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<document id="C8FD9A9772E94757DF722CFB32B55464" ID-DOI="10.11646/zootaxa.5405.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10603267" ID-ZooBank="EA51F733-DA41-4750-8DDB-F8B23837C5B6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1706773131114" checkinUser="plazi" docAuthor="Bayartogtokh, Badamdorj &amp; Bae, Yang-Seop" docDate="2024" docId="03E987B6FF8E0D1AFF110CB5FEAB80EB" docLanguage="en" docName="zootaxa.5405.2.1.pdf" docOrigin="Zootaxa 5405 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5405.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Humerobates ulleungdoensis Bayartogtokh &amp; Bae 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="161" masterDocId="FFD0FFCEFF890D10FF860E3BFFFA8226" masterDocTitle="New findings of poronotic oribatid mites (Acari: Oribatida) from Korea" masterLastPageNumber="184" masterPageNumber="151" pageNumber="158" updateTime="1707135444103" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:titleInfo id="3E0A976AD0B9CDE769AF2B6400DD1F20">
<mods:title id="8F8AB85FDD9F47F6D1356C13C10ED3FA">New findings of poronotic oribatid mites (Acari: Oribatida) from Korea</mods:title>
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<mods:name id="0B66E5E435238BFC920DB5E5732C576D" type="personal">
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<mods:roleTerm id="BD0148970EEEEB84E3C0E1DDC3CA8FDD">Author</mods:roleTerm>
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<mods:namePart id="5B0C924C387B032D097BA5F390690E15">Bayartogtokh, Badamdorj</mods:namePart>
<mods:affiliation id="BDC7E2AF51232941C20797DBF10E9DAE">Department of Biology, School of Arts and Sciences, National University of Mongolia, Ulaanbaatar, 14201 Mongolia</mods:affiliation>
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<mods:namePart id="5C5EAA614416693308C7886568E1775B">Bae, Yang-Seop</mods:namePart>
<mods:affiliation id="C7F329A7D8D373985A5CC514F09FA6D2">College of Life Sciences and Bioengineering, Incheon National University, 119 Academy-ro, Yeonsu-gu, Incheon, 22012, South Korea</mods:affiliation>
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<mods:title id="4BCE83E6514DB0C5B3FC4013E2454E5D">Zootaxa</mods:title>
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<mods:date id="2710DCDB0DD7D1822F12EB4282268626">2024</mods:date>
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<mods:number id="1555BEA5BD3A6BAD9B81988E25352C81">2024-01-30</mods:number>
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<mods:number id="356370A7963CBE22814D8769CFBACF2A">5405</mods:number>
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<mods:url id="9FD8EFF5E6E263ED1C04BEBA3BB35311">http://dx.doi.org/10.11646/zootaxa.5405.2.1</mods:url>
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<mods:classification id="465D362F97D8FD61D60578B3BA146E7D">journal article</mods:classification>
<mods:identifier id="1A5787C980CB12DF6B24A43D6B791B64" type="DOI">10.11646/zootaxa.5405.2.1</mods:identifier>
<mods:identifier id="80F6C2E428CAD60F2D1FB26C6CCF9B6F" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E987B6FF8E0D1AFF110CB5FEAB80EB" ID-DOI="http://doi.org/10.5281/zenodo.10603255" ID-Zenodo-Dep="10603255" LSID="urn:lsid:plazi:treatment:03E987B6FF8E0D1AFF110CB5FEAB80EB" httpUri="http://treatment.plazi.org/id/03E987B6FF8E0D1AFF110CB5FEAB80EB" lastPageId="10" lastPageNumber="161" pageId="7" pageNumber="158">
<subSubSection id="C35A652BFF8E0D17FF110CB5FDA7808F" box="[151,605,654,681]" pageId="7" pageNumber="158" type="nomenclature">
<paragraph id="8BFF36A0FF8E0D17FF110CB5FDA7808F" blockId="7.[151,605,654,717]" box="[151,605,654,681]" pageId="7" pageNumber="158">
<heading id="D0B781CCFF8E0D17FF110CB5FDA7808F" bold="true" box="[151,605,654,681]" fontSize="11" level="1" pageId="7" pageNumber="158" reason="1">
<emphasis id="B934EAB2FF8E0D17FF110CB5FDA7808F" bold="true" box="[151,605,654,681]" pageId="7" pageNumber="158">
<taxonomicName id="4C404D23FF8E0D17FF110CB5FE02808F" authority="Bayartogtokh &amp; Bae, 2024" authorityName="Bayartogtokh &amp; Bae" authorityYear="2024" box="[151,504,654,681]" class="Arachnida" family="Humerobatidae" genus="Humerobates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="ulleungdoensis" status="sp. nov.">
<emphasis id="B934EAB2FF8E0D17FF110CB5FE02808F" bold="true" box="[151,504,654,681]" italics="true" pageId="7" pageNumber="158">Humerobates ulleungdoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757C9FF8E0D17FE790CB4FDA7808F" box="[511,605,655,681]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A652BFF8E0D17FF110C88FB4D842F" pageId="7" pageNumber="158" type="description">
<paragraph id="8BFF36A0FF8E0D17FF110C88FEBF80EB" blockId="7.[151,605,654,717]" box="[151,325,690,717]" pageId="7" pageNumber="158">
(
<figureCitation id="137B2A25FF8E0D17FF190C88FEF080EB" box="[159,266,690,717]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[259,1328,181,1912]" captionTargetId="figure-20@8.[250,1337,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Line drawing images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603281" httpUri="https://zenodo.org/record/10603281/files/figure.png" pageId="7" pageNumber="158">Figures 5</figureCitation>
