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<document id="FC111868064CF2D6A324FE3E26480BE0" ID-DOI="10.5281/zenodo.178756" ID-GBIF-Dataset="f9ea8775-9b2d-4364-b20a-4738aab6a54c" ID-ISSN="1175-5326" ID-Zenodo-Dep="178756" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920987951" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2007" docId="03E987D1FFBAFFE4FF178B023696FD41" docLanguage="en" docName="zt01604p020.pdf" docOrigin="Zootaxa 1604" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Harpalus giacomazzoi subsp. giacomazzoi Kataev &amp; Wrase 1996" docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="FFD0FFA9FFB5FFF4FF8088153517FFD3" masterDocTitle="Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="16" updateTime="1698229235843" updateUser="plazi">
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<mods:title id="ADDBFF5C3DE1A8666B260C0F53779326">Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)</mods:title>
</mods:titleInfo>
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<mods:namePart id="91319CE68C0730B8F1BCD520CD68E60A">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="512DB5AAC13E23F17BA3E4E157BD333A">Liang, Hongbin</mods:namePart>
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<mods:date id="DCE5D1ABA43AD8D30B6200AF4CD6D147">2007</mods:date>
<mods:detail id="E1DDBB9AA9CB51BC450441E9EFCD2F3F" type="volume">
<mods:number id="6894E84FD1E6B6BE75016A1BFEFAE535">1604</mods:number>
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<treatment id="03E987D1FFBAFFE4FF178B023696FD41" ID-DOI="http://doi.org/10.5281/zenodo.6250248" ID-GBIF-Taxon="119373716" ID-Zenodo-Dep="6250248" LSID="urn:lsid:plazi:treatment:03E987D1FFBAFFE4FF178B023696FD41" httpUri="http://treatment.plazi.org/id/03E987D1FFBAFFE4FF178B023696FD41" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="C35A654CFFBAFFFBFF178B02343AFC87" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BFF36C7FFBAFFFBFF178B023675FCE2" blockId="15.[151,866,791,852]" box="[151,866,791,817]" pageId="15" pageNumber="16">
<heading id="D0B781ABFFBAFFFBFF178B023675FCE2" bold="true" box="[151,866,791,817]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="4C404D44FFBAFFFBFF178B023675FCE2" ID-CoL="8MJZ6" authority="Kataev &amp; Wrase, 1996" authorityName="Kataev &amp; Wrase" authorityYear="1996" box="[151,866,791,817]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="giacomazzoi" subSpecies="giacomazzoi">
<emphasis id="B934EAD5FFBAFFFBFF178B023675FCE2" bold="true" box="[151,866,791,817]" pageId="15" pageNumber="16">
<emphasis id="B934EAD5FFBAFFFBFF178B023757FCE2" bold="true" box="[151,576,791,817]" italics="true" pageId="15" pageNumber="16">Harpalus giacomazzoi giacomazzoi</emphasis>
<bibRefCitation id="EFD14B36FFBAFFFBFDC78B023675FCE2" author="Kataev" box="[583,866,791,817]" pageId="15" pageNumber="20" refString="Kataev, B. M. &amp; Wrase, D. W. (1996) A new Dicheirotrichus Duv. and a new Harpalus Latr. (Coleoptera, Carabidae) from Sichuan, China. Linzer biologische Beitrage, 28 (1), 517 - 523." type="journal article" year="1996">Kataev &amp; Wrase, 1996</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFF36C7FFBAFFFBFF178B2F343AFC87" blockId="15.[151,866,791,852]" box="[151,301,826,852]" pageId="15" pageNumber="16">
(
