203 lines
26 KiB
XML
203 lines
26 KiB
XML
<document id="DE132083EE8BCFD1DC6B08BCCA987D65" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza & Husemann, Martin" docDate="2023" docId="03DC6C76CC364C2CF7DFFB69FB48FBC8" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Metarbelodes Strand 1909" docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="8" updateTime="1699487052491" updateUser="plazi">
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<mods:title id="64D002E56907105B49E030B8CCC7A4B6">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
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<mods:name id="2D31CD49DBCFEDE65D93990E42C5BF87" type="personal">
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<mods:namePart id="F6AB1D1443ABE93B88DB3A8394F63944">Lehmann, Ingo</mods:namePart>
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<mods:nameIdentifier id="A63616971CC80057CC24805DF927D107" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
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<mods:namePart id="A462E7F43AEC7AC672E05D4CD46B31EE">Zahiri, Reza</mods:namePart>
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<mods:nameIdentifier id="B0F401791B751E71B795CDA77F475E24" type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2E371EA93E61E2BC91BD38591ED9D528">Husemann, Martin</mods:namePart>
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<mods:nameIdentifier id="1A4F14F94F9229F0552263F64DE91D5D" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
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<mods:title id="A5370EF654BEF45573D35BB9D4CE934A">Zootaxa</mods:title>
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<mods:date id="6C9B76D97A2696CE99D25E1A9AD69FDC">2023</mods:date>
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<mods:number id="F456978D05B2693D81531AA4C9FE472F">2023-04-18</mods:number>
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<mods:number id="C33BCFDAEBCE2550D6E9290035B9D82D">5267</mods:number>
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<mods:url id="30FF222922289CA635B937386DC23301">http://dx.doi.org/10.11646/zootaxa.5267.1.1</mods:url>
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<mods:classification id="D333C4CB56BA1F12BC1C56699D45808C">journal article</mods:classification>
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<treatment id="03DC6C76CC364C2CF7DFFB69FB48FBC8" ID-DOI="http://doi.org/10.5281/zenodo.7840738" ID-Zenodo-Dep="7840738" LSID="urn:lsid:plazi:treatment:03DC6C76CC364C2CF7DFFB69FB48FBC8" httpUri="http://treatment.plazi.org/id/03DC6C76CC364C2CF7DFFB69FB48FBC8" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<subSubSection id="C36F8EEBCC364C23F7DFFB69FD17FB56" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8BCADD60CC364C23F7DFFB69FE7CFB15" blockId="7.[151,482,1158,1185]" box="[151,482,1158,1185]" pageId="7" pageNumber="8">
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<heading id="D0826A0CCC364C23F7DFFB69FE7CFB15" bold="true" box="[151,482,1158,1185]" fontSize="11" level="2" pageId="7" pageNumber="8" reason="4">
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<taxonomicName id="4C75A6E3CC364C23F7DFFB69FE7CFB15" authority="Strand, 1909" authorityName="Strand" authorityYear="1909" box="[151,482,1158,1185]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B9010172CC364C23F7DFFB69FE7CFB15" bold="true" box="[151,482,1158,1185]" pageId="7" pageNumber="8">
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<emphasis id="B9010172CC364C23F7DFFB69FEA4FB14" bold="true" box="[151,314,1158,1185]" italics="true" pageId="7" pageNumber="8">Metarbelodes</emphasis>
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<bibRefCitation id="EFE4A091CC364C23F609FB69FE7CFB15" author="Strand, E." box="[321,482,1158,1184]" pageId="7" pageNumber="8" pagination="118 - 121" refId="ref83019" refString="Strand, E. (1909) Fam. Metarbelidae (Hollandiidae). In: Lepidoptera aus Deutsch-Ostafrika gesammelt von Herrn Oberleutnant Wintgens. Deutsche Entomologische Zeitschrift " Iris ", 22, 118 - 121." type="journal article" year="1909">Strand, 1909</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BCADD60CC364C23F7DFFB24FD17FB56" blockId="7.[151,1308,1227,1282]" box="[151,649,1227,1251]" pageId="7" pageNumber="8">
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<emphasis id="B9010172CC364C23F7DFFB24FDA9FB56" box="[151,567,1227,1251]" italics="true" pageId="7" pageNumber="8">Deutsche Entomologische Zeitschrift Iris</emphasis>
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<emphasis id="B9010172CC364C23F575FB24FDC8FB56" bold="true" box="[573,598,1227,1251]" pageId="7" pageNumber="8">22</emphasis>
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: 119
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F8EEBCC364C23F7DFFB05FA82FAB7" box="[151,1308,1258,1282]" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="8BCADD60CC364C23F7DFFB05FA82FAB7" blockId="7.[151,1308,1227,1282]" box="[151,1308,1258,1282]" pageId="7" pageNumber="8">
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<typeStatus id="54CE63C2CC364C23F7DFFB05FF57FAB7" box="[151,201,1258,1282]" pageId="7" pageNumber="8">Type</typeStatus>
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species:
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<taxonomicName id="4C75A6E3CC364C23F66EFB05FE0EFAB7" authorityName="Holland" authorityYear="1893" box="[294,400,1258,1282]" class="Insecta" family="Cossidae" genus="Metarbela" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B9010172CC364C23F66EFB05FE0EFAB7" box="[294,400,1258,1282]" italics="true" pageId="7" pageNumber="8">Metarbela</emphasis>
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</taxonomicName>
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?