&amp;
<figureCitation id="137B2A25FF8E0D17FEAB0C88FEC680EB" box="[301,316,691,717]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[304,1283,183,1919]" captionTargetId="figure-17@9.[304,1283,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Compound microscopic images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603283" httpUri="https://zenodo.org/record/10603283/files/figure.png" pageId="7" pageNumber="158">6</figureCitation>
)
</paragraph>
<paragraph id="8BFF36A0FF8E0D17FF110CC0FBE281EF" blockId="7.[151,1437,762,2013]" pageId="7" pageNumber="158">
<emphasis id="B934EAB2FF8E0D17FF110CC0FEF38132" box="[151,265,763,788]" italics="true" pageId="7" pageNumber="158">Diagnosis</emphasis>
. Body length 502–567 μm, width 378–421. Rostrum bidentate. Prolamellar ridge absent. All prodorsal setae long, distinctly barbed. Bothridial seta with short peduncle and large, clavate head, with spiculate surface. Lamella narrow, with distinct cusp like large lateral tooth; translamella absent. Tutorium large, narrowed basally, with longitudinal striations, distally with prominent cusp, not dentate. Notogastral setae represented by their alveoli; pteromorph large, curved ventrad, with line of desclerotization; porose areas circular or subcircular, with indistinct rims. Anal and adanal setae represented by their alveoli; legs heterotridactylous.
</paragraph>
<paragraph id="8BFF36A0FF8E0D17FF410DE8FE8A8637" blockId="7.[151,1437,762,2013]" pageId="7" pageNumber="158">
<emphasis id="B934EAB2FF8E0D17FF410DE8FE9381CA" box="[199,361,979,1004]" italics="true" pageId="7" pageNumber="158">Measurements</emphasis>
. Body length: 502–567 (536) μm, width of notogaster 378–421 (396) μm, length of notogaster 454–502 (471) μm.
</paragraph>
<paragraph id="8BFF36A0FF8E0D17FF410A20FA7D867F" blockId="7.[151,1437,762,2013]" pageId="7" pageNumber="158">
<emphasis id="B934EAB2FF8E0D17FF410A20FEBF8612" box="[199,325,1051,1076]" italics="true" pageId="7" pageNumber="158">Integument</emphasis>
. Body colour yellowish brown; microtuberculate over whole body and leg segments; excrescences of cerotegument granular; restricted to region between pteromorph, pedotectum I, tutorium and lateral side of body.
</paragraph>
<paragraph id="8BFF36A0FF8E0D17FF410A58FE50875F" blockId="7.[151,1437,762,2013]" pageId="7" pageNumber="158">
<emphasis id="B934EAB2FF8E0D17FF410A58FEBE865B" box="[199,324,1123,1149]" italics="true" pageId="7" pageNumber="158">Prodorsum</emphasis>
(
<figureCitation id="137B2A25FF8E0D17FED20A58FE20865B" box="[340,474,1122,1149]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[259,1328,181,1912]" captionTargetId="figure-20@8.[250,1337,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Line drawing images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603281" httpUri="https://zenodo.org/record/10603281/files/figure.png" pageId="7" pageNumber="158">Figs. 5A, C</figureCitation>
,
<figureCitation id="137B2A25FF8E0D17FE610A58FDD4865B" box="[487,558,1123,1149]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[304,1283,183,1919]" captionTargetId="figure-17@9.[304,1283,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Compound microscopic images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603283" httpUri="https://zenodo.org/record/10603283/files/figure.png" pageId="7" pageNumber="158">6A, C</figureCitation>
). Rostrum bidentate; prolamellar ridge absent. Rostral seta (
<emphasis id="B934EAB2FF8E0D17FB590A58FB0D865A" box="[1247,1271,1123,1148]" italics="true" pageId="7" pageNumber="158">ro</emphasis>
) barbed, ~109 μm long. Lamella narrow, with distinct cusp like large lateral tooth; translamella absent. Lamellar seta (
<emphasis id="B934EAB2FF8E0D17FA920ABCFAD28687" box="[1300,1320,1159,1185]" italics="true" pageId="7" pageNumber="158">le</emphasis>
) ~147 μm long, barbed. Interlamellar seta (
<emphasis id="B934EAB2FF8E0D17FD850A90FDE386E2" box="[515,537,1195,1220]" italics="true" pageId="7" pageNumber="158">in</emphasis>
) ~155 μm long, barbed. Exobothridial seta (ex) ~38 μm long, barbed. Bothridial seta (
<emphasis id="B934EAB2FF8E0D17FF520AF4FF1786CF" box="[212,237,1231,1257]" italics="true" pageId="7" pageNumber="158">bs</emphasis>
) ~42 μm long, with short peduncle and large, clavate head, with spiculate surface. Bothridium oval, with ventrolateral and ventromedial scales. Tutorium large, ~197 μm long, narrowed basally, with longitudinal striations, distally with prominent cusp, not dentate. Humerosejugal porose area
<emphasis id="B934EAB2FF8E0D17FC370B2CFC358717" box="[945,975,1303,1329]" italics="true" pageId="7" pageNumber="158">AJ</emphasis>
present, hardly discernible; porose areas
<emphasis id="B934EAB2FF8E0D17FF110B00FF468773" box="[151,188,1339,1365]" italics="true" pageId="7" pageNumber="158">Am</emphasis>
,
<emphasis id="B934EAB2FF8E0D17FF4B0B00FF168773" box="[205,236,1339,1365]" italics="true" pageId="7" pageNumber="158">Ah</emphasis>
not evident. Genal tooth narrow, pointed distally. Pedotectum I large, with convex dorsal margin, partly covering base of seta
<emphasis id="B934EAB2FF8E0D17FE0C0B64FE59875E" box="[394,419,1375,1400]" italics="true" pageId="7" pageNumber="158">ex</emphasis>
.
