1D056FC44F9B5" box="[840,933,1560,1586]" pageId="454" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFE692B8EFF3ED075FDBFF9D1" box="[151,606,1595,1622]" pageId="454" pageNumber="455" type="description">
<paragraph id="8BFF36F4FE692B8EFF3ED075FDBFF9D1" blockId="454.[151,933,1559,1622]" box="[151,606,1595,1622]" pageId="454" pageNumber="455">
(
<figureCitation id="137B2A71FE692B8EFF36D075FEF9F9D2" box="[159,280,1595,1621]" captionStart="FIGURE 186" captionStartId="452.[151,250,1111,1136]" captionTargetBox="[156,1410,178,1088]" captionTargetId="figure@452.[151,1436,174,1099]" captionTargetPageId="452" captionText="FIGURE 186. Habitus of Gonatopus species holotypes. A: G. pluvialis Olmi, ♀ in dorsal view. B, C: G. ruvidus Olmi, ♀ in dorsal (B) and lateral (C) view. D: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov.: ♀ in dorsal view; mesosoma sculpture. Scale bar: A: 3.00 mm." httpUri="https://zenodo.org/record/3337048/files/figure.png" pageId="454" pageNumber="455">Figs 186D</figureCitation>
,
<figureCitation id="137B2A71FE692B8EFE8DD072FEB1F9D1" box="[292,336,1596,1622]" captionStart="FIGURE 188" captionStartId="455.[151,250,1667,1692]" captionTargetBox="[156,1431,697,1654]" captionTargetId="figure@455.[151,1436,697,1655]" captionTargetPageId="455" captionText="FIGURE 188. Gonatopus bellicosus Olmi, Copeland &amp; van Noort sp. nov., ♀ holotype: habitus in dorsal (A) and lateral (C) view; head and mesosoma in dorsal view (B); head in frontal view (D)." httpUri="https://zenodo.org/record/3337052/files/figure.png" pageId="454" pageNumber="455">188</figureCitation>
,
<figureCitation id="137B2A71FE692B8EFEF5D072FE7CF9D1" box="[348,413,1596,1622]" captionStart="FIGURE 189" captionStartId="457.[151,250,1629,1654]" captionTargetBox="[157,1429,199,1616]" captionTargetId="figure@457.[151,1436,191,1617]" captionTargetPageId="457" captionText="FIGURE 189. Habitus of Gonatopus species, ♀ holotypes. A: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov. in lateral view; mesosoma sculpture. B, C: G. marojejyanus Olmi, Copeland &amp; van Noort sp. nov. in dorsal (B) and lateral (C) view; mesosoma sculpture. D–F: G. subrufulus Olmi: head and mesosoma in dorsal view (D); habitus in lateral view (E); head in frontal view (F)." httpUri="https://zenodo.org/record/3337054/files/figure.png" pageId="454" pageNumber="455">189A</figureCitation>
,
<figureCitation id="137B2A71FE692B8EFE01D072FE0BF9D1" box="[424,490,1596,1622]" captionStart="FIGURE 191" captionStartId="460.[151,250,996,1021]" captionTargetBox="[157,1431,182,978]" captionTargetId="figure@460.[151,1436,174,984]" captionTargetPageId="460" captionText="FIGURE 191. Chelae of Gonatopus species holotypes.A: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov. B: G. danielssoni Olmi. C: G. emelyanovi Ponomarenko &amp; Olmi. D: G. ibayensis Olmi. E: G. gazensis Olmi. Scale bar: A: 0.11 mm; B, D: 0.09 mm; C: 0.08 mm; E: 0.14 mm." httpUri="https://zenodo.org/record/3337058/files/figure.png" pageId="454" pageNumber="455">191A</figureCitation>
;