<figureCitation id="137B2A42FFBAFFFBFF1F8B2F3432FC87" box="[159,293,826,852]" captionStart="FIGURES 45 53" captionStartId="16.[151,269,1784,1808]" captionTargetBox="[152,1427,721,1783]" captionTargetId="figure@16.[151,1436,685,1862]" captionTargetPageId="16" captionText="FIGURES 45 53. Harpalus giacomazzoi, median lobe of aedeagus (47, 50, 53, terminal lamella). 45 50, H. g. giacomazzoi (45 47, Sichuan, Dêgê Co.; 48 50, Sichuan, Yargong); 51 53, H. g. gracilis ssp. n. (Tibet, Markam Co., holotype). 45, 47 48, 50 51, 53, dorsal aspect); 46, 49, 52, lateral aspect). Scales: A = 0.5 mm (Figs 47, 50, 53), B = 1.0 mm (Figs 45 46, 48 49, 51 52)." httpUri="https://zenodo.org/record/178763/files/figure.png" pageId="15" pageNumber="16">Figs 4553</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFBAFFFBFF178B9F308AFC72" box="[151,1437,906,929]" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8BFF36C7FFBAFFFBFF178B9F308AFC72" blockId="15.[151,1437,906,929]" box="[151,1437,906,929]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB5011E9FFBAFFFBFF178B9F308AFC72" box="[151,1437,906,929]" pageId="15" pageNumber="16">
<taxonomicName id="4C404D44FFBAFFFBFF178B9F37A0FC72" ID-CoL="3JSL6" authority="Kataev &amp; Wrase, 1996: 519" authorityName="Kataev &amp; Wrase" authorityPageNumber="519" authorityYear="1996" box="[151,695,906,929]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="giacomazzoi">
<emphasis id="B934EAD5FFBAFFFBFF178B9F349FFC72" box="[151,392,906,929]" italics="true" pageId="15" pageNumber="16">Harpalus giacomazzoi</emphasis>
<treatmentCitation id="0AE110D6FFBAFFFBFE0D8B9F37A0FC72" author="Kataev" box="[397,695,906,929]" page="519" pageId="15" pageNumber="16" year="1996">
<bibRefCitation id="EFD14B36FFBAFFFBFE0D8B9F3795FC72" author="Kataev" box="[397,642,906,929]" pageId="15" pageNumber="20" refString="Kataev, B. M. &amp; Wrase, D. W. (1996) A new Dicheirotrichus Duv. and a new Harpalus Latr. (Coleoptera, Carabidae) from Sichuan, China. Linzer biologische Beitrage, 28 (1), 517 - 523." type="journal article" year="1996">Kataev &amp; Wrase, 1996</bibRefCitation>
: 519
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54FB8865FFBAFFFBFD418B9F37E1FC72" box="[705,758,906,929]" pageId="15" pageNumber="16">Type</typeStatus>
locality:
<quantity id="4CB89B22FFBAFFFBFCDA8B9F368AFC72" box="[858,925,906,929]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="km" value="50.0">50 km</quantity>
E Batang, road LitangBatang, Sichuan,
<collectingCountry id="F3577657FFBAFFFBFAD58B9F308FFC72" box="[1365,1432,906,929]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFBAFFFBFF178BC737F0FA77" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BFF36C7FFBAFFFBFF178BC734AEFBEF" blockId="15.[151,1437,978,2004]" pageId="15" pageNumber="16">
<emphasis id="B934EAD5FFBAFFFBFF178BC73451FC3F" bold="true" box="[151,326,978,1004]" pageId="15" pageNumber="16">
<typeStatus id="54FB8865FFBAFFFBFF178BC735C3FC3F" box="[151,212,978,1004]" pageId="15" pageNumber="16">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54FB8865FFBAFFFBFED48BC734D1FC3F" box="[340,454,978,1004]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
: ɗ,
<collectingCountry id="F3577657FFBAFFFBFE748BC7372AFC3F" box="[500,573,978,1004]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
, Sichuan, road LitangBatang,
<quantity id="4CB89B22FFBAFFFBFC2F8BC736EDFC3F" box="[943,1018,978,1004]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="km" value="50.0">50 km</quantity>
E Batang,
<date id="FFFE1007FFBAFFFBFBFD8BC7302CFC3F" box="[1149,1339,978,1004]" pageId="15" pageNumber="16" value="1992-07-14" valueMax="1992-07-16" valueMin="1992-07-14">1416.VII.1992</date>
, E. Giacomazzo leg. (cWR).
<typeStatus id="54FB8865FFBAFFFBFE158BEF371BFBC7" box="[405,524,1018,1044]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
: 1 ɗ, same data as
<typeStatus id="54FB8865FFBAFFFBFD668BEF365BFBC7" box="[742,844,1018,1044]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
(cSCI); 1 ɗ, 1 Ψ, Batang,
<quantity id="4CB89B22FFBAFFFBFBFF8BEF31CCFBC7" box="[1151,1243,1018,1044]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="16" unit="m" value="2500.0">2500 m</quantity>
,
<date id="FFFE1007FFBAFFFBFB658BEF3060FBC7" box="[1253,1399,1018,1044]" pageId="15" pageNumber="16" value="1992-07-15">15.VII.1992</date>
, E. Giacomazzo leg. (cSCI).
</paragraph>
<paragraph id="8BFF36C7FFBAFFFBFF478C5F35F6FAFF" blockId="15.[151,1437,978,2004]" pageId="15" pageNumber="16">
<emphasis id="B934EAD5FFBAFFFBFF478C5F3728FBB7" bold="true" box="[199,575,1098,1124]" pageId="15" pageNumber="16">Additional material examined</emphasis>
.
<collectingCountry id="F3577657FFBAFFFBFDD08C5F378DFBB7" box="[592,666,1098,1124]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
. SICHUAN: 5 ɗ, 2 Ψ, Dêgê Co.,
<quantity id="4CB89B22FFBAFFFBFBAC8C5F31C4FBB7" box="[1068,1235,1098,1124]" metricMagnitude="3" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="15" pageNumber="16" unit="m" value="3200.0" valueMax="3300.0" valueMin="3100.0">31003300 m</quantity>
,
<date id="FFFE1007FFBAFFFBFB5F8C5F3075FBB7" box="[1247,1378,1098,1124]" pageId="15" pageNumber="16" value="1983-07-04">4.VII.1983</date>
, Niu Chunlai (
<specimenCount id="9D46FD4EFFBAFFFBFE848C673480FB5F" box="[260,407,1138,1164]" pageId="15" pageNumber="16" type="generic">6 specimens</specimenCount>
) and Wang Shuyong (
<specimenCount id="9D46FD4EFFBAFFFBFD1D8C673635FB5F" box="[669,802,1138,1164]" pageId="15" pageNumber="16" type="generic">1 specimen</specimenCount>
) leg. (
<collectionCode id="ED51AE02FFBAFFFBFCEC8C6736B7FB5F" box="[876,928,1138,1164]" pageId="15" pageNumber="16">IZB</collectionCode>
); 1 Ψ, Dêgê Co.,
<quantity id="4CB89B22FFBAFFFBFBE88C6731D3FB5F" box="[1128,1220,1138,1164]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="15" pageNumber="16" unit="m" value="3200.0">3200 m</quantity>
,
<date id="FFFE1007FFBAFFFBFB4F8C673045FB5F" box="[1231,1362,1138,1164]" pageId="15" pageNumber="16" value="1983-07-04">4.VII.1983</date>
, Chen Yuanqing leg. (
<collectionCode id="ED51AE02FFBAFFFBFECF8C8F3494FB67" box="[335,387,1178,1204]" pageId="15" pageNumber="16">IZB</collectionCode>
); 7 Ψ, Batang,
<quantity id="4CB89B22FFBAFFFBFDBC8C8F378DFB67" box="[572,666,1178,1204]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="16" unit="m" value="2500.0">2500 m</quantity>
,
<date id="FFFE1007FFBAFFFBFD288C8F3653FB67" box="[680,836,1178,1204]" pageId="15" pageNumber="16" value="1982-08-13">13.VIII.1982</date>
, Wang Shuyong leg. (