</paragraph>
<paragraph id="8BFF36A0FF8E0D17FF410BB8FB4D842F" blockId="7.[151,1437,762,2013]" pageId="7" pageNumber="158">
<emphasis id="B934EAB2FF8E0D17FF410BB8FEB987BA" box="[199,323,1411,1436]" italics="true" pageId="7" pageNumber="158">Notogaster</emphasis>
(
<figureCitation id="137B2A25FF8E0D17FED40BB8FE4F87BB" box="[338,437,1410,1437]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[259,1328,181,1912]" captionTargetId="figure-20@8.[250,1337,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Line drawing images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603281" httpUri="https://zenodo.org/record/10603281/files/figure.png" pageId="7" pageNumber="158">Figs. 5A</figureCitation>
,
<figureCitation id="137B2A25FF8E0D17FE390BB8FE1F87BB" box="[447,485,1411,1437]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[304,1283,183,1919]" captionTargetId="figure-17@9.[304,1283,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Compound microscopic images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603283" httpUri="https://zenodo.org/record/10603283/files/figure.png" pageId="7" pageNumber="158">6A</figureCitation>
). Longer than wide, ratio about 1.2:1.0; anterior margin broadly rounded. Pteromorph large, curved ventrad, with line of desclerotization. Ten pairs of notogastral setae represented by their alveoli. Four pairs of porose areas circular or subcircular, with indistinct rims. Opisthonotal gland opening (
<emphasis id="B934EAB2FF8E0D17FB3A0BF0FB1A87C3" box="[1212,1248,1483,1509]" italics="true" pageId="7" pageNumber="158">gla</emphasis>
) and lyrifissures
<emphasis id="B934EAB2FF8E0D17FF110BD4FF57842E" box="[151,173,1519,1544]" italics="true" pageId="7" pageNumber="158">ia</emphasis>
,
<emphasis id="B934EAB2FF8E0D17FF3D0BD4FF2D842E" box="[187,215,1519,1544]" italics="true" pageId="7" pageNumber="158">im</emphasis>
,
<emphasis id="B934EAB2FF8E0D17FF630BD4FF01842F" box="[229,251,1519,1545]" italics="true" pageId="7" pageNumber="158">ih</emphasis>
,
<emphasis id="B934EAB2FF8E0D17FE8F0BD4FED0842E" box="[265,298,1519,1544]" italics="true" pageId="7" pageNumber="158">ips</emphasis>
and
<emphasis id="B934EAB2FF8E0D17FEE60BD4FE8C842E" box="[352,374,1519,1544]" italics="true" pageId="7" pageNumber="158">ip</emphasis>
well developed, in normal positions. Posterior notogastral tectum present.
</paragraph>
</subSubSection>
<subSubSection id="C35A652BFF8E0D17FF410828FD97856B" pageId="7" pageNumber="158" type="reference_group">
<paragraph id="8BFF36A0FF8E0D17FF410828FEB684BF" blockId="7.[151,1437,762,2013]" pageId="7" pageNumber="158">
<emphasis id="B934EAB2FF8E0D17FF410828FEAC840B" box="[199,342,1555,1581]" italics="true" pageId="7" pageNumber="158">Gnathosoma</emphasis>
(
<figureCitation id="137B2A25FF8E0D17FEE00828FE33840B" box="[358,457,1554,1581]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[259,1328,181,1912]" captionTargetId="figure-20@8.[250,1337,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Line drawing images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603281" httpUri="https://zenodo.org/record/10603281/files/figure.png" pageId="7" pageNumber="158">Figs. 5B</figureCitation>
,
<figureCitation id="137B2A25FF8E0D17FE500828FE01840B" box="[470,507,1555,1581]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[304,1283,183,1919]" captionTargetId="figure-17@9.[304,1283,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Compound microscopic images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603283" httpUri="https://zenodo.org/record/10603283/files/figure.png" pageId="7" pageNumber="158">6B</figureCitation>
). Typical for family (
<emphasis id="B934EAB2FF8E0D17FD6B0828FCF4840A" box="[749,782,1555,1580]" italics="true" pageId="7" pageNumber="158">e.g</emphasis>
.,
<bibRefCitation id="EFD14B51FF8E0D17FCA20828FB5E840B" author="Behan-Pelletier, V. M. &amp; Mahunka, S." box="[804,1188,1555,1581]" pageId="7" pageNumber="158" pagination="9 - 16" refId="ref11393" refString="Behan-Pelletier, V. M. &amp; Mahunka, S. (1993) Description of Humerobates setosus sp. n. (Acari: Humerobatidae) from South Africa. Folia Entomologica Hungarica, 44, 9 - 16." type="journal article" year="1993">Behan-Pelletier &amp; Mahunka 1993</bibRefCitation>
;
<bibRefCitation id="EFD14B51FF8E0D17FB370828FA6B840B" author="Ermilov, S. G. &amp; Sidorchuk, E. A. &amp; Rybalov, L. B." box="[1201,1425,1555,1581]" pageId="7" pageNumber="158" pagination="323 - 343" refId="ref11594" refString="Ermilov, S. G., Sidorchuk, E. A. &amp; Rybalov, L. B. (2011) New oribatid mites of the superfamily Ceratozetoidea (Acari: Oribatida) from Ethiopia. African Invertebrates, 52 (2), 323 - 343. https: // doi. org / 10.5733 / afin. 052.0208" type="journal article" year="2011">
Ermilov
<emphasis id="B934EAB2FF8E0D17FA920828FABC840B" box="[1300,1350,1555,1581]" italics="true" pageId="7" pageNumber="158">et al</emphasis>
. 2011
</bibRefCitation>
). Subcapitulum wider than long; subcapitular setae
<emphasis id="B934EAB2FF8E0D17FD4C080CFD228476" box="[714,728,1591,1616]" italics="true" pageId="7" pageNumber="158">a</emphasis>
,
<emphasis id="B934EAB2FF8E0D17FD60080CFD008476" box="[742,762,1591,1616]" italics="true" pageId="7" pageNumber="158">m</emphasis>
and
<emphasis id="B934EAB2FF8E0D17FCB4080CFCBA8477" box="[818,832,1591,1617]" italics="true" pageId="7" pageNumber="158">h</emphasis>
setiform, smooth, 21–24 μm long. Palp and chelicera typical for family; palpal setation: 0–2–1–3–9(+ω), solenidion bacilliform; cheliceral setae setiform, barbed;
<emphasis id="B934EAB2FF8E0D17FAF20860FA668453" box="[1396,1436,1627,1653]" italics="true" pageId="7" pageNumber="158">cha</emphasis>
longer than
<emphasis id="B934EAB2FF8E0D17FE9B0844FEBF84BF" box="[285,325,1663,1689]" italics="true" pageId="7" pageNumber="158">chb</emphasis>
.
</paragraph>
<paragraph id="8BFF36A0FF8E0D17FF410898FD97856B" blockId="7.[151,1437,762,2013]" pageId="7" pageNumber="158">
<emphasis id="B934EAB2FF8E0D17FF410898FE78849A" box="[199,386,1699,1725]" italics="true" pageId="7" pageNumber="158">Epimeral region</emphasis>
(
<figureCitation id="137B2A25FF8E0D17FE130898FE00849B" box="[405,506,1698,1725]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[259,1328,181,1912]" captionTargetId="figure-20@8.[250,1337,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Line drawing images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603281" httpUri="https://zenodo.org/record/10603281/files/figure.png" pageId="7" pageNumber="158">Figs. 5B</figureCitation>
,
<figureCitation id="137B2A25FF8E0D17FD8C0898FDD4849B" box="[522,558,1699,1725]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[304,1283,183,1919]" captionTargetId="figure-17@9.[304,1283,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Compound microscopic images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603283" httpUri="https://zenodo.org/record/10603283/files/figure.png" pageId="7" pageNumber="158">6B</figureCitation>
). Epimeres without muscle sigillae; apodemes
<emphasis id="B934EAB2FF8E0D17FBD70898FB6A849B" box="[1105,1168,1699,1725]" italics="true" pageId="7" pageNumber="158">apo.2</emphasis>
,
<emphasis id="B934EAB2FF8E0D17FB270898FB1F849A" box="[1185,1253,1699,1724]" italics="true" pageId="7" pageNumber="158">apo.sj</emphasis>
and
<emphasis id="B934EAB2FF8E0D17FAA50898FA9B849B" box="[1315,1377,1699,1725]" italics="true" pageId="7" pageNumber="158">apo.3</emphasis>
well developed. Epimeral setae smooth, 18–35 μm long, setal formula: 3–1–3–3; seta
<emphasis id="B934EAB2FF8E0D17FBA508FCFBC384C7" box="[1059,1081,1735,1761]" italics="true" pageId="7" pageNumber="158">lb</emphasis>
,
<emphasis id="B934EAB2FF8E0D17FBC108FCFB9984C7" box="[1095,1123,1735,1761]" italics="true" pageId="7" pageNumber="158">3b</emphasis>
,
<emphasis id="B934EAB2FF8E0D17FBF708FCFB7184C7" box="[1137,1163,1735,1761]" italics="true" pageId="7" pageNumber="158">3c</emphasis>
and
<emphasis id="B934EAB2FF8E0D17FB4408FCFB2584C7" box="[1218,1247,1735,1761]" italics="true" pageId="7" pageNumber="158">4b</emphasis>
distinctly longer than other setae. Discidium large, projected distally, but not sharp; custodium short, extending anteriad to level of posterior margin of leg acetabulum II. Circumpedal carina short, not reaching to base of pedotectum I, but complete laterally, reaching margin of ventral plate.