<figureCitation id="137B2A71FE692B8EFE5CD072FDB5F9D1" box="[501,596,1596,1622]" captionStart="FIGURE 192" captionStartId="460.[151,250,1697,1722]" captionTargetBox="[159,1421,1114,1671]" captionTargetId="figure@460.[151,1435,1107,1685]" captionTargetPageId="460" captionText="FIGURE 192. Gonatopus species: ♀ holotypes. A, B: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov.: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal view (A); mesosoma in lateral view (B). C, D: G. danielssoni Olmi: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal (C) and lateral (D) view. E: G. emelyanovi Ponomarenko &amp; Olmi: mesosoma in lateral view. F: G. gazensis Olmi: head in frontal view. G, H: G. karooensis Olmi, Copeland &amp; van Noort sp. nov.: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal view (G); mesosoma in lateral view (H). I: G. orangensis Olmi: mesoscutum + mesoscutellum + metapectal- propodeal disc + propodeal declivity in dorsal view. Scale bar:A: 0.84 mm; B: 1.21 mm; C: 0.74 mm; D: 0.72 mm; E: 1.16 mm; F: 0.38 mm; G: 0.65 mm; H: 0.98 mm; I: 0.79 mm." httpUri="https://zenodo.org/record/3337060/files/figure.png" pageId="454" pageNumber="455">192A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFE692B8EFF3ED0CDFD60F88D" pageId="454" pageNumber="455" type="diagnosis">
<paragraph id="8BFF36F4FE692B8EFF3ED0CDFD60F88D" blockId="454.[151,1437,1667,2018]" pageId="454" pageNumber="455">
<emphasis id="B934EAE6FE692B8EFF3ED0CDFEEBF919" bold="true" box="[151,266,1667,1694]" pageId="454" pageNumber="455">Diagnosis</emphasis>
. Apterous
<specimenCount id="9D46FD7DFE692B8EFE2AD0CAFE79F919" box="[387,408,1668,1694]" pageId="454" pageNumber="455" type="female">♀</specimenCount>
of
<taxonomicName id="4C404D77FE692B8EFE12D0CAFDD7F919" authorityName="Ljungh" authorityYear="1810" box="[443,566,1668,1694]" class="Insecta" family="Dryinidae" genus="Gonatopus" kingdom="Animalia" order="Hymenoptera" pageId="454" pageNumber="455" phylum="Arthropoda" rank="genus" superFamily="Chrysidoidea">
<emphasis id="B934EAE6FE692B8EFE12D0CAFDD7F919" box="[443,566,1668,1694]" italics="true" pageId="454" pageNumber="455">Gonatopus</emphasis>
</taxonomicName>
with palpal formula 6/3; pronotum crossed by strong transverse furrow; metanotum about coplanar to mesoscutellum (
<figureCitation id="137B2A71FE692B8EFDDCD0E9FD0CF945" box="[629,749,1703,1730]" captionStart="FIGURE 192" captionStartId="460.[151,250,1697,1722]" captionTargetBox="[159,1421,1114,1671]" captionTargetId="figure@460.[151,1435,1107,1685]" captionTargetPageId="460" captionText="FIGURE 192. Gonatopus species: ♀ holotypes. A, B: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov.: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal view (A); mesosoma in lateral view (B). C, D: G. danielssoni Olmi: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal (C) and lateral (D) view. E: G. emelyanovi Ponomarenko &amp; Olmi: mesosoma in lateral view. F: G. gazensis Olmi: head in frontal view. G, H: G. karooensis Olmi, Copeland &amp; van Noort sp. nov.: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal view (G); mesosoma in lateral view (H). I: G. orangensis Olmi: mesoscutum + mesoscutellum + metapectal- propodeal disc + propodeal declivity in dorsal view. Scale bar:A: 0.84 mm; B: 1.21 mm; C: 0.74 mm; D: 0.72 mm; E: 1.16 mm; F: 0.38 mm; G: 0.65 mm; H: 0.98 mm; I: 0.79 mm." httpUri="https://zenodo.org/record/3337060/files/figure.png" pageId="454" pageNumber="455">Fig. 192B</figureCitation>