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<taxonomicName id="4C75A6E3CC364C23F6E9FB05FD25FAB7" authority="Aurivillius, 1901" authorityName="Aurivillius" authorityYear="1901" box="[417,699,1258,1282]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="umtaliana">
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<emphasis id="B9010172CC364C23F6E9FB05FD96FAB7" box="[417,520,1258,1282]" italics="true" pageId="7" pageNumber="8">umtaliana</emphasis>
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<bibRefCitation id="EFE4A091CC364C23F545FB05FD25FAB7" author="Aurivillius, C." box="[525,699,1258,1282]" pageId="7" pageNumber="8" pagination="126 - 128" refId="ref75057" refString="Aurivillius, C. (1901) Diagnosen neuer Lepidopteren aus Afrika. Fam. Hollandiidae. Entomologisk Tidskrift, 22 (2), 126 - 128." type="journal article" year="1901">Aurivillius, 1901</bibRefCitation>
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</taxonomicName>
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(
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<emphasis id="B9010172CC364C23F581FB05FC31FAB7" box="[713,943,1258,1282]" italics="true" pageId="7" pageNumber="8">Entomologisk Tidskrift</emphasis>
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<emphasis id="B9010172CC364C23F4FDFB05FC50FAB7" bold="true" box="[949,974,1258,1282]" pageId="7" pageNumber="8">22</emphasis>
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(2): 127, Fig. 29), by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F8EEBCC364C23F7DFFADFFE8AFA4A" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph id="8BCADD60CC364C23F7DFFADFFE8AFA4A" blockId="7.[151,1437,1328,2044]" pageId="7" pageNumber="8">
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<emphasis id="B9010172CC364C23F7DFFADFFE94FAFF" bold="true" box="[151,266,1328,1354]" pageId="7" pageNumber="8">Diagnosis</emphasis>
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.
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<taxonomicName id="4C75A6E3CC364C23F652FADFFE2AFAFF" authorityName="Strand" authorityYear="1909" box="[282,436,1328,1354]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B9010172CC364C23F652FADFFE2AFAFF" box="[282,436,1328,1354]" italics="true" pageId="7" pageNumber="8">Metarbelodes</emphasis>
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</taxonomicName>
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is considered monotypic at present. The male genitalia can be distinguished from other metarbelids by a long, hollow, well sclerotized, thorn-like process that originates at the costal margin and is bent strongly downwards towards the ventral edge of valva (autapomorphy). The base of the thorn-like process has a broad, long, open slit dorsally, with a weakly-sclerotized projection attached to it. The forewing has a strong, continuous, tubular CuP fold. Based on the latter,
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<taxonomicName id="4C75A6E3CC364C23F585FA2EFCFBFA6F" authorityName="Aurivillius" authorityYear="1901" box="[717,869,1472,1498]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="umtaliana">
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<emphasis id="B9010172CC364C23F585FA2EFCFBFA6F" box="[717,869,1472,1498]" italics="true" pageId="7" pageNumber="8">M. umtaliana</emphasis>
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</taxonomicName>
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is likely more closely related to
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<taxonomicName id="4C75A6E3CC364C23F392FA2FFAD9FA6F" authorityName="Lehmann & Zahiri & Husemann" authorityYear="2023" box="[1242,1351,1472,1498]" class="Insecta" family="Metarbelidae" genus="Zambezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B9010172CC364C23F392FA2FFAD9FA6F" box="[1242,1351,1472,1498]" italics="true" pageId="7" pageNumber="8">Zambezia</emphasis>
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</taxonomicName>
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than to
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<taxonomicName id="4C75A6E3CC364C23F7DFFA0BFE93FA4B" authorityName="Lehmann & Zahiri & Husemann" authorityYear="2023" box="[151,269,1508,1534]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B9010172CC364C23F7DFFA0BFE93FA4B" box="[151,269,1508,1534]" italics="true" pageId="7" pageNumber="8">Lukeniana</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F8EEBCC364C2CF78FF9E7FB48FBC8" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
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<paragraph id="8BCADD60CC364C23F78FF9E7FAE0F907" blockId="7.[151,1437,1328,2044]" pageId="7" pageNumber="8">
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<emphasis id="B9010172CC364C23F78FF9E7FECFF997" bold="true" box="[199,337,1544,1570]" pageId="7" pageNumber="8">Description</emphasis>
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.
