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<document id="94A33797277BE0B35DFF361A5D8C17C4" ID-DOI="10.5281/zenodo.178756" ID-GBIF-Dataset="f9ea8775-9b2d-4364-b20a-4738aab6a54c" ID-ISSN="1175-5326" ID-Zenodo-Dep="178756" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920987951" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2007" docId="03E987D1FFB8FFF9FF178BCA31CDF8C7" docLanguage="en" docName="zt01604p020.pdf" docOrigin="Zootaxa 1604" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Harpalus kaznakovi subsp. lilliputa Kataev &amp; Liang, 2007, ssp. n." docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="FFD0FFA9FFB5FFF4FF8088153517FFD3" masterDocTitle="Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="14" updateTime="1698229235843" updateUser="plazi">
<mods:mods id="4C1CD3BF87CFB4CDA8DDA2D96615C8C2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7DA6AA84176582D374B298B3BC022612">
<mods:title id="26156915F6AD95DAF994595402A1F1E3">Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)</mods:title>
</mods:titleInfo>
<mods:name id="9EEB50BF4CBF87009AC59E9E65FDA510" type="personal">
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<mods:namePart id="1CE13E8A0F90A33FA4FA358D26B214D0">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="00472F3746A05D2117A7BAF287700A9F">Liang, Hongbin</mods:namePart>
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<mods:typeOfResource id="8E1C1D2245EE2B1410A1FD56499C3551">text</mods:typeOfResource>
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<mods:title id="F0923AC5890203042937806EC2798D55">Zootaxa</mods:title>
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<mods:part id="14B13F5DCBA6D6BF8FA6499BC2E9B003">
<mods:date id="363CDB6F005AE4E2AD6CF5427C047D92">2007</mods:date>
<mods:detail id="F8EFC681F900BDE0F3390C087A023FFF" type="volume">
<mods:number id="F6D56FEF74AF756EE84FB702F521D3EB">1604</mods:number>
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<mods:classification id="DB400CDEC75BAB893C98BD35F0038017">journal article</mods:classification>
<mods:identifier id="3208336DC2CEBA4E1BE3F176FE08D792" type="DOI">10.5281/zenodo.178756</mods:identifier>
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<mods:identifier id="6DE5395FA92061BF1E486C90117B3921" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="307D30768A5FA1C20DBC5386F783950A" type="Zenodo-Dep">178756</mods:identifier>
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<treatment id="03E987D1FFB8FFF9FF178BCA31CDF8C7" ID-DOI="http://doi.org/10.5281/zenodo.6250244" ID-GBIF-Taxon="119373719" ID-Zenodo-Dep="6250244" LSID="urn:lsid:plazi:treatment:03E987D1FFB8FFF9FF178BCA31CDF8C7" httpUri="http://treatment.plazi.org/id/03E987D1FFB8FFF9FF178BCA31CDF8C7" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="C35A654CFFB8FFF9FF178BCA3492FBCF" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BFF36C7FFB8FFF9FF178BCA3753FC2A" blockId="13.[151,580,991,1052]" box="[151,580,991,1017]" pageId="13" pageNumber="14">
<heading id="D0B781ABFFB8FFF9FF178BCA3753FC2A" bold="true" box="[151,580,991,1017]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B934EAD5FFB8FFF9FF178BCA3753FC2A" bold="true" box="[151,580,991,1017]" pageId="13" pageNumber="14">