</paragraph>
</subSubSection>
<subSubSection id="C35A652BFF8E0D1AFF41096CFEAB80EB" lastPageId="10" lastPageNumber="161" pageId="7" pageNumber="158" type="description">
<paragraph id="8BFF36A0FF8E0D17FF41096CFBAE85FB" blockId="7.[151,1437,762,2013]" pageId="7" pageNumber="158">
<emphasis id="B934EAB2FF8E0D17FF41096CFE698556" box="[199,403,1879,1905]" italics="true" pageId="7" pageNumber="158">Anogenital region</emphasis>
(
<figureCitation id="137B2A25FF8E0D17FE23096CFDF38557" box="[421,521,1878,1905]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[259,1328,181,1912]" captionTargetId="figure-20@8.[250,1337,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Line drawing images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603281" httpUri="https://zenodo.org/record/10603281/files/figure.png" pageId="7" pageNumber="158">Figs. 5B</figureCitation>
,
<figureCitation id="137B2A25FF8E0D17FD9E096CFDC68557" box="[536,572,1879,1905]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[304,1283,183,1919]" captionTargetId="figure-17@9.[304,1283,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Compound microscopic images of Humerobates ulleundoensis sp. nov. A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma." figureDoi="http://doi.org/10.5281/zenodo.10603283" httpUri="https://zenodo.org/record/10603283/files/figure.png" pageId="7" pageNumber="158">6B</figureCitation>
). Genital and aggenital setae well developed; six pairs of genital setae,
<emphasis id="B934EAB2FF8E0D17FAFE096CFA7C8556" box="[1400,1414,1879,1904]" italics="true" pageId="7" pageNumber="158">g</emphasis>
<subScript id="17C434E5FF8E0D17FA000952FA748551" attach="left" box="[1414,1422,1897,1911]" fontSize="6" pageId="7" pageNumber="158">1</subScript>
–
<emphasis id="B934EAB2FF8E0D17FF110940FF5F85B2" box="[151,165,1915,1940]" italics="true" pageId="7" pageNumber="158">g</emphasis>
<subScript id="17C434E5FF8E0D17FF2309B6FF5785BD" attach="left" box="[165,173,1933,1947]" fontSize="6" pageId="7" pageNumber="158">4</subScript>
inserted close to anterior margin of genital plate,
<emphasis id="B934EAB2FF8E0D17FD7F0940FCFD85B2" box="[761,775,1915,1940]" italics="true" pageId="7" pageNumber="158">g</emphasis>
<subScript id="17C434E5FF8E0D17FC8109B6FCF585BD" attach="left" box="[775,783,1933,1947]" fontSize="6" pageId="7" pageNumber="158">5</subScript>
and
<emphasis id="B934EAB2FF8E0D17FCCA0940FCA085B2" box="[844,858,1915,1940]" italics="true" pageId="7" pageNumber="158">g</emphasis>
<subScript id="17C434E5FF8E0D17FCDC09B6FC9885BD" attach="left" box="[858,866,1933,1947]" fontSize="6" pageId="7" pageNumber="158">6</subScript>
close to posterior margin. Anal and adanal setae represented by their alveoli;
<emphasis id="B934EAB2FF8E0D17FE5109A4FE09859F" box="[471,499,1951,1977]" italics="true" pageId="7" pageNumber="158">ad</emphasis>
<subScript id="17C434E5FF8E0D17FE75098AFE018599" attach="left" box="[499,507,1969,1983]" fontSize="6" pageId="7" pageNumber="158">1</subScript>
and
<emphasis id="B934EAB2FF8E0D17FDB709A4FDB7859F" box="[561,589,1951,1977]" italics="true" pageId="7" pageNumber="158">ad</emphasis>
<subScript id="17C434E5FF8E0D17FDCB098AFDAF8599" attach="left" box="[589,597,1969,1983]" fontSize="6" pageId="7" pageNumber="158">2</subScript>
inserted posterior,
<emphasis id="B934EAB2FF8E0D17FCAA09A4FCB2859F" box="[812,840,1951,1977]" italics="true" pageId="7" pageNumber="158">ad</emphasis>
<subScript id="17C434E5FF8E0D17FCCE098AFCAA8599" attach="left" box="[840,848,1969,1983]" fontSize="6" pageId="7" pageNumber="158">3</subScript>
lateral to anal aperture. Lyrifissure
<emphasis id="B934EAB2FF8E0D17FB6609A4FAFE859F" box="[1248,1284,1951,1977]" italics="true" pageId="7" pageNumber="158">iad</emphasis>
located close and parallel to anterolateral margin of anal aperture. Postanal porose area small, oval.