<collectionCode id="ED51AE02FFBAFFFBFBD08C8F3193FB67" box="[1104,1156,1178,1204]" pageId="15" pageNumber="16">IZB</collectionCode>
); 23 ɗ, 6 Ψ, &quot;Yun-nan: Tsekou et Se-Tchouen, Principante de Batang: Yargong, J.A. Soulie, 1907&quot; (
<collectionCode id="ED51AE02FFBAFFFBFBD38CD731A6FB0F" LSID="urn:lsid:biocol.org:col:34988" box="[1107,1201,1218,1244]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
,
<collectionCode id="ED51AE02FFBAFFFBFB448CD73012FB0F" box="[1220,1285,1218,1244]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="15" pageNumber="16">ZISP</collectionCode>
); 5 ɗ, 5 Ψ, Xiangcheng Co.,
<quantity id="4CB89B22FFBAFFFBFEE48CFF34A8FAD7" box="[356,447,1258,1284]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="15" pageNumber="16" unit="m" value="2900.0">2900 m</quantity>
,
<date id="FFFE1007FFBAFFFBFE4B8CFF3744FAD7" box="[459,595,1258,1284]" pageId="15" pageNumber="16" value="1982-06-18">18.VI.1982</date>
, Wang Shuyong (
<specimenCount id="9D46FD4EFFBAFFFBFCA38CFF36A2FAD7" box="[803,949,1258,1284]" pageId="15" pageNumber="16" type="generic">8 specimens</specimenCount>
) and Zhang Xuezhong (
<specimenCount id="9D46FD4EFFBAFFFBFB4F8CFF3075FAD7" box="[1231,1378,1258,1284]" pageId="15" pageNumber="16" type="generic">2 specimens</specimenCount>
) leg. (
<collectionCode id="ED51AE02FFBAFFFBFF208D0735C2FAFF" box="[160,213,1298,1324]" pageId="15" pageNumber="16">IZB</collectionCode>
).
</paragraph>
<paragraph id="8BFF36C7FFBAFFFBFF478D2F37F0FA77" blockId="15.[151,1437,978,2004]" pageId="15" pageNumber="16">
Questionable locality (probably mislabelled and actually Sichuan): 1 ɗ, 1 Ψ, &quot;Peking, Westberge, Exp. Stötzner&quot;, &quot;
<typeStatus id="54FB8865FFBAFFFBFE9F8D773440FAAF" box="[287,343,1378,1404]" pageId="15" pageNumber="16">type</typeStatus>
&quot; [male] or &quot;cotype&quot; [female], &quot;
<emphasis id="B934EAD5FFBAFFFBFD5A8D763643FAAF" box="[730,852,1379,1404]" italics="true" pageId="15" pageNumber="16">pekinensis</emphasis>
Schaub.&quot; [according to our data, this taxon has never been described by E. Schauberger] (
<collectionCode id="ED51AE02FFBAFFFBFD068D9F37CCFA77" box="[646,731,1418,1444]" pageId="15" pageNumber="16">OÖLL</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFBAFFFBFF478DA7349AF8E7" pageId="15" pageNumber="16" type="description">
<paragraph id="8BFF36C7FFBAFFFBFF478DA7349AF8E7" blockId="15.[151,1437,978,2004]" pageId="15" pageNumber="16">
<emphasis id="B934EAD5FFBAFFFBFF478DA7344FFA1F" bold="true" box="[199,344,1458,1484]" pageId="15" pageNumber="16">Description</emphasis>
. Size variable: body length
<quantity id="4CB89B22FFBAFFFBFD1F8DA73629FA1F" box="[671,830,1458,1484]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" metricValueMax="10.8" metricValueMin="7.8" pageId="15" pageNumber="16" unit="mm" value="9.3" valueMax="10.8" valueMin="7.8">7.810.8 mm</quantity>