<taxonomicName id="4C404D44FFB8FFF9FF178BCA34E7FC2A" ID-CoL="8MC2N" box="[151,496,991,1017]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="kaznakovi" status="subsp. nov." subSpecies="lilliputa">
<emphasis id="B934EAD5FFB8FFF9FF178BCA34E7FC2A" bold="true" box="[151,496,991,1017]" italics="true" pageId="13" pageNumber="14">Harpalus kaznakovi lilliputa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757AEFFB8FFF9FE768BCA3753FC2A" box="[502,580,991,1017]" pageId="13" pageNumber="14" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF36C7FFB8FFF9FF178C173492FBCF" blockId="13.[151,580,991,1052]" box="[151,389,1026,1052]" pageId="13" pageNumber="14">
(
<figureCitation id="137B2A42FFB8FFF9FF1F8C173431FBCF" box="[159,294,1026,1052]" captionStart="FIGURES 25 29" captionStartId="12.[151,269,1777,1801]" captionTargetBox="[153,1389,532,1776]" captionTargetId="figure@12.[151,1436,532,1843]" captionTargetPageId="12" captionText="FIGURES 25 29. Harpalus kaznakovi. 25 27, H. k. kaznakovi (25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28 29, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 25 26, 28, habitus, 27 29, left metepisternum. Scales = 1.0 mm (A: Figs 25 26, 28; B: 27, 29)." httpUri="https://zenodo.org/record/178761/files/figure.png" pageId="13" pageNumber="14">Figs 2829</figureCitation>
,
<figureCitation id="137B2A42FFB8FFF9FEB28C173469FBCF" box="[306,382,1026,1052]" captionStart="FIGURES 30 44" captionStartId="14.[151,269,1898,1922]" captionTargetBox="[186,1385,219,1897]" captionTargetId="figure@14.[151,1436,212,1938]" captionTargetPageId="14" captionText="FIGURES 30 44. Harpalus kaznakovi, median lobe of aedeagus (32 33, 38, 41, 44, terminal lamella). 30 41, H. k. kaznakovi (30 38, Sichuan, Garzê Co.; 39 41, Sichuan, Dêgê Co.). 42 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 34, 36, 38 39, 41 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 33, 38, 41, 44), B = 1.0 mm (Figs 30 31, 34 37, 39 40, 42 43)." httpUri="https://zenodo.org/record/178762/files/figure.png" pageId="13" pageNumber="14">4244</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFB8FFF9FF178C4734FAFB6F" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BFF36C7FFB8FFF9FF178C4734FAFB6F" blockId="13.[151,1437,1106,1812]" pageId="13" pageNumber="14">
<emphasis id="B934EAD5FFB8FFF9FF178C473453FBBF" bold="true" box="[151,324,1106,1132]" pageId="13" pageNumber="14">
<typeStatus id="54FB8865FFB8FFF9FF178C4735C5FBBF" box="[151,210,1106,1132]" pageId="13" pageNumber="14">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54FB8865FFB8FFF9FED48C4734D1FBBF" box="[340,454,1106,1132]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
: 1 ɗ,
<collectingCountry id="F3577657FFB8FFF9FD898C473745FBBF" box="[521,594,1106,1132]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
, Tibet, Zogang Co.,
<quantity id="4CB89B22FFB8FFF9FCC98C4736B2FBBF" box="[841,933,1106,1132]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="14" unit="m" value="3800.0">3800 m</quantity>
,
<date id="FFFE1007FFB8FFF9FC308C473124FBBF" box="[944,1075,1106,1132]" pageId="13" pageNumber="14" value="1976-07-03">3.VII.1976</date>
, Zhang Xuezhong leg. (
<collectionCode id="ED51AE02FFB8FFF9FACD8C473095FBBF" box="[1357,1410,1106,1132]" pageId="13" pageNumber="14">IZB</collectionCode>
).