</paragraph>
<caption id="DF3F6628FF810D18FF1109ACFC5385F5" ID-DOI="http://doi.org/10.5281/zenodo.10603281" ID-Zenodo-Dep="10603281" httpUri="https://zenodo.org/record/10603281/files/figure.png" pageId="8" pageNumber="159" startId="8.[151,250,1943,1967]" targetBox="[259,1328,181,1912]" targetPageId="8" targetType="figure">
<paragraph id="8BFF36A0FF810D18FF1109ACFC7B8589" blockId="8.[151,937,1943,2003]" box="[151,897,1943,1967]" pageId="8" pageNumber="159">
<emphasis id="B934EAB2FF810D18FF1109ACFEE98589" bold="true" box="[151,275,1943,1967]" pageId="8" pageNumber="159">FIGURE 5.</emphasis>
Line drawing images of
<taxonomicName id="4C404D23FF810D18FD9709ACFCD08589" authority="Bayartogtokh &amp; Bae, 2024" authorityName="Bayartogtokh &amp; Bae" authorityYear="2024" box="[529,810,1943,1967]" class="Arachnida" family="Humerobatidae" genus="Humerobates" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="ulleundoensis" status="sp. nov.">
<emphasis id="B934EAB2FF810D18FD9709ACFCD08589" box="[529,810,1943,1967]" italics="true" pageId="8" pageNumber="159">Humerobates ulleundoensis</emphasis>
</taxonomicName>
<emphasis id="B934EAB2FF810D18FCB709ACFC7B8589" bold="true" box="[817,897,1943,1967]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="A20757C9FF810D18FCB709ACFC7B8589" box="[817,897,1943,1967]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFF36A0FF810D18FF110980FC5385F5" blockId="8.[151,937,1943,2003]" box="[151,937,1979,2003]" pageId="8" pageNumber="159">A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma.</paragraph>
</caption>
<caption id="DF3F6628FF800D19FF1109ACFC5385F5" ID-DOI="http://doi.org/10.5281/zenodo.10603283" ID-Zenodo-Dep="10603283" httpUri="https://zenodo.org/record/10603283/files/figure.png" pageId="9" pageNumber="160" startId="9.[151,250,1943,1967]" targetBox="[304,1283,183,1919]" targetPageId="9" targetType="figure">
<paragraph id="8BFF36A0FF800D19FF1109ACFC168596" blockId="9.[151,1004,1943,2003]" box="[151,1004,1943,1968]" pageId="9" pageNumber="160">
<emphasis id="B934EAB2FF800D19FF1109ACFEE98589" bold="true" box="[151,275,1943,1967]" pageId="9" pageNumber="160">FIGURE 6.</emphasis>
Compound microscopic images of
<taxonomicName id="4C404D23FF800D19FDFA09ACFC6F8589" authority="Bayartogtokh &amp; Bae, 2024" authorityName="Bayartogtokh &amp; Bae" authorityYear="2024" box="[636,917,1943,1967]" class="Arachnida" family="Humerobatidae" genus="Humerobates" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="ulleundoensis" status="sp. nov.">
<emphasis id="B934EAB2FF800D19FDFA09ACFC6F8589" box="[636,917,1943,1967]" italics="true" pageId="9" pageNumber="160">Humerobates ulleundoensis</emphasis>
</taxonomicName>
<emphasis id="B934EAB2FF800D19FC1A09A3FC168596" bold="true" box="[924,1004,1944,1968]" pageId="9" pageNumber="160">
<taxonomicNameLabel id="A20757C9FF800D19FC1A09A3FC168596" box="[924,1004,1944,1968]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFF36A0FF800D19FF110980FC5385F5" blockId="9.[151,1004,1943,2003]" box="[151,937,1979,2003]" pageId="9" pageNumber="160">A—Dorsal view; B—Ventral view of body; C—Lateral view of proterosoma.</paragraph>
</caption>
<paragraph id="8BFF36A0FF830D1AFF410EACFE8F82F3" blockId="10.[151,1437,151,717]" pageId="10" pageNumber="161">
<emphasis id="B934EAB2FF830D1AFF410EACFF068296" box="[199,252,151,176]" italics="true" pageId="10" pageNumber="161">Legs</emphasis>
. Medial claw thicker than lateral claws, tarsi without pulvillus. Structure and setation of legs same as those of previous species.
</paragraph>
<paragraph id="8BFF36A0FF830D1AFF410EE4FB1F8343" blockId="10.[151,1437,151,717]" pageId="10" pageNumber="161">
<emphasis id="B934EAB2FF830D1AFF410EE4FE6182DF" box="[199,411,223,249]" italics="true" pageId="10" pageNumber="161">Material examined</emphasis>
.