); metapectal-propodeal disc brown-black, with disc slightly humped (
<figureCitation id="137B2A71FE692B8EFEA8D085FE98F961" box="[257,377,1739,1766]" captionStart="FIGURE 189" captionStartId="457.[151,250,1629,1654]" captionTargetBox="[157,1429,199,1616]" captionTargetId="figure@457.[151,1436,191,1617]" captionTargetPageId="457" captionText="FIGURE 189. Habitus of Gonatopus species, ♀ holotypes. A: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov. in lateral view; mesosoma sculpture. B, C: G. marojejyanus Olmi, Copeland &amp; van Noort sp. nov. in dorsal (B) and lateral (C) view; mesosoma sculpture. D–F: G. subrufulus Olmi: head and mesosoma in dorsal view (D); habitus in lateral view (E); head in frontal view (F)." httpUri="https://zenodo.org/record/3337054/files/figure.png" pageId="454" pageNumber="455">Fig. 189A</figureCitation>
); enlarged claw with apex very broad (
<figureCitation id="137B2A71FE692B8EFC99D085FC45F961" box="[816,932,1739,1766]" captionStart="FIGURE 191" captionStartId="460.[151,250,996,1021]" captionTargetBox="[157,1431,182,978]" captionTargetId="figure@460.[151,1436,174,984]" captionTargetPageId="460" captionText="FIGURE 191. Chelae of Gonatopus species holotypes.A: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov. B: G. danielssoni Olmi. C: G. emelyanovi Ponomarenko &amp; Olmi. D: G. ibayensis Olmi. E: G. gazensis Olmi. Scale bar: A: 0.11 mm; B, D: 0.09 mm; C: 0.08 mm; E: 0.14 mm." httpUri="https://zenodo.org/record/3337058/files/figure.png" pageId="454" pageNumber="455">Fig. 191A</figureCitation>
) and one big subdistal tooth; protarsomere 5 with one short row of lamellae (
<figureCitation id="137B2A71FE692B8EFE54D0A1FD95F88D" box="[509,628,1775,1802]" captionStart="FIGURE 191" captionStartId="460.[151,250,996,1021]" captionTargetBox="[157,1431,182,978]" captionTargetId="figure@460.[151,1436,174,984]" captionTargetPageId="460" captionText="FIGURE 191. Chelae of Gonatopus species holotypes.A: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov. B: G. danielssoni Olmi. C: G. emelyanovi Ponomarenko &amp; Olmi. D: G. ibayensis Olmi. E: G. gazensis Olmi. Scale bar: A: 0.11 mm; B, D: 0.09 mm; C: 0.08 mm; E: 0.14 mm." httpUri="https://zenodo.org/record/3337058/files/figure.png" pageId="454" pageNumber="455">Fig. 191A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFE692B8FFF6ED15DFE81FE54" lastPageId="455" lastPageNumber="456" pageId="454" pageNumber="455" type="description">
<paragraph id="8BFF36F4FE692B8FFF6ED15DFC25FE28" blockId="454.[151,1437,1667,2018]" lastBlockId="455.[151,1437,153,684]" lastPageId="455" lastPageNumber="456" pageId="454" pageNumber="455">
<emphasis id="B934EAE6FE692B8EFF6ED15DFEB8F8A9" bold="true" box="[199,345,1811,1838]" pageId="454" pageNumber="455">Description.</emphasis>
<specimenCount id="9D46FD7DFE692B8EFEC8D15AFE97F8A9" box="[353,374,1812,1838]" pageId="454" pageNumber="455" type="female">♀</specimenCount>
(