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<emphasis id="B9010172CC364C23F617F9E7FE02F997" box="[351,412,1544,1570]" italics="true" pageId="7" pageNumber="8">Head</emphasis>
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: Rough-scaled, with light ochre hair-like scales below compound eyes on fronto-clypeus in male; a pair of pits absent from lower fronto-clypeus, a pair of conical projections present on lower fronto-clypeus; pits behind labial palpi ovoid; labial palpi long, almost as long as eye diameter, consisting of three segments, 2
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<superScript id="7C007028CC364C23F2C4F9BFFA02F9EB" attach="none" box="[1420,1436,1616,1630]" fontSize="6" pageId="7" pageNumber="8">nd</superScript>
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segment longest, 1.5–2.3 longer than the basal segment, 3
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<superScript id="7C007028CC364C23F46AF99BFCAEF937" attach="left" box="[802,816,1652,1666]" fontSize="6" pageId="7" pageNumber="8">rd</superScript>
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segment shortest, 0.5 length of basal segment; antenna bipectinate in male, branches narrow, 3.5 width of shaft, densely scaled cream; flagellum scaled, cream colored.
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</paragraph>
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<paragraph id="8BCADD60CC364C2CF78FF953FDE6FDAC" blockId="7.[151,1437,1328,2044]" lastBlockId="8.[151,1437,150,1149]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<emphasis id="B9010172CC364C23F78FF953FE86F963" box="[199,280,1724,1750]" italics="true" pageId="7" pageNumber="8">Thorax</emphasis>
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: Densely covered with light ochre hair-like scales on patagia and tegulae, slightly glossy, tegulae not pronounced; crest on metathorax small, light ochre. Forelegs and midlegs ochre with long dense hair-like scales. Epiphyses absent. Hindlegs with one pair of narrow tibial spurs, slightly unequal in length (outer spur
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<quantity id="4C8D7085CC364C23F251F8EBFAEFF8AA" box="[1305,1393,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="8" unit="mm" value="1.4">1.4 mm</quantity>
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, inner spur
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<quantity id="4C8D7085CC364C23F7B3F8C7FECBF8F7" box="[251,341,1832,1858]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="8" unit="mm" value="1.3">1.3 mm</quantity>
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); pretarsus with a pair of pulvilli. Forewing upperside with ground colour dark ochre mixed with cream, not glossy, with dark brown, sometimes almost black (= sepia) vertical striae along costa and on the outer half of the forewing upper side; a geometric design is absent (a geometric design was defined by
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<bibRefCitation id="EFE4A091CC364C23F396F89FFA06F83F" author="Lehmann, I." box="[1246,1432,1904,1930]" pageId="7" pageNumber="8" pagination="3 - 81" refId="ref79498" refString="Lehmann, I. (2010 b) s. n. In: A revision of the genus Arbelodes Karsch (Lepidoptera: Cossoidea: Metarbelidae) from southeastcentral and southern Africa with the description of thirteen new species. Published by the author, Hamburg & Wismar, pp. 3 - 81, 8 b / w pls., 5 colour pls. Available from http: // www. biodiversitylibrary. org / bibliography / 79419 (accessed 30 March 2021)" type="journal article" year="2010">Lehmann 2010b</bibRefCitation>
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, e.g., the occurrence of any triangular-like or leaf-like patterns); a simple, not contrasting pattern is present and comprises a straight sepia terminal line, not parallel to termen, from near apex to dorsum, strongly bent towards base of wing at M