<typeStatus id="54FB8865FFB8FFF9FF178C6F3419FB47" box="[151,270,1146,1172]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
: 1 ɗ, same data as
<typeStatus id="54FB8865FFB8FFF9FE7F8C6F377FFB47" box="[511,616,1146,1172]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
,
<date id="FFFE1007FFB8FFF9FDF78C6F37EFFB47" box="[631,760,1146,1172]" pageId="13" pageNumber="14" value="1976-07-02">2.VII.1976</date>
(
<collectionCode id="ED51AE02FFB8FFF9FC8B8C6F3628FB47" box="[779,831,1146,1172]" pageId="13" pageNumber="14">IZB</collectionCode>
); 2 ɗ, Lhünzhub Co.,
<quantity id="4CB89B22FFB8FFF9FBD38C6F31A5FB47" box="[1107,1202,1146,1172]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="13" pageNumber="14" unit="m" value="4200.0">4200 m</quantity>
,
<date id="FFFE1007FFB8FFF9FB418C6F305EFB47" box="[1217,1353,1146,1172]" pageId="13" pageNumber="14" value="1960-06-13">13.VI.1960</date>
, Wang Chunguang leg. (
<collectionCode id="ED51AE02FFB8FFF9FEE08CB73484FB6F" box="[352,403,1186,1212]" pageId="13" pageNumber="14">IZB</collectionCode>
,
<collectionCode id="ED51AE02FFB8FFF9FE1F8CB734F7FB6F" box="[415,480,1186,1212]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="13" pageNumber="14">ZISP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFB8FFF9FF478CDF34C2F9A7" pageId="13" pageNumber="14" type="description">
<paragraph id="8BFF36C7FFB8FFF9FF478CDF34C2F9A7" blockId="13.[151,1437,1106,1812]" pageId="13" pageNumber="14">
<emphasis id="B934EAD5FFB8FFF9FF478CDF344FFB37" bold="true" box="[199,344,1226,1252]" pageId="13" pageNumber="14">Description</emphasis>
. Size very small: body length
<quantity id="4CB89B22FFB8FFF9FD478CDF364EFB37" box="[711,857,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.8" metricValueMin="5.2" pageId="13" pageNumber="14" unit="mm" value="5.5" valueMax="5.8" valueMin="5.2">5.25.8 mm</quantity>
. Habitus:
<figureCitation id="137B2A42FFB8FFF9FC538CDF313CFB37" box="[979,1067,1226,1252]" captionStart="FIGURES 25 29" captionStartId="12.[151,269,1777,1801]" captionTargetBox="[153,1389,532,1776]" captionTargetId="figure@12.[151,1436,532,1843]" captionTargetPageId="12" captionText="FIGURES 25 29. Harpalus kaznakovi. 25 27, H. k. kaznakovi (25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28 29, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 25 26, 28, habitus, 27 29, left metepisternum. Scales = 1.0 mm (A: Figs 25 26, 28; B: 27, 29)." httpUri="https://zenodo.org/record/178761/files/figure.png" pageId="13" pageNumber="14">Fig. 28</figureCitation>
. Head relatively large (in male WHmax/WPmax = 0.690.71 and WHmin/WPmax = 0.580.60), with convex eyes (WHmax/WHmin = 1.191.20). Pronotum relatively narrow, 1.421.50 as wide as long, with sides converging posteriad in basal half and basal angles rounded at apices; basal foveae small and narrow; base of pronotum usually almost impunctate, at most with few very fine punctures around basal foveae and along lateral furrows basally. Elytra relatively long, 1.321.42 times as long as wide, 2.402.62 times as long and 1.171.34 times as wide as pronotum; humeral denticle small; preapical sinuation moderate, parascutellar pore present; each third interval with one discal pore. Metepisterna (
<figureCitation id="137B2A42FFB8FFF9FDED8DF737D3FA2F" box="[621,708,1506,1532]" captionStart="FIGURES 25 29" captionStartId="12.[151,269,1777,1801]" captionTargetBox="[153,1389,532,1776]" captionTargetId="figure@12.[151,1436,532,1843]" captionTargetPageId="12" captionText="FIGURES 25 29. Harpalus kaznakovi. 25 27, H. k. kaznakovi (25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28 29, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 25 26, 28, habitus, 27 29, left metepisternum. Scales = 1.0 mm (A: Figs 25 26, 28; B: 27, 29)." httpUri="https://zenodo.org/record/178761/files/figure.png" pageId="13" pageNumber="14">Fig. 29</figureCitation>