<materialsCitation id="3B283CFDFF830D1AFE2C0EE4FB1F8343" collectingDate="2022-05-26" collectionCode="T" collectorName="Coll. &amp; G. Lee." country="South Korea" county="Engler's" latitude="37.49309" location="San" longLatPrecision="1" longitude="130.82797" municipality="Japanese" pageId="10" pageNumber="161" specimenCount="13" specimenCount-female="5" specimenCount-male="2" stateProvince="Gyeongbuk" typeStatus="holotype">
<typeStatus id="54FB8802FF830D1AFE2C0EE4FDE882DF" box="[426,530,223,249]" pageId="10" pageNumber="161" type="holotype">Holotype</typeStatus>
(female) and
<specimenCount id="9D46FD29FF830D1AFD280EE4FCBB82DF" box="[686,833,223,249]" count="6" pageId="10" pageNumber="161" type="generic" typeStatus="paratype">six paratypes</specimenCount>
(
<specimenCount id="9D46FD29FF830D1AFCD60EE4FC2182DF" box="[848,987,223,249]" count="5" pageId="10" pageNumber="161" type="female">five females</specimenCount>
and
<specimenCount id="9D46FD29FF830D1AFB940EE4FB7082DF" box="[1042,1162,223,249]" count="2" pageId="10" pageNumber="161" type="male">two males</specimenCount>
):
<location id="8E9F607BFF830D1AFB1A0EE4FB3382DF" LSID="urn:lsid:plazi:treatment:03E987B6FF8E0D1AFF110CB5FEAB80EB:8E9F607BFF830D1AFB1A0EE4FB3382DF" box="[1180,1225,223,249]" country="South Korea" county="Engler's" latitude="37.49309" longLatPrecision="1" longitude="130.82797" municipality="Japanese" name="San" pageId="10" pageNumber="161" stateProvince="Gyeongbuk">San</location>
171,
<location id="8E9F607BFF830D1AFB780EE4FAA482DF" LSID="urn:lsid:plazi:treatment:03E987B6FF8E0D1AFF110CB5FEAB80EB:8E9F607BFF830D1AFB780EE4FAA482DF" box="[1278,1374,223,249]" country="South Korea" county="Engler's" latitude="37.49309" longLatPrecision="1" longitude="130.82797" municipality="Japanese" name="Taeha-ri" pageId="10" pageNumber="161" stateProvince="Gyeongbuk">Taeha-ri</location>
,
<location id="8E9F607BFF830D1AFAEF0EE4FF1C833B" LSID="urn:lsid:plazi:treatment:03E987B6FF8E0D1AFF110CB5FEAB80EB:8E9F607BFF830D1AFAEF0EE4FF1C833B" country="South Korea" county="Engler's" latitude="37.49309" longLatPrecision="1" longitude="130.82797" municipality="Japanese" name="Seo-myeon" pageId="10" pageNumber="161" stateProvince="Gyeongbuk">Seo-myeon</location>
,
<location id="8E9F607BFF830D1AFF760F38FE7A833B" LSID="urn:lsid:plazi:treatment:03E987B6FF8E0D1AFF110CB5FEAB80EB:8E9F607BFF830D1AFF760F38FE7A833B" box="[240,384,259,285]" country="South Korea" county="Engler's" latitude="37.49309" longLatPrecision="1" longitude="130.82797" municipality="Japanese" name="Ulleung-gun" pageId="10" pageNumber="161" stateProvince="Gyeongbuk">Ulleung-gun</location>
,
<collectingRegion id="4984F842FF830D1AFE0D0F38FD33833B" box="[395,713,259,285]" country="South Korea" name="Gyeongbuk" pageId="10" pageNumber="161">North Gyeongsang Province</collectingRegion>
,
<collectingCountry id="F3577630FF830D1AFD520F38FCE1833B" box="[724,795,259,285]" name="South Korea" pageId="10" pageNumber="161">Korea</collectingCountry>
,
<geoCoordinate id="EE745067FF830D1AFCA30F39FC29833B" box="[805,979,258,285]" degrees="37" direction="north" minutes="29" orientation="latitude" pageId="10" pageNumber="161" precision="1" seconds="35.13" value="37.49309">37°29’35.13”N</geoCoordinate>
,
<geoCoordinate id="EE745067FF830D1AFC580F38FB62833B" box="[990,1176,259,285]" degrees="130" direction="east" minutes="49" orientation="longitude" pageId="10" pageNumber="161" precision="1" seconds="40.71" value="130.82797">130°49’40.71”E</geoCoordinate>
, soil and litter of mixed forest composed by
<location id="8E9F607BFF830D1AFEFB0F1CFE298367" LSID="urn:lsid:plazi:treatment:03E987B6FF8E0D1AFF110CB5FEAB80EB:8E9F607BFF830D1AFEFB0F1CFE298367" box="[381,467,295,321]" country="South Korea" county="Engler's" latitude="37.49309" longLatPrecision="1" longitude="130.82797" municipality="Japanese" name="Siebold" pageId="10" pageNumber="161" stateProvince="Gyeongbuk">Siebold</location>
humlock (
<taxonomicName id="4C404D23FF830D1AFDC80F1CFC9A8367" authority="Carriere" authorityName="Carriere" box="[590,864,295,321]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="species" species="sieboldii">
<emphasis id="B934EAB2FF830D1AFDC80F1CFD0D8367" box="[590,759,295,321]" italics="true" pageId="10" pageNumber="161">Tsuga sieboldii</emphasis>
Carrière
</taxonomicName>
),
<collectingMunicipality id="6B9BACDAFF830D1AFCF20F1CFC228367" box="[884,984,295,321]" pageId="10" pageNumber="161">Japanese</collectingMunicipality>
white pine (
<taxonomicName id="4C404D23FF830D1AFBE10F1CFEFE8343" authority="Siebold &amp; Zuccarini" authorityName="Siebold &amp; Zuccarini" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="species" species="parviflora">