<figureCitation id="137B2A71FE692B8EFE2CD15DFE1EF8AA" box="[389,511,1811,1837]" captionStart="FIGURE 186" captionStartId="452.[151,250,1111,1136]" captionTargetBox="[156,1410,178,1088]" captionTargetId="figure@452.[151,1436,174,1099]" captionTargetPageId="452" captionText="FIGURE 186. Habitus of Gonatopus species holotypes. A: G. pluvialis Olmi, ♀ in dorsal view. B, C: G. ruvidus Olmi, ♀ in dorsal (B) and lateral (C) view. D: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov.: ♀ in dorsal view; mesosoma sculpture. Scale bar: A: 3.00 mm." httpUri="https://zenodo.org/record/3337048/files/figure.png" pageId="454" pageNumber="455">Figs 186D</figureCitation>
,
<figureCitation id="137B2A71FE692B8EFDA5D15AFDD9F8A9" box="[524,568,1812,1838]" captionStart="FIGURE 188" captionStartId="455.[151,250,1667,1692]" captionTargetBox="[156,1431,697,1654]" captionTargetId="figure@455.[151,1436,697,1655]" captionTargetPageId="455" captionText="FIGURE 188. Gonatopus bellicosus Olmi, Copeland &amp; van Noort sp. nov., ♀ holotype: habitus in dorsal (A) and lateral (C) view; head and mesosoma in dorsal view (B); head in frontal view (D)." httpUri="https://zenodo.org/record/3337052/files/figure.png" pageId="454" pageNumber="455">188</figureCitation>
,
<figureCitation id="137B2A71FE692B8EFDECD15AFD69F8A9" box="[581,648,1812,1838]" captionStart="FIGURE 189" captionStartId="457.[151,250,1629,1654]" captionTargetBox="[157,1429,199,1616]" captionTargetId="figure@457.[151,1436,191,1617]" captionTargetPageId="457" captionText="FIGURE 189. Habitus of Gonatopus species, ♀ holotypes. A: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov. in lateral view; mesosoma sculpture. B, C: G. marojejyanus Olmi, Copeland &amp; van Noort sp. nov. in dorsal (B) and lateral (C) view; mesosoma sculpture. D–F: G. subrufulus Olmi: head and mesosoma in dorsal view (D); habitus in lateral view (E); head in frontal view (F)." httpUri="https://zenodo.org/record/3337054/files/figure.png" pageId="454" pageNumber="455">189A</figureCitation>
). Apterous; body length
<quantity id="4CB89B11FE692B8EFC37D15AFC19F8A9" box="[926,1016,1812,1838]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="454" pageNumber="455" unit="mm" value="3.7">3.7 mm</quantity>
. Head brown-black, except mandible testaceous and one ferruginous longitudinal stripe between antennal toruli; antenna brown (antennomere 10 missing); mesosoma brown-black, except mesoscutum testaceous; metasoma brown; legs brown, except tarsi, part of procoxa and protrochanter testaceous.Antenna clavate; antennomeres in following proportions: 11:6:25:16:13:12:9:8:7 (antennomere 10 missing in
<typeStatus id="54FB8856FE692B8EFE7DD1EDFDDAF839" box="[468,571,1955,1982]" pageId="454" pageNumber="455" type="holotype">holotype</typeStatus>
). Head (
<figureCitation id="137B2A71FE692B8EFD3ED1EDFCEBF839" box="[663,778,1955,1982]" captionStart="FIGURE 188" captionStartId="455.[151,250,1667,1692]" captionTargetBox="[156,1431,697,1654]" captionTargetId="figure@455.[151,1436,697,1655]" captionTargetPageId="455" captionText="FIGURE 188. Gonatopus bellicosus Olmi, Copeland &amp; van Noort sp. nov., ♀ holotype: habitus in dorsal (A) and lateral (C) view; head and mesosoma in dorsal view (B); head in frontal view (D)." httpUri="https://zenodo.org/record/3337052/files/figure.png" pageId="454" pageNumber="455">Fig. 188D</figureCitation>