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<subScript id="17F1DF25CC364C23F662F801FEACF849" attach="left" box="[298,306,2030,2044]" fontSize="6" pageId="7" pageNumber="8">3</subScript>
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; 1A+2A and discocellular vein sepia; CuA
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<subScript id="17F1DF25CC364C23F450F801FCBEF849" attach="left" box="[792,800,2030,2044]" fontSize="6" pageId="7" pageNumber="8">2</subScript>
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is distinctly marked white or cream, narrow edged sepia above. Hindwing cream-coloured, glossy. Fringe long, cilia up to
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<quantity id="4C8D7085CC394C2CF43FFF79FC51FF04" box="[887,975,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="mm" value="1.5">1.5 mm</quantity>
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, cream-coloured, glossy. Forewing venation (
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<figureCitation id="134EC1E5CC394C2CF79AFF54FEBAFF60" box="[210,292,187,213]" captionStart="FIGURE 1" captionStartId="9.[152,255,1793,1817]" captionTargetBox="[152,1436,777,1769]" captionTargetId="figure-310@9.[152,1436,777,1769]" captionTargetPageId="9" captionText="FIGURE 1. a. Zambezia jennyhuntae, holotype, male, Zimbabwe, Manicaland Province, Mutare (formerly Umtali); b. Z. durrellbarnesi, holotype, female, Zimbabwe, Manicaland Province, Bvumba Mountains; c. Z. darrelplowesi, holotype, female, Zimbabwe, Manicaland Province, Mutare (formerly Umtali), Hospital (today Kopje House); d. Metarbelodes umtaliana, holotype, male, Zimbabwe, Manicaland Province, Old Umtali or Umtali (today Mutare)." figureDoi="http://doi.org/10.5281/zenodo.7840788" httpUri="https://zenodo.org/record/7840788/files/figure.png" pageId="8" pageNumber="9">Fig. 1d</figureCitation>
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;
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<figureCitation id="134EC1E5CC394C2CF678FF54FEE5FF60" box="[304,379,187,213]" captionStart="FIGURE 9" captionStartId="59.[151,250,1714,1738]" captionTargetBox="[235,1352,188,1685]" captionTargetId="figure-19@59.[223,1364,181,1690]" captionTargetPageId="59" captionText="FIGURE 9. Wing venation of: a. Zambezia darrelplowesi, female, holotype (Zambezia = subgroup G1 in Lehmann 2019b); b. Lukeniana obliqualinea, female, Kenya, County Nairobi, Ngong (Lukeniana = subgroup G3 in Lehmann 2019b); c. Zambezia madambae, male, holotype; d. Z. diredaouaensis, male, paratype; e. Lukeniana tubiraensis, male, holotype; f. Metarbelodes umtaliana, male, holotype (Metarbelodes = subgroup G2 in Lehmann 2019b). The areole (blue arrows) is a possible plesiomorphy (in Lehmann 2019b, p. 24), the continuous CuP fold (red arrows), partly sclerotized, is a very rare apomorphy (in Lehmann 2019b, p. 25) among species of Metarbelidae. It has been found only in species close to or to the South of the Zambezi River. In the same areas an intact CuP on the forewing occurs only in basal species (as defined by Lehmann 2019b, pp. 24 and 25) of Metarbelidae (Lehmann 2019b). The complete loss of the CuP in the forewing is another apomorphy (Lehmann 2019b, p. 322) that occurs in the majority of species of Metarbelidae in theAfrotropical Region and Oriental Region (seeFigure 39).Drawings by I.L." figureDoi="http://doi.org/10.5281/zenodo.7840807" httpUri="https://zenodo.org/record/7840807/files/figure.png" pageId="8" pageNumber="9">Fig. 9f</figureCitation>
|
||
) with 1A+2A long forked at base; CuP a strong, tube-like, continuous unsclerotized fold; basal part of CuA
|
||