. Head moderate (WHmax/WPmax = 0.550.63 and WHmin/WPmax = 0.460.52). Pronotum variable in form, from nearly oval to more or less trapezium shaped, but in all cases more strongly narrowed apicad than basad and with base notably wider than apex; basal angles narrowly or widely rounded. Elytra 1.331.45 times as long as wide, 2.282.57 times as long and 1.071.16 times as wide as pronotum; preapical sinuation weak or moderately deep. Wings reduced to scale extending to about 1/3 3/4 of elytra. Three last abdominal sternites glabrous or with scattered additional short setae. Metacoxae with or without additional setae medially. Metafemur with 410 setigerous pores along posterior margin. Median lobe of aedeagus with terminal lamella notably widened apically (
<figureCitation id="137B2A42FFBAFFFBFB288EDF303BF937" box="[1192,1324,1738,1764]" captionStart="FIGURES 45 53" captionStartId="16.[151,269,1784,1808]" captionTargetBox="[152,1427,721,1783]" captionTargetId="figure@16.[151,1436,685,1862]" captionTargetPageId="16" captionText="FIGURES 45 53. Harpalus giacomazzoi, median lobe of aedeagus (47, 50, 53, terminal lamella). 45 50, H. g. giacomazzoi (45 47, Sichuan, Dêgê Co.; 48 50, Sichuan, Yargong); 51 53, H. g. gracilis ssp. n. (Tibet, Markam Co., holotype). 45, 47 48, 50 51, 53, dorsal aspect); 46, 49, 52, lateral aspect). Scales: A = 0.5 mm (Figs 47, 50, 53), B = 1.0 mm (Figs 45 46, 48 49, 51 52)." httpUri="https://zenodo.org/record/178763/files/figure.png" pageId="15" pageNumber="16">Fig. 47, 50</figureCitation>
); internal sac with moderately wide longitudinal spiny patch and with or without two extremely small separate teeth (
<figureCitation id="137B2A42FFBAFFFBFF1F8F0F3497F8E7" box="[159,384,1818,1844]" captionStart="FIGURES 45 53" captionStartId="16.[151,269,1784,1808]" captionTargetBox="[152,1427,721,1783]" captionTargetId="figure@16.[151,1436,685,1862]" captionTargetPageId="16" captionText="FIGURES 45 53. Harpalus giacomazzoi, median lobe of aedeagus (47, 50, 53, terminal lamella). 45 50, H. g. giacomazzoi (45 47, Sichuan, Dêgê Co.; 48 50, Sichuan, Yargong); 51 53, H. g. gracilis ssp. n. (Tibet, Markam Co., holotype). 45, 47 48, 50 51, 53, dorsal aspect); 46, 49, 52, lateral aspect). Scales: A = 0.5 mm (Figs 47, 50, 53), B = 1.0 mm (Figs 45 46, 48 49, 51 52)." httpUri="https://zenodo.org/record/178763/files/figure.png" pageId="15" pageNumber="16">Figs 4546, 4849</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFBAFFFBFF478F5731A3F88F" box="[199,1204,1858,1884]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BFF36C7FFBAFFFBFF478F5731A3F88F" blockId="15.[151,1437,978,2004]" box="[199,1204,1858,1884]" pageId="15" pageNumber="16">
<emphasis id="B934EAD5FFBAFFFBFF478F573477F88F" bold="true" box="[199,352,1858,1884]" pageId="15" pageNumber="16">Distribution</emphasis>
. The nominotypical subspecies is distributed in western part of Sichuan.
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFBAFFE4FF478F7F31CCFE21" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="8BFF36C7FFBAFFE4FF478F7F31CCFE21" blockId="15.[151,1437,978,2004]" lastBlockId="16.[151,1437,152,658]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B934EAD5FFBAFFFBFF478F7F342AF857" bold="true" box="[199,317,1898,1924]" pageId="15" pageNumber="16">Variation</emphasis>
. In the specimens from Dêgê County, the pronotum is only slightly narrower than elytra (WPmax/WE = 1.071.08), the metacoxae are with many additional setae medially, the abdominal sternites are covered with scattered additional short setae, the metafemur is with 810 setigerous pores along posterior margin and the aedeagus has separate teeth in the internal sac larger than in the specimens from other known localities (
<figureCitation id="137B2A42FFA5FFE4FE9088D5348DFF09" box="[272,410,192,218]" captionStart="FIGURES 45 53" captionStartId="16.[151,269,1784,1808]" captionTargetBox="[152,1427,721,1783]" captionTargetId="figure@16.[151,1436,685,1862]" captionTargetPageId="16" captionText="FIGURES 45 53. Harpalus giacomazzoi, median lobe of aedeagus (47, 50, 53, terminal lamella). 45 50, H. g. giacomazzoi (45 47, Sichuan, Dêgê Co.; 48 50, Sichuan, Yargong); 51 53, H. g. gracilis ssp. n. (Tibet, Markam Co., holotype). 45, 47 48, 50 51, 53, dorsal aspect); 46, 49, 52, lateral aspect). Scales: A = 0.5 mm (Figs 47, 50, 53), B = 1.0 mm (Figs 45 46, 48 49, 51 52)." httpUri="https://zenodo.org/record/178763/files/figure.png" pageId="16" pageNumber="17">Figs 4546</figureCitation>
). In the specimens from Batang County including the
<typeStatus id="54FB8865FFA5FFE4FB9888D5315CFF09" box="[1048,1099,192,218]" pageId="16" pageNumber="17">type</typeStatus>
series, the pronotum is variable but usually notably narrower than elytra (WPmax/WE = 1.091.16), the metacoxae are with or without additional setae medially, the abdominal sternites are usually glabrous, at most with few very short setae on three penultimate sternites, the metafemur possesses 47 setigerous pores along posterior margin and the separate teeth in the internal sac of the aedeagus are small or absent (
<figureCitation id="137B2A42FFA5FFE4FC308975312BFEA9" box="[944,1084,352,378]" captionStart="FIGURES 45 53" captionStartId="16.[151,269,1784,1808]" captionTargetBox="[152,1427,721,1783]" captionTargetId="figure@16.[151,1436,685,1862]" captionTargetPageId="16" captionText="FIGURES 45 53. Harpalus giacomazzoi, median lobe of aedeagus (47, 50, 53, terminal lamella). 45 50, H. g. giacomazzoi (45 47, Sichuan, Dêgê Co.; 48 50, Sichuan, Yargong); 51 53, H. g. gracilis ssp. n. (Tibet, Markam Co., holotype). 45, 47 48, 50 51, 53, dorsal aspect); 46, 49, 52, lateral aspect). Scales: A = 0.5 mm (Figs 47, 50, 53), B = 1.0 mm (Figs 45 46, 48 49, 51 52)." httpUri="https://zenodo.org/record/178763/files/figure.png" pageId="16" pageNumber="17">Figs 4849</figureCitation>
). Among the specimens from Batang County, the specimens from “Yargong” are characterized by deeper preapical sinuation of elytra. The specimens examined from Xiangcheng County and “Peking” are similar to those from Batang County (WPmax/WE = 1.101.16) but the metacoxae seem to be constantly without additional setae.
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFA5FFE4FF4789EA3696FD41" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BFF36C7FFA5FFE4FF4789EA3696FD41" blockId="16.[151,1437,152,658]" pageId="16" pageNumber="17">
<emphasis id="B934EAD5FFA5FFE4FF4789EA342EFDCA" bold="true" box="[199,313,511,537]" pageId="16" pageNumber="17">Remarks</emphasis>
. The most important distinctive characters of the nominotypical subspecies are the pronotum with comparatively wide base and the terminal lamella of the aedeagus notably widened apically. This taxon is highly variable in many of its morphological characteristics and it is possible that it will be divided into two or several subspecies when more populations will be examined.
</paragraph>
</subSubSection>
</treatment>
</document>