) excavated, dull, granulate; frontal line complete; occipital carina absent; POL = 1; OL = 2.5; OOL = 9; greatest breadth of lateral ocelli longer than POL (2:1). Palpal for-mula 6/3. Pronotum shiny, finely punctate, unsculptured among punctures, crossed by strong transverse furrow. Mesoscutum shiny, unsculptured, with two strong lateral pointed apophyses (
<figureCitation id="137B2A71FE682B8FFBB9D6F3FB6BFF50" box="[1040,1162,189,215]" captionStart="FIGURE 188" captionStartId="455.[151,250,1667,1692]" captionTargetBox="[156,1431,697,1654]" captionTargetId="figure@455.[151,1436,697,1655]" captionTargetPageId="455" captionText="FIGURE 188. Gonatopus bellicosus Olmi, Copeland &amp; van Noort sp. nov., ♀ holotype: habitus in dorsal (A) and lateral (C) view; head and mesosoma in dorsal view (B); head in frontal view (D)." httpUri="https://zenodo.org/record/3337052/files/figure.png" pageId="455" pageNumber="456">Figs 188B</figureCitation>
,
<figureCitation id="137B2A71FE682B8FFB30D6F3FB3AFF50" box="[1177,1243,189,215]" captionStart="FIGURE 192" captionStartId="460.[151,250,1697,1722]" captionTargetBox="[159,1421,1114,1671]" captionTargetId="figure@460.[151,1435,1107,1685]" captionTargetPageId="460" captionText="FIGURE 192. Gonatopus species: ♀ holotypes. A, B: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov.: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal view (A); mesosoma in lateral view (B). C, D: G. danielssoni Olmi: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal (C) and lateral (D) view. E: G. emelyanovi Ponomarenko &amp; Olmi: mesosoma in lateral view. F: G. gazensis Olmi: head in frontal view. G, H: G. karooensis Olmi, Copeland &amp; van Noort sp. nov.: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal view (G); mesosoma in lateral view (H). I: G. orangensis Olmi: mesoscutum + mesoscutellum + metapectal- propodeal disc + propodeal declivity in dorsal view. Scale bar:A: 0.84 mm; B: 1.21 mm; C: 0.74 mm; D: 0.72 mm; E: 1.16 mm; F: 0.38 mm; G: 0.65 mm; H: 0.98 mm; I: 0.79 mm." httpUri="https://zenodo.org/record/3337060/files/figure.png" pageId="455" pageNumber="456">192A</figureCitation>
). Mesoscutellum shiny, flat. Metanotum about coplanar to mesoscutellum (
<figureCitation id="137B2A71FE682B8FFC82D6AFFC44FF7C" box="[811,933,225,251]" captionStart="FIGURE 188" captionStartId="455.[151,250,1667,1692]" captionTargetBox="[156,1431,697,1654]" captionTargetId="figure@455.[151,1436,697,1655]" captionTargetPageId="455" captionText="FIGURE 188. Gonatopus bellicosus Olmi, Copeland &amp; van Noort sp. nov., ♀ holotype: habitus in dorsal (A) and lateral (C) view; head and mesosoma in dorsal view (B); head in frontal view (D)." httpUri="https://zenodo.org/record/3337052/files/figure.png" pageId="455" pageNumber="456">Figs 188C</figureCitation>
,