<emphasis id="B934EAB2FF830D1AFBE10F1CFADB8367" box="[1127,1313,295,321]" italics="true" pageId="10" pageNumber="161">Pinus parviflora</emphasis>
Siebold &amp; Zuccarini
</taxonomicName>
) and
<collectingCounty id="629E4E2CFF830D1AFEC40F70FE5A8343" box="[322,416,331,357]" pageId="10" pageNumber="161">Engler’s</collectingCounty>
beech (
<taxonomicName id="4C404D23FF830D1AFE7F0F70FC7D8343" authority="Seemen ex Diels" authorityName="Diels" box="[505,903,331,357]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="species" species="engleriana">
<emphasis id="B934EAB2FF830D1AFE7F0F70FD458343" box="[505,703,331,357]" italics="true" pageId="10" pageNumber="161">Fagus engleriana</emphasis>
Seemen ex Diels
</taxonomicName>
),
<date id="FFFE1060FF830D1AFC1C0F70FBC88343" box="[922,1074,331,357]" pageId="10" pageNumber="161" value="2022-05-26">
<collectingDate id="EFBAE988FF830D1AFC1C0F70FBC88343" box="[922,1074,331,357]" pageId="10" pageNumber="161" value="2022-05-26">26 May 2022</collectingDate>
</date>
,
<collectorName id="26B55376FF830D1AFBBB0F70FB8F8343" box="[1085,1141,331,357]" pageId="10" pageNumber="161">Coll.</collectorName>
<collectionCode id="ED51AE65FF830D1AFBFD0F70FB748343" box="[1147,1166,331,357]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="161">T</collectionCode>
.
<collectorName id="26B55376FF830D1AFB140F70FB1F8343" box="[1170,1253,331,357]" pageId="10" pageNumber="161">G. Lee.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BFF36A0FF830D1AFF410F54FAAE838B" blockId="10.[151,1437,151,717]" pageId="10" pageNumber="161">
<emphasis id="B934EAB2FF830D1AFF410F54FEC583AF" box="[199,319,367,393]" italics="true" pageId="10" pageNumber="161">Etymology</emphasis>
. The specific name
<taxonomicName id="4C404D23FF830D1AFDB60F54FD1E83AF" authorityName="Bayartogtokh &amp; Bae" authorityYear="2024" box="[560,740,367,393]" class="Arachnida" family="Humerobatidae" genus="Humerobates" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="ulleungdoensis">
“
<emphasis id="B934EAB2FF830D1AFDBA0F54FD1E83AF" box="[572,740,367,393]" italics="true" pageId="10" pageNumber="161">ulleungdoensis</emphasis>
</taxonomicName>
” refers to the name of the island Ulleungdo, which mainly compose the Ulleung county, located in the east of Korean peninsula, from which we recovered this species.
</paragraph>
<paragraph id="8BFF36A0FF830D1AFF410F8CFDE6801B" blockId="10.[151,1437,151,717]" pageId="10" pageNumber="161">
<emphasis id="B934EAB2FF830D1AFF410F8CFED383F7" box="[199,297,439,465]" italics="true" pageId="10" pageNumber="161">Remarks</emphasis>
. The present new species clearly differs from all other known species of
<taxonomicName id="4C404D23FF830D1AFBD40F8CFB1283F7" authorityName="Sellnick" authorityYear="1928" box="[1106,1256,439,465]" class="Arachnida" family="Humerobatidae" genus="Humerobates" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="161" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAB2FF830D1AFBD40F8CFB1283F7" box="[1106,1256,439,465]" italics="true" pageId="10" pageNumber="161">Humerobates</emphasis>
</taxonomicName>
in the relatively small circular or subcircular porose areas with indistinct rims; development of large cusp-like lateral tooth of lamellar cusp; ventrally bent pteromorph with no radiating vein-like ridges; and absence of anal and adanal setae, which represented by their alveoli.
</paragraph>
<paragraph id="8BFF36A0FF830D1AFF410C7CFEAB80EB" blockId="10.[151,1437,151,717]" pageId="10" pageNumber="161">
In general appearance, this species is similar to
<taxonomicName id="4C404D23FF830D1AFD670C7CFC4D8047" authority="Bayartogtokh &amp; Bae, 2024" authorityName="Bayartogtokh &amp; Bae" authorityYear="2024" box="[737,951,583,609]" class="Arachnida" family="Humerobatidae" genus="Humerobates" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="aokii" status="sp. nov.">
<emphasis id="B934EAB2FF830D1AFD670C7CFC4D8047" box="[737,951,583,609]" italics="true" pageId="10" pageNumber="161">Humerobates aokii</emphasis>
</taxonomicName>
<emphasis id="B934EAB2FF830D1AFC380C7CFBE28047" bold="true" box="[958,1048,583,609]" pageId="10" pageNumber="161">
<taxonomicNameLabel id="A20757C9FF830D1AFC380C7CFBE28047" box="[958,1048,583,609]" pageId="10" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However, it differs from the latter species in the distinctly bidentate rostrum; relatively long interlamellar seta, which is reaching tip of rostrum; relatively smaller bothridial seta; proximally conspicuously narrowed and distally not striated tutorium, and much larger body size.
</paragraph>
</subSubSection>
</treatment>
</document>
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