<subScript id="17F1DF25CC394C2CF656FF1EFEB8FF4A" attach="left" box="[286,294,241,255]" fontSize="6" pageId="8" pageNumber="9">2</subScript>
|
||
originating from two-thirds of lower median; CuA
|
||
<subScript id="17F1DF25CC394C2CF42DFF1EFCF3FF4A" attach="left" box="[869,877,241,255]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
|
||
originating from hind margin of posterior cell
|
||
<subScript id="17F1DF25CC394C2CF23BFF1EFAE9FF4A" attach="left" box="[1395,1399,241,255]" fontSize="6" pageId="8" pageNumber="9">,</subScript>
|
||
M
|
||
<subScript id="17F1DF25CC394C2CF2DCFF1EFA02FF4A" attach="left" box="[1428,1436,241,255]" fontSize="6" pageId="8" pageNumber="9">3</subScript>
|
||
and M
|
||
<subScript id="17F1DF25CC394C2CF797FEFAFF79FE96" attach="left" box="[223,231,277,291]" fontSize="6" pageId="8" pageNumber="9">2</subScript>
|
||
originating from apical angle of posterior cell, both are separated at base; M
|
||
<subScript id="17F1DF25CC394C2CF37EFEFAFBA0FE96" attach="left" box="[1078,1086,277,291]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
|
||
originating from anterior angle of median cell and close to base of a relatively large areole; R
|
||
<subScript id="17F1DF25CC394C2CF403FED6FCCDFEF2" attach="left" box="[843,851,313,327]" fontSize="6" pageId="8" pageNumber="9">5</subScript>
|
||
originates from posterior angle of areole; R
|
||
<subScript id="17F1DF25CC394C2CF277FED6FAD9FEF2" attach="left" box="[1343,1351,313,327]" fontSize="6" pageId="8" pageNumber="9">4</subScript>
|
||
+R
|
||
<subScript id="17F1DF25CC394C2CF221FED6FAEFFEF2" attach="left" box="[1385,1393,313,327]" fontSize="6" pageId="8" pageNumber="9">3</subScript>
|
||
+R
|
||
<subScript id="17F1DF25CC394C2CF2DCFED6FA02FEF2" attach="left" box="[1428,1436,313,327]" fontSize="6" pageId="8" pageNumber="9">2</subScript>
|
||
on a long stalk that is originating from anterior angle of areole close to base of R
|
||
<subScript id="17F1DF25CC394C2CF354FEB2FBBAFEDE" attach="left" box="[1052,1060,349,363]" fontSize="6" pageId="8" pageNumber="9">5</subScript>
|
||
; R
|
||
<subScript id="17F1DF25CC394C2CF30DFEB2FBD3FEDE" attach="left" box="[1093,1101,349,363]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
|
||
originating from anterior part of median cell; Sc more or less parallel to R
|
||
<subScript id="17F1DF25CC394C2CF5CDFE6EFD13FE3A" attach="left" box="[645,653,385,399]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
|
||
. Hindwing—3A present; 1A+2A rudimentary; CuP present but weak; CuA
|
||
<subScript id="17F1DF25CC394C2CF784FE4AFF4AFE06" attach="left" box="[204,212,421,435]" fontSize="6" pageId="8" pageNumber="9">2</subScript>
|
||
originating from two-thirds of lower median of posterior cell; CuA
|
||
<subScript id="17F1DF25CC394C2CF48BFE4AFC55FE06" attach="left" box="[963,971,421,435]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
|
||
originating from hind margin of posterior cell; M
|
||
<subScript id="17F1DF25CC394C2CF7A0FE26FF6EFE62" attach="left" box="[232,240,457,471]" fontSize="6" pageId="8" pageNumber="9">3</subScript>
|
||
and M
|
||
<subScript id="17F1DF25CC394C2CF60AFE26FED4FE62" attach="left" box="[322,330,457,471]" fontSize="6" pageId="8" pageNumber="9">2</subScript>
|
||
originating from apical angle of posterior cell, separated; M
|
||
<subScript id="17F1DF25CC394C2CF4BFFE26FC61FE62" attach="left" box="[1015,1023,457,471]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
|
||
+Rs originating from apical angle of anterior cell, long-stalked; a short bar present from Rs to Sc+R
|
||
<subScript id="17F1DF25CC394C2CF404FE02FCCAFE4E" attach="left" box="[844,852,493,507]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
|
||
; a discocellular vein on both forewing and hindwing present. Retinaculum and frenulum absent.