<figureCitation id="137B2A71FE682B8FFC1AD6AFFC12FF7C" box="[947,1011,225,251]" captionStart="FIGURE 192" captionStartId="460.[151,250,1697,1722]" captionTargetBox="[159,1421,1114,1671]" captionTargetId="figure@460.[151,1435,1107,1685]" captionTargetPageId="460" captionText="FIGURE 192. Gonatopus species: ♀ holotypes. A, B: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov.: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal view (A); mesosoma in lateral view (B). C, D: G. danielssoni Olmi: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal (C) and lateral (D) view. E: G. emelyanovi Ponomarenko &amp; Olmi: mesosoma in lateral view. F: G. gazensis Olmi: head in frontal view. G, H: G. karooensis Olmi, Copeland &amp; van Noort sp. nov.: mesoscutum + mesoscutellum + metapectal-propodeal disc + propodeal declivity in dorsal view (G); mesosoma in lateral view (H). I: G. orangensis Olmi: mesoscutum + mesoscutellum + metapectal- propodeal disc + propodeal declivity in dorsal view. Scale bar:A: 0.84 mm; B: 1.21 mm; C: 0.74 mm; D: 0.72 mm; E: 1.16 mm; F: 0.38 mm; G: 0.65 mm; H: 0.98 mm; I: 0.79 mm." httpUri="https://zenodo.org/record/3337060/files/figure.png" pageId="455" pageNumber="456">192B</figureCitation>
). Metapectal-propodeal disc swollen, shiny, finely punctate, unsculptured among punctures; propodeal declivity transversely striate. Mesopleuron shiny, unsculptured. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 19:3:7:20:34. Enlarged claw (
<figureCitation id="137B2A71FE682B8FFD42D703FCBEFEE0" box="[747,863,333,359]" captionStart="FIGURE 191" captionStartId="460.[151,250,996,1021]" captionTargetBox="[157,1431,182,978]" captionTargetId="figure@460.[151,1436,174,984]" captionTargetPageId="460" captionText="FIGURE 191. Chelae of Gonatopus species holotypes.A: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov. B: G. danielssoni Olmi. C: G. emelyanovi Ponomarenko &amp; Olmi. D: G. ibayensis Olmi. E: G. gazensis Olmi. Scale bar: A: 0.11 mm; B, D: 0.09 mm; C: 0.08 mm; E: 0.14 mm." httpUri="https://zenodo.org/record/3337058/files/figure.png" pageId="455" pageNumber="456">Fig. 191A</figureCitation>
) with apex very broad, one big subdistal tooth and one row of eight lamellae + two bristles. Protarsomere 5 (
<figureCitation id="137B2A71FE682B8FFCB7D73FFC73FE0C" box="[798,914,369,395]" captionStart="FIGURE 191" captionStartId="460.[151,250,996,1021]" captionTargetBox="[157,1431,182,978]" captionTargetId="figure@460.[151,1436,174,984]" captionTargetPageId="460" captionText="FIGURE 191. Chelae of Gonatopus species holotypes.A: G. bellicosus Olmi, Copeland &amp; van Noort sp. nov. B: G. danielssoni Olmi. C: G. emelyanovi Ponomarenko &amp; Olmi. D: G. ibayensis Olmi. E: G. gazensis Olmi. Scale bar: A: 0.11 mm; B, D: 0.09 mm; C: 0.08 mm; E: 0.14 mm." httpUri="https://zenodo.org/record/3337058/files/figure.png" pageId="455" pageNumber="456">Fig. 191A</figureCitation>
) with one row of 10 lamellae situated in distal half of protarsomere 5; apex with at least 25 lamellae. Tibial spurs 1/0/1.
</paragraph>
<paragraph id="8BFF36F4FE682B8FFF6ED7F7FE81FE54" blockId="455.[151,1437,153,684]" box="[199,352,441,467]" pageId="455" pageNumber="456">♂. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C35A657FFE682B8FFF6ED792FC02FDB8" pageId="455" pageNumber="456" type="materials_examined">
<paragraph id="8BFF36F4FE682B8FFF6ED792FC02FDB8" blockId="455.[151,1437,153,684]" pageId="455" pageNumber="456">
<emphasis id="B934EAE6FE682B8FFF6ED792FE48FE70" bold="true" box="[199,425,476,503]" pageId="455" pageNumber="456">Material examined</emphasis>
.
<emphasis id="B934EAE6FE682B8FFE1FD793FE10FE70" bold="true" box="[438,497,477,503]" pageId="455" pageNumber="456">Type</emphasis>
:
<materialsCitation id="3B283CA9FE682B8FFE56D793FC02FDB8" ID-GBIF-Occurrence="2275275452" collectedFrom="low vegetation, montane rainforest" collectingDate="2003-11-18" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" latitude="-14.439333" location="Antranohofa" longLatPrecision="13" longitude="49.739334" municipality="Marojejy National Park" pageId="455" pageNumber="456" specimenCode="CASTYPE19451" specimenCount="1326" specimenCount-female="1" specimenCount-male="1325" stateProvince="Antsiranana" typeStatus="holotype">
<specimenCount id="9D46FD7DFE682B8FFE56D793FDF5FE70" box="[511,532,477,503]" pageId="455" pageNumber="456" type="female">♀</specimenCount>
<typeStatus id="54FB8856FE682B8FFDB2D793FD9CFE70" box="[539,637,477,503]" pageId="455" pageNumber="456" type="holotype">holotype</typeStatus>
(
<specimenCode id="DBE69E8FFE682B8FFD22D792FCB5FE70" box="[651,852,476,503]" pageId="455" pageNumber="456">CASTYPE19451</specimenCode>
):
<collectingCountry id="F3577664FE682B8FFCCFD793FBD3FE70" box="[870,1074,477,503]" name="Madagascar" pageId="455" pageNumber="456">MADAGASCAR</collectingCountry>
:
<collectingRegion id="4984F816FE682B8FFB93D793FB25FE70" box="[1082,1220,477,503]" country="Madagascar" name="Antsiranana" pageId="455" pageNumber="456">Antsiranana</collectingRegion>
,
<collectingMunicipality id="6B9BAC8EFE682B8FFB66D793FF2CFD9C" pageId="455" pageNumber="456">Marojejy National Park</collectingMunicipality>
,
<location id="8E9F602FFE682B8FFF71D44FFE88FD9C" LSID="urn:lsid:plazi:treatment:03E987E2FE692B8FFF3ED056FA71FD2C:8E9F602FFE682B8FFF71D44FFE88FD9C" box="[216,361,513,539]" country="Madagascar" latitude="-14.439333" longLatPrecision="13" longitude="49.739334" municipality="Marojejy National Park" name="Antranohofa" pageId="455" pageNumber="456" stateProvince="Antsiranana">Antranohofa</location>
,
<location id="8E9F602FFE682B8FFEDCD44FFBFDFD9C" LSID="urn:lsid:plazi:treatment:03E987E2FE692B8FFF3ED056FA71FD2C:8E9F602FFE682B8FFEDCD44FFBFDFD9C" box="[373,1052,513,539]" country="Madagascar" latitude="-14.439333" longLatPrecision="13" longitude="49.739334" municipality="Marojejy National Park" name="26.6 km 31 ° NNE Andapa, 10.7 km 318 ° NW Manantenina" pageId="455" pageNumber="456" stateProvince="Antsiranana">
<quantity id="4CB89B11FE682B8FFEDCD44FFE33FD9C" box="[373,466,513,539]" metricMagnitude="4" metricUnit="m" metricValue="2.66" pageId="455" pageNumber="456" unit="km" value="26.6">26.6 km</quantity>
31° NNE Andapa,
<quantity id="4CB89B11FE682B8FFD06D44FFCEDFD9C" box="[687,780,513,539]" metricMagnitude="4" metricUnit="m" metricValue="1.0699999999999998" pageId="455" pageNumber="456" unit="km" value="10.7">10.7 km</quantity>
318° NW Manantenina
</location>
,
<geoCoordinate id="EE745033FE682B8FFB80D44FFB49FD9C" box="[1065,1192,513,539]" degrees="14" direction="south" minutes="26.36" orientation="latitude" pageId="455" pageNumber="456" precision="9" value="-14.439333">14°26.36’S</geoCoordinate>
<geoCoordinate id="EE745033FE682B8FFB19D44FFAD2FD9C" box="[1200,1331,513,539]" degrees="49" direction="east" minutes="44.36" orientation="longitude" pageId="455" pageNumber="456" precision="9" value="49.739334">49°44.36’E</geoCoordinate>
,
<specimenCount id="9D46FD7DFE682B8FFA96D44EFA79FD9C" box="[1343,1432,512,539]" pageId="455" pageNumber="456" type="male">
<quantity id="4CB89B11FE682B8FFA96D44EFA79FD9C" box="[1343,1432,512,539]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="455" pageNumber="456" unit="m" value="1325.0">1325 m</quantity>
</specimenCount>
,
<collectingDate id="EFBAE9DCFE682B8FFF3ED46BFEF8FDB8" box="[151,281,549,575]" pageId="455" pageNumber="456" value="2003-11-18">
<date id="FFFE1034FE682B8FFF3ED46BFEF8FDB8" box="[151,281,549,575]" pageId="455" pageNumber="456" value="2003-11-18">18.XI.2003</date>
</collectingDate>
, on
<collectedFrom id="385C7509FE682B8FFEE1D46BFD30FDB8" box="[328,721,549,575]" pageId="455" pageNumber="456">low vegetation, montane rainforest</collectedFrom>
,
<collectorName id="26B55322FE682B8FFD75D46BFCBAFDB8" box="[732,859,549,575]" pageId="455" pageNumber="456">
B.L.
<vernacularName id="054346DAFE682B8FFCBCD46BFCBAFDB8" baseAuthorityName="Erxleben" baseAuthorityYear="1777" box="[789,859,549,575]" canonicalName="Martes pennanti" class="Mammalia" family="Mustelidae" genus="Martes" kingdom="Animalia" language="eng" order="Carnivora" pageId="455" pageNumber="456" phylum="Chordata" rank="species" species="pennanti">Fisher</vernacularName>
</collectorName>
leg. (
<collectionCode id="ED51AE31FE682B8FFC33D46BFC37FDB8" box="[922,982,549,575]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="455" pageNumber="456">CAS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFE682B8FFF6ED407FE6FFDE4" box="[199,398,585,611]" pageId="455" pageNumber="456" type="biology_ecology">
<paragraph id="8BFF36F4FE682B8FFF6ED407FE6FFDE4" blockId="455.[151,1437,153,684]" box="[199,398,585,611]" pageId="455" pageNumber="456">
<emphasis id="B934EAE6FE682B8FFF6ED407FEEBFDE4" bold="true" box="[199,266,585,611]" pageId="455" pageNumber="456">Hosts</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFE682B8FFF6ED422FE14FD00" box="[199,501,620,647]" pageId="455" pageNumber="456" type="distribution">
<paragraph id="8BFF36F4FE682B8FFF6ED422FE14FD00" blockId="455.[151,1437,153,684]" box="[199,501,620,647]" pageId="455" pageNumber="456">
<emphasis id="B934EAE6FE682B8FFF6ED422FEB8FD00" bold="true" box="[199,345,620,647]" pageId="455" pageNumber="456">Distribution</emphasis>
. Madagascar.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFE682B8FFF6ED4DFFA71FD2C" box="[199,1424,657,684]" pageId="455" pageNumber="456" type="etymology">
<paragraph id="8BFF36F4FE682B8FFF6ED4DFFA71FD2C" blockId="455.[151,1437,153,684]" box="[199,1424,657,684]" pageId="455" pageNumber="456">
<emphasis id="B934EAE6FE682B8FFF6ED4DFFEA9FD2C" bold="true" box="[199,328,657,683]" pageId="455" pageNumber="456">Etymology</emphasis>
. The species is named
<emphasis id="B934EAE6FE682B8FFDE2D4DFFD5AFD2C" box="[587,699,657,683]" italics="true" pageId="455" pageNumber="456">bellicosus</emphasis>
(Latin adjective; = aggressive), because of its aggressive aspect.
</paragraph>
</subSubSection>
</treatment>
</document>
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