) approximately as long as wide and more strongly than in
<taxonomicName id="4C404D44FFB8FFF9FAFF8DF6349EF9F7" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="kaznakovi" subSpecies="kaznakovi">
<emphasis id="B934EAD5FFB8FFF9FAFF8DF6349EF9F7" italics="true" pageId="13" pageNumber="14">H. kaznakovi kaznakovi</emphasis>
</taxonomicName>
narrowed posteriad. Metacoxae without additional setae. Median lobe of aedeagus (
<figureCitation id="137B2A42FFB8FFF9FAE78E1F35F5F99F" captionStart="FIGURES 30 44" captionStartId="14.[151,269,1898,1922]" captionTargetBox="[186,1385,219,1897]" captionTargetId="figure@14.[151,1436,212,1938]" captionTargetPageId="14" captionText="FIGURES 30 44. Harpalus kaznakovi, median lobe of aedeagus (32 33, 38, 41, 44, terminal lamella). 30 41, H. k. kaznakovi (30 38, Sichuan, Garzê Co.; 39 41, Sichuan, Dêgê Co.). 42 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 34, 36, 38 39, 41 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 33, 38, 41, 44), B = 1.0 mm (Figs 30 31, 34 37, 39 40, 42 43)." httpUri="https://zenodo.org/record/178762/files/figure.png" pageId="13" pageNumber="14">Figs 4243</figureCitation>
) notably bent just after basal bulb; terminal lamella (
<figureCitation id="137B2A42FFB8FFF9FCE18E2736AEF99F" box="[865,953,1586,1612]" captionStart="FIGURES 30 44" captionStartId="14.[151,269,1898,1922]" captionTargetBox="[186,1385,219,1897]" captionTargetId="figure@14.[151,1436,212,1938]" captionTargetPageId="14" captionText="FIGURES 30 44. Harpalus kaznakovi, median lobe of aedeagus (32 33, 38, 41, 44, terminal lamella). 30 41, H. k. kaznakovi (30 38, Sichuan, Garzê Co.; 39 41, Sichuan, Dêgê Co.). 42 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 34, 36, 38 39, 41 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 33, 38, 41, 44), B = 1.0 mm (Figs 30 31, 34 37, 39 40, 42 43)." httpUri="https://zenodo.org/record/178762/files/figure.png" pageId="13" pageNumber="14">Fig. 44</figureCitation>
) comparatively short, with sides usually slightly converging apicad.
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFB8FFF9FF478E973117F94F" box="[199,1024,1666,1692]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BFF36C7FFB8FFF9FF478E973117F94F" blockId="13.[151,1437,1106,1812]" box="[199,1024,1666,1692]" pageId="13" pageNumber="14">
<emphasis id="B934EAD5FFB8FFF9FF478E973477F94F" bold="true" box="[199,352,1666,1692]" pageId="13" pageNumber="14">Distribution</emphasis>
. This new subspecies occurs in eastern and central Tibet.
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFB8FFF9FF478EBF314DF917" box="[199,1114,1706,1732]" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BFF36C7FFB8FFF9FF478EBF314DF917" blockId="13.[151,1437,1106,1812]" box="[199,1114,1706,1732]" pageId="13" pageNumber="14">
<emphasis id="B934EAD5FFB8FFF9FF478EBF345BF917" bold="true" box="[199,332,1706,1732]" pageId="13" pageNumber="14">Etymology</emphasis>
. The subspecies epithet refers to the small body of this new taxon.
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFB8FFF9FF478EC731CDF8C7" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BFF36C7FFB8FFF9FF478EC731CDF8C7" blockId="13.[151,1437,1106,1812]" pageId="13" pageNumber="14">
<emphasis id="B934EAD5FFB8FFF9FF478EC7342EF93F" bold="true" box="[199,313,1746,1772]" pageId="13" pageNumber="14">Remarks</emphasis>
. The new subspecies distinctly differs from the nominotypical one in the narrower metepisterna, the smaller body and the terminal lamella of aedeagus more clearly converging apicad.
</paragraph>
</subSubSection>
</treatment>
</document>