|
||
</paragraph>
|
||
<paragraph id="8BCADD60CC394C2CF78FFDCCFA03FBEC" blockId="8.[151,1437,150,1149]" pageId="8" pageNumber="9">
|
||
<emphasis id="B9010172CC394C2CF78FFDCCFEAFFD88" box="[199,305,547,573]" italics="true" pageId="8" pageNumber="9">Abdomen</emphasis>
|
||
: Externally primarily covered by cream and light ochre, hair-like scales; abdominal tuft cream, short, up to 0.25 length of abdomen. Male genitalia (
|
||
<figureCitation id="134EC1E5CC394C2CF5DEFDA8FD6FFDD4" box="[662,753,583,609]" captionStart="FIGURE 11" captionStartId="62.[151,250,1774,1798]" captionTargetBox="[162,1436,606,1750]" captionTargetId="figure-205@62.[151,1436,597,1750]" captionTargetPageId="62" captionText="FIGURE 11. a. Zambezia madambae, holotype, male, Botswana, Chobe District Council, Kasane, Chobe River, Chobe Rapids; b. Z. jennyhuntae, holotype, male, Zimbabwe, Manicaland Province, Mutare (formerly Umtali); c. Metarbelodes umtaliana, holotype, male, Zimbabwe, Manicaland Province, Old Umtali or Umtali (today Mutare); d1. Lukeniana mzuzuensis, paratype, female, Malawi, Northern Region, Mzuzu, Nkho-rongo, female genitalia with ductus bursae and corpus bursae, both thinly membranous and without any structures; d2. Detail of female genitalia of L. mzuzuensis, paratype, with narrow bandson lateroventral surface of segment 8 (cf. red arrows); e. L. obliqualinea, holotype, male, Kenya, County Nairobi, Nairobi (picture taken by Kim Buckmaster, NHMUK, London (picture taken in 1996).." figureDoi="http://doi.org/10.5281/zenodo.7840814" httpUri="https://zenodo.org/record/7840814/files/figure.png" pageId="8" pageNumber="9">Fig. 11c</figureCitation>
|
||
) with tegumen fused to vinculum, forming a firm ring. Uncus large, ovoid, comprised of a pair of narrowly rounded lobes, short setae ventrally, basal edge of uncus is not straight but has a crescent-shape in horizontal position at middle. Gnathos short, well above costa, covered with short toothlike processes at distal end, hand-shaped, not connected by a sclerotized band basally. Valva ovoid with broad base, costa lacking setae, inner side of median sector with few setae; large, setose, weakly-sclerotized projection attached to thorn-like process from near base of valva and along costal margin; thorn-like process prominent and hollow; at end of its base with long open slit dorsally where it is connected by a thin membrane to weakly-sclerotized projection; thorn-like process originating from outer side of valva near costa, and strongly bent downwards towards ventral edge of valva. Sacculus strongly reduced to one-third near base of valva. A short emargination (less than 25% length of valva) extending between weakly-sclerotized projection and thorn-like process. Transtilla absent; semi-transtilla short, rectangular with a somewhat leaf-shaped, attached to costa of valva and opposite of vinculum, without setae. Juxta nearly twice as large as saccus with two acuminate tips and a short process at each tip, deeply emargination between tips (90% the length of juxta). Vinculum broad at saccus, twice as large as the saccus, forming a plate-like structure. Saccus digitate, gently rounded caudally. Phallus simple, 1.5 longer than basal width of valva, slightly trumpet-shaped, bilobate, with a cleft at each end. Female postabdominal structure and genitalia unknown.
|
||
</paragraph>
|
||
<paragraph id="8BCADD60CC394C2CF78FFB8DFB48FBC8" blockId="8.[151,1437,150,1149]" box="[199,1238,1122,1149]" pageId="8" pageNumber="9">
|
||
<emphasis id="B9010172CC394C2CF78FFB8DFE19FBC9" bold="true" box="[199,391,1122,1148]" pageId="8" pageNumber="9">Species richness</emphasis>
|
||
. As defined herein, the genus includes only the
|
||
<typeStatus id="54CE63C2CC394C2CF4EAFB8CFC4CFBC8" box="[930,978,1123,1149]" pageId="8" pageNumber="9">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C75A6E3CC394C2CF370FB8CFB4EFBC8" authorityName="Aurivillius" authorityYear="1901" box="[1080,1232,1123,1149]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="umtaliana">
|
||
<emphasis id="B9010172CC394C2CF370FB8CFB4EFBC8" box="[1080,1232,1123,1149]" italics="true" pageId="8" pageNumber="9">M. umtaliana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |