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<document id="0916218E38459B669CBFB90E9F8E6E53" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza &amp; Husemann, Martin" docDate="2023" docId="03DC6C76CC154C02F7DFFA01FC66F998" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lukeniana stueningi Lehmann, Zahiri &amp; Husemann 2023, sp. nov." docType="treatment" docUuid="F99BEB0F-3BDF-48D9-9086-FF69285E4811" docUuidSource="ZooBank" docVersion="3" lastPageNumber="39" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="37" updateTime="1699487052491" updateUser="plazi">
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<mods:title id="308BFD39D5528BC66CFC0CB86401F3F3">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
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<mods:namePart id="28A9A43526B2B8819E56CFDAE08B9FAF">Lehmann, Ingo</mods:namePart>
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<mods:namePart id="BD30627FB84EC24D9D10423C8DBB7EB9">Zahiri, Reza</mods:namePart>
<mods:nameIdentifier id="575CCD69981FD755B858C8D47CD83975" type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D8D1F3117019013F691291CEB5EB3082">Husemann, Martin</mods:namePart>
<mods:nameIdentifier id="4C3768A3853F55F011B6B08846AE7EBE" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
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<mods:date id="F7A59B0A5BF7A8B09EA1942C4F397AA3">2023</mods:date>
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<heading id="D0826A0CCC154C00F7DFFA01FC1BF9BC" bold="true" box="[151,901,1518,1545]" fontSize="11" level="2" pageId="36" pageNumber="37" reason="4">
<emphasis id="B9010172CC154C00F7DFFA01FC1BF9BC" bold="true" box="[151,901,1518,1545]" pageId="36" pageNumber="37">
<taxonomicName id="4C75A6E3CC154C00F7DFFA01FCBEF9BC" ID-CoL="B8R9P" authority="Lehmann, Zahiri &amp; Husemann" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[151,800,1518,1545]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="stueningi" status="sp. nov.">
<emphasis id="B9010172CC154C00F7DFFA01FE0DF9BC" bold="true" box="[151,403,1518,1545]" italics="true" pageId="36" pageNumber="37">Lukeniana stueningi</emphasis>
Lehmann, Zahiri &amp; Husemann
</taxonomicName>
<taxonomicNameLabel id="A232BC09CC154C00F46FFA00FC1BF9BC" box="[807,901,1519,1545]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BCADD60CC154C00F7DFF9FCFE80F998" blockId="36.[151,956,1518,1617]" box="[151,286,1555,1581]" pageId="36" pageNumber="37">
<figureCitation id="134EC1E5CC154C00F7DFF9FCFF73F998" box="[151,237,1555,1581]" captionStart="FIGURE 6" captionStartId="37.[151,250,1791,1815]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-19@37.[151,1436,181,1767]" captionTargetPageId="37" captionText="FIGURE 6. a. Lukeniana kammeri, holotype, male, Kenya, County Kiambu, Thika; b. L. mbalaensis, paratype, female, Zambia, Northern Province, Mbala; c. L. mbalaensis, holotype, male, Zambia, Northern Province, Mbala; d. L. hausmanni, paratype, male, Zambia, Northern Province, 23 km southeast Mbala; e. L. lenzi, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area; f. L. robplowesi, holotype, male, Zambia, Northern Province, Mbala; g. L. utaheidenreichae, holotype, male, Mozambique, Manica Province, Barr De Chikamba Real; h. L. stueningi, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest." figureDoi="http://doi.org/10.5281/zenodo.7840800" httpUri="https://zenodo.org/record/7840800/files/figure.png" pageId="36" pageNumber="37">Figs 6h</figureCitation>
,
<figureCitation id="134EC1E5CC154C00F7B0F9FCFE80F998" box="[248,286,1555,1581]" captionStart="FIGURE 12" captionStartId="63.[151,250,1330,1354]" captionTargetBox="[160,1427,191,1293]" captionTargetId="figure-19@63.[151,1436,181,1306]" captionTargetPageId="63" captionText="FIGURE 12. a. Lukeniana mzuzuensis, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo; b. L. mbalaensis, holotype, male, Zambia, Northern Province, Mbala; c. L. robplowesi, holotype, male, Zambia, Northern Province, Mbala; d. L. hausmanni, paratype, male, Zambia, Northern Province, 23 km southeast Mbala; e. L. lenzi, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area; f. L. stueningi, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest." figureDoi="http://doi.org/10.5281/zenodo.7840816" httpUri="https://zenodo.org/record/7840816/files/figure.png" pageId="36" pageNumber="37">12f</figureCitation>
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urn:lsid:zoobank.org:act:
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<materialsCitation id="3B1DD73DCC154C00F7DFF990FD2AF908" collectionCode="LIB, ZFMK" county="Analysis of Biodiversity Change" location="Zamibia" municipality="Leibniz-Institute" pageId="36" pageNumber="37" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE63C2CC154C00F7DFF990FF51F92C" box="[151,207,1663,1689]" pageId="36" pageNumber="37">Type</typeStatus>
locality and repository:
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,
<location id="8EAA8BBBCC154C00F51CF990FB1EF92C" LSID="urn:lsid:plazi:treatment:03DC6C76CC154C02F7DFFA01FC66F998:8EAA8BBBCC154C00F51CF990FB1EF92C" box="[596,1152,1663,1689]" county="Analysis of Biodiversity Change" municipality="Leibniz-Institute" name="Zoological Research Museum Alexander Koenig" pageId="36" pageNumber="37">Zoological Research Museum Alexander Koenig</location>
,
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for the
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(
<collectionCode id="ED6445A5CC154C00F55CF94CFDDDF908" box="[532,579,1699,1725]" country="Liberia" lsid="urn:lsid:biocol.org:col:14548" name="University of Liberia" pageId="36" pageNumber="37" type="Herbarium">LIB</collectionCode>
/
<collectionCode id="ED6445A5CC154C00F511F94CFD32F908" box="[601,684,1699,1725]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="36" pageNumber="37" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="C36F8EEBCC154C00F7DFF905FB80F8F8" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BCADD60CC154C00F7DFF905FB80F8F8" blockId="36.[151,1436,1770,2013]" pageId="36" pageNumber="37">
<emphasis id="B9010172CC154C00F7DFF905FE1CF8B1" bold="true" box="[151,386,1770,1796]" pageId="36" pageNumber="37">Material examined.</emphasis>
<materialsCitation id="3B1DD73DCC154C00F6C3F904FB80F8F8" collectingDate="2012-10-18" collectingDateMax="2012-10-20" collectingDateMin="2012-10-18" collectionCode="R, ZFMK" collectorName="J. Murphy &amp; I. Lehmann" country="Zambia" county="Isoka District" elevation="2080" location="Nyika National Park" municipality="Nyika Plateau" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Muchinga" typeStatus="holotype">
<typeStatus id="54CE63C2CC154C00F6C3F904FE6DF8B0" box="[395,499,1771,1797]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F3629DF0CC154C00F50BF904FD05F8B0" box="[579,667,1771,1797]" name="Zambia" pageId="36" pageNumber="37">Zambia</collectingCountry>
,
<collectingRegion id="49B11382CC154C00F5E0F904FC16F8B0" box="[680,904,1771,1797]" country="Zambia" name="Muchinga" pageId="36" pageNumber="37">Muchinga Province</collectingRegion>
,
<collectingCounty id="62ABA5ECCC154C00F4DEF904FBACF8B0" box="[918,1074,1771,1797]" pageId="36" pageNumber="37">Isoka District</collectingCounty>
,
<collectingMunicipality id="6BAE471ACC154C00F377F904FB7FF8B0" box="[1087,1249,1771,1797]" pageId="36" pageNumber="37">Nyika Plateau</collectingMunicipality>
,
<location id="8EAA8BBBCC154C00F3A7F904FF53F89C" LSID="urn:lsid:plazi:treatment:03DC6C76CC154C02F7DFFA01FC66F998:8EAA8BBBCC154C00F3A7F904FF53F89C" country="Zambia" county="Isoka District" municipality="Nyika Plateau" name="Nyika National Park" pageId="36" pageNumber="37" stateProvince="Muchinga">Nyika National Park</location>
,
<location id="8EAA8BBBCC154C00F794F8E0FE2CF89C" LSID="urn:lsid:plazi:treatment:03DC6C76CC154C02F7DFFA01FC66F998:8EAA8BBBCC154C00F794F8E0FE2CF89C" box="[220,434,1807,1833]" country="Zambia" county="Isoka District" municipality="Nyika Plateau" name="Manyanjere Forest" pageId="36" pageNumber="37" stateProvince="Muchinga">Manyanjere Forest</location>
(=misspelling of
<location id="8EAA8BBBCC154C00F5CBF8E0FCC3F89C" LSID="urn:lsid:plazi:treatment:03DC6C76CC154C02F7DFFA01FC66F998:8EAA8BBBCC154C00F5CBF8E0FCC3F89C" box="[643,861,1807,1833]" country="Zambia" county="Isoka District" municipality="Nyika Plateau" name="Manyenjere Forest" pageId="36" pageNumber="37" stateProvince="Muchinga">Manyenjere Forest</location>
),
<quantity id="4C8D7085CC154C00F43CF8E0FC48F89C" box="[884,982,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.08" pageId="36" pageNumber="37" unit="m" value="2080.0">
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, 1035′24 S, 3339′40 E,
<date id="FFCBFBA0CC154C00F3BAF8E0FF4CF8F8" pageId="36" pageNumber="37" value="2012-10-18" valueMax="2012-10-20" valueMin="2012-10-18">
<collectingDate id="EF8F0248CC154C00F3BAF8E0FF4CF8F8" pageId="36" pageNumber="37" value="2012-10-18" valueMax="2012-10-20" valueMin="2012-10-18">18–20 October 2012</collectingDate>
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,
<collectionCode id="ED6445A5CC154C00F795F8DCFF6DF8F8" box="[221,243,1843,1869]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="36" pageNumber="37">R</collectionCode>
.
<collectorName id="2680B8B6CC154C00F7B6F8DCFEEFF8F8" box="[254,369,1843,1869]" pageId="36" pageNumber="37">J. Murphy</collectorName>
leg., genitalia slide number 29/082015
<collectorName id="2680B8B6CC154C00F466F8DCFC31F8F8" box="[814,943,1843,1869]" pageId="36" pageNumber="37">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC154C00F4F6F8DCFB8CF8F8" box="[958,1042,1843,1869]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="36" pageNumber="37" type="Museum">ZFMK</collectionCode>
).
</materialsCitation>
</paragraph>
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<subSubSection id="C36F8EEBCC154C02F78FF8B9FE3CFC8C" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37" type="description">
<paragraph id="8BCADD60CC154C00F78FF8B9FACBF868" blockId="36.[151,1436,1770,2013]" pageId="36" pageNumber="37">
<emphasis id="B9010172CC154C00F78FF8B9FEC7F8C5" bold="true" box="[199,345,1878,1904]" pageId="36" pageNumber="37">Description.</emphasis>
Male.
<emphasis id="B9010172CC154C00F6E0F8B8FE7BF8C4" box="[424,485,1879,1905]" italics="true" pageId="36" pageNumber="37">Head</emphasis>
: Yellow-ochre mixed with sepia hair-like scales towards eyes; long, dense, hair-like, dark ochre scales between eyes; eyes vinaceous with black patches; antenna 0.50 length of forewing, bipectinate, with branches 5.0 width of shaft, covered with cream-coloured scales laterally, shaft covered with cream-coloured scales dorsally; antennal tips not spatulate and with two long scales, bending towards apex; labial palpi dark ochre.
</paragraph>
<caption id="DF0A8DE8CC144C01F7DFF910FD3FF87E" ID-DOI="http://doi.org/10.5281/zenodo.7840800" ID-Zenodo-Dep="7840800" httpUri="https://zenodo.org/record/7840800/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,1791,1815]" targetBox="[151,1436,181,1767]" targetPageId="37">
<paragraph id="8BCADD60CC144C01F7DFF910FD3FF87E" blockId="37.[151,1437,1791,1995]" pageId="37" pageNumber="38">
<emphasis id="B9010172CC144C01F7DFF910FD89F8A2" bold="true" box="[151,535,1791,1815]" pageId="37" pageNumber="38">
FIGURE 6. a.
<taxonomicName id="4C75A6E3CC144C01F676F910FD89F8A2" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[318,535,1791,1815]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC144C01F676F910FD89F8A2" bold="true" box="[318,535,1791,1815]" italics="true" pageId="37" pageNumber="38">Lukeniana kammeri</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Kenya, County Kiambu, Thika;
<emphasis id="B9010172CC144C01F36EF910FB47F8A2" bold="true" box="[1062,1241,1791,1815]" pageId="37" pageNumber="38">
b.
<taxonomicName id="4C75A6E3CC144C01F30CF910FB47F8A2" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1092,1241,1791,1815]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mbalaensis">
<emphasis id="B9010172CC144C01F30CF910FB47F8A2" bold="true" box="[1092,1241,1791,1815]" italics="true" pageId="37" pageNumber="38">L. mbalaensis</emphasis>
</taxonomicName>
</emphasis>
, paratype, female, Zambia, Northern Province, Mbala;
<emphasis id="B9010172CC144C01F546F8CCFD24F88E" bold="true" box="[526,698,1827,1851]" pageId="37" pageNumber="38">
c.
<taxonomicName id="4C75A6E3CC144C01F560F8CCFD24F88E" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[552,698,1827,1851]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mbalaensis">
<emphasis id="B9010172CC144C01F560F8CCFD24F88E" bold="true" box="[552,698,1827,1851]" italics="true" pageId="37" pageNumber="38">L. mbalaensis</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Zambia, Northern Province, Mbala;
<emphasis id="B9010172CC144C01F3AEF8CCFA08F88E" bold="true" box="[1254,1430,1827,1851]" pageId="37" pageNumber="38">
d.
<taxonomicName id="4C75A6E3CC144C01F24AF8CCFA08F88E" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1282,1430,1827,1851]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hausmanni">
<emphasis id="B9010172CC144C01F24AF8CCFA08F88E" bold="true" box="[1282,1430,1827,1851]" italics="true" pageId="37" pageNumber="38">L. hausmanni</emphasis>
</taxonomicName>
</emphasis>
, paratype, male, Zambia, Northern Province, 23 km southeast Mbala;
<emphasis id="B9010172CC144C01F42AF8A8FC54F8EA" bold="true" box="[866,970,1863,1887]" pageId="37" pageNumber="38">
e.
<taxonomicName id="4C75A6E3CC144C01F433F8A8FC54F8EA" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[891,970,1863,1887]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lenzi">
<emphasis id="B9010172CC144C01F433F8A8FC54F8EA" bold="true" box="[891,970,1863,1887]" italics="true" pageId="37" pageNumber="38">L. lenzi</emphasis>
</taxonomicName>
</emphasis>
, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area;
<emphasis id="B9010172CC144C01F682F884FDECF836" bold="true" box="[458,626,1899,1923]" pageId="37" pageNumber="38">
f.
<taxonomicName id="4C75A6E3CC144C01F6AAF884FDECF836" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[482,626,1899,1923]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="robplowesi">
<emphasis id="B9010172CC144C01F6AAF884FDECF836" bold="true" box="[482,626,1899,1923]" italics="true" pageId="37" pageNumber="38">L. robplowesi</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Zambia, Northern Province, Mbala;
<emphasis id="B9010172CC144C01F3EDF884FA08F836" bold="true" box="[1189,1430,1899,1923]" pageId="37" pageNumber="38">
g.
<taxonomicName id="4C75A6E3CC144C01F389F884FA08F836" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[1217,1430,1899,1923]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="utaheidenreichae">
<emphasis id="B9010172CC144C01F389F884FA08F836" bold="true" box="[1217,1430,1899,1923]" italics="true" pageId="37" pageNumber="38">L. utaheidenreichae</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Mozambique, Manica Province, Barr De Chikamba Real;
<emphasis id="B9010172CC144C01F4C7F860FBB6F812" bold="true" box="[911,1064,1935,1959]" pageId="37" pageNumber="38">
h.
<taxonomicName id="4C75A6E3CC144C01F4E2F860FBB6F812" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[938,1064,1935,1959]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC144C01F4E2F860FBB6F812" bold="true" box="[938,1064,1935,1959]" italics="true" pageId="37" pageNumber="38">L. stueningi</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest.
</paragraph>
</caption>
<paragraph id="8BCADD60CC174C02F78FFF78FAC0FE18" blockId="38.[151,1437,151,1581]" pageId="38" pageNumber="39">
<emphasis id="B9010172CC174C02F78FFF78FE86FF04" box="[199,280,151,177]" italics="true" pageId="38" pageNumber="39">Thorax</emphasis>
: Patagia and tegulae not shiny; long hair-like scales of ochre mixed with sepia; a small crest of ivoryyellow scales on metathorax. Hindlegs ochre with fine hair-like scales; one pair of narrow tibial spurs present, outer spur
<emphasis id="B9010172CC174C02F647FF30FEAEFF4D" box="[271,304,223,248]" italics="true" pageId="38" pageNumber="39">ca.</emphasis>
1.0 mm, inner spur
<emphasis id="B9010172CC174C02F55AFF30FDADFF4D" box="[530,563,223,248]" italics="true" pageId="38" pageNumber="39">ca.</emphasis>
<quantity id="4C8D7085CC174C02F572FF30FD0DFF4C" box="[570,659,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="38" pageNumber="39" unit="mm" value="0.6">0.6 mm</quantity>
. Forewing length 15.0 mm (wingspan
<quantity id="4C8D7085CC174C02F30AFF31FB34FF4C" box="[1090,1194,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="38" pageNumber="39" unit="mm" value="32.5">32.5 mm</quantity>
). Forewing upperside ochre; costal margin sepia, without striae; termen without lunules; terminal and subterminal lines absent; faded striae of dark ochre on almost entire wing; CuA
<subScript id="17F1DF25CC174C02F5E1FED6FD2FFEF2" attach="left" box="[681,689,313,327]" fontSize="6" pageId="38" pageNumber="39">2</subScript>
with narrow edge of sepia above; all remaining veins not distinctly coloured; cilia long,
<quantity id="4C8D7085CC174C02F636FEA5FE48FED0" box="[382,470,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="39" unit="mm" value="1.5">1.5 mm</quantity>
, in forewing and hindwing, cream-coloured, shiny. Underside of forewing not roughly scaled and with many fine hair-like scales, cream-coloured, glossy, costal margin dark ochre along costa, without striae. Hindwing upperside cream-coloured and glossy. Underside as in forewing but without darkened costa.
</paragraph>
<paragraph id="8BCADD60CC174C02F78FFE58FB5AFCA0" blockId="38.[151,1437,151,1581]" pageId="38" pageNumber="39">
<emphasis id="B9010172CC174C02F78FFE58FEAFFE64" box="[199,305,439,465]" italics="true" pageId="38" pageNumber="39">Abdomen</emphasis>
: Cream-coloured and ochre, glossy; abdominal tuft short, one-fifth abdominal length. Genitalia (
<figureCitation id="134EC1E5CC174C02F227FE58FF23FE40" captionStart="FIGURE 12" captionStartId="63.[151,250,1330,1354]" captionTargetBox="[160,1427,191,1293]" captionTargetId="figure-19@63.[151,1436,181,1306]" captionTargetPageId="63" captionText="FIGURE 12. a. Lukeniana mzuzuensis, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo; b. L. mbalaensis, holotype, male, Zambia, Northern Province, Mbala; c. L. robplowesi, holotype, male, Zambia, Northern Province, Mbala; d. L. hausmanni, paratype, male, Zambia, Northern Province, 23 km southeast Mbala; e. L. lenzi, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area; f. L. stueningi, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest." figureDoi="http://doi.org/10.5281/zenodo.7840816" httpUri="https://zenodo.org/record/7840816/files/figure.png" pageId="38" pageNumber="39">Fig. 12f</figureCitation>
) with uncus lobes bearing rounded tips, both short and long setae ventrally, basal edge of uncus strongly bent at middle, triangular in ventral view, emargination broad,
<emphasis id="B9010172CC174C02F44FFE10FCB7FDAD" box="[775,809,511,536]" italics="true" pageId="38" pageNumber="39">ca.</emphasis>
3.5 as broad as one tip of uncus lobe; gnathos arms 1.5 as long as basal width of valva; valvae elongate, oval, costa without setae; sacculus without setae; almost 50% of inner side of valva with a weakly-sclerotized projection, setose, with rectangular tip shorter than single thorn-like process below; the latter hollow and with tiny setae; a long, ovoid emargination extending between weakly-sclerotized projection and thorn-like process,
<emphasis id="B9010172CC174C02F514FD60FDE3FD1D" box="[604,637,655,680]" italics="true" pageId="38" pageNumber="39">ca.</emphasis>
65% length of valva; ventral side of valva not bent in middle. Saccus short, triangular, opposite an unsually broad ventral part of vinculum, rectangular,
<emphasis id="B9010172CC174C02F37FFD5CFBC6FD79" box="[1079,1112,691,716]" italics="true" pageId="38" pageNumber="39">ca.</emphasis>
8 larger than saccus. Juxta 5 larger than saccus, between tips an unusually shallow emargination, only 25% of length of juxta. Phallus 1.3 as long as width of valva, not trumpet-like, not bent in middle, slightly bilobed with a cleft at each end.
</paragraph>
<paragraph id="8BCADD60CC174C02F78FFCF0FE3CFC8C" blockId="38.[151,1437,151,1581]" box="[199,418,799,825]" pageId="38" pageNumber="39">
<emphasis id="B9010172CC174C02F78FFCF0FEBBFC8C" bold="true" box="[199,293,799,825]" pageId="38" pageNumber="39">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC174C02F78FFCADFB56FB70" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="8BCADD60CC174C02F78FFCADFB56FB70" blockId="38.[151,1437,151,1581]" pageId="38" pageNumber="39">
<emphasis id="B9010172CC174C02F78FFCADFEDCFCE9" bold="true" box="[199,322,834,860]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
<taxonomicName id="4C75A6E3CC174C02F601FCACFDB3FCE9" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[329,557,835,861]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC174C02F601FCACFDB3FCE9" box="[329,557,835,861]" italics="true" pageId="38" pageNumber="39">Lukeniana stueningi</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C75A6E3CC174C02F5B5FCACFC42FCE8" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[765,988,835,861]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="utaheidenreichae">
<emphasis id="B9010172CC174C02F5B5FCACFC42FCE8" box="[765,988,835,861]" italics="true" pageId="38" pageNumber="39">L. utaheidenreichae</emphasis>
</taxonomicName>
with which it shares at least six characters: (i) the gnathos arms are very broad, as broad as the semi-transtilla; (ii) the ventral side of the end of the gnathos arms, specifically the palmate structures, are covered with scales dorsally and also ventrally; (iii) the valvae are elongate and ovoid; (iv) the weakly-sclerotized projection is shorter than the thorn-like process below and covers less than 30% of the inner side of valva; (v) an unsually broad ventral part of vinculum is rectangular; and (vi) the sacculus is only half of the length of ventral edge of valva. Diagnostic characters for
<taxonomicName id="4C75A6E3CC174C02F312FC18FB41FBA5" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1114,1247,1015,1040]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC174C02F312FC18FB41FBA5" box="[1114,1247,1015,1040]" italics="true" pageId="38" pageNumber="39">L. stueningi</emphasis>
</taxonomicName>
include the long emargination of the valva, 40% of the length of valva in
<taxonomicName id="4C75A6E3CC174C02F45BFBF4FC6CFB80" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[787,1010,1051,1077]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="utaheidenreichae">
<emphasis id="B9010172CC174C02F45BFBF4FC6CFB80" box="[787,1010,1051,1077]" italics="true" pageId="38" pageNumber="39">L. utaheidenreichae</emphasis>
</taxonomicName>
versus 65% in
<taxonomicName id="4C75A6E3CC174C02F3E8FBF4FABAFB81" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1184,1316,1051,1076]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC174C02F3E8FBF4FABAFB81" box="[1184,1316,1051,1076]" italics="true" pageId="38" pageNumber="39">L. stueningi</emphasis>
</taxonomicName>
; the broad ventral part of the vinculum, which is slightly shorter in length than the basal width of one valva in
<taxonomicName id="4C75A6E3CC174C02F247FBD0FA08FBED" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1295,1430,1087,1112]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC174C02F247FBD0FA08FBED" box="[1295,1430,1087,1112]" italics="true" pageId="38" pageNumber="39">L. stueningi</emphasis>
</taxonomicName>
, but is 1.4 as long as the basal width of one valva in
<taxonomicName id="4C75A6E3CC174C02F5A4FB8CFC52FBC8" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[748,972,1123,1149]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="utaheidenreichae">
<emphasis id="B9010172CC174C02F5A4FB8CFC52FBC8" box="[748,972,1123,1149]" italics="true" pageId="38" pageNumber="39">L. utaheidenreichae</emphasis>
</taxonomicName>
; the narrowly finger-shaped saccus in
<taxonomicName id="4C75A6E3CC174C02F2CEFB8CFEC6FB14" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="utaheidenreichae">
<emphasis id="B9010172CC174C02F2CEFB8CFEC6FB14" italics="true" pageId="38" pageNumber="39">L. utaheidenreichae</emphasis>
</taxonomicName>
that is triangular in
<taxonomicName id="4C75A6E3CC174C02F577FB68FD5AFB15" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[575,708,1159,1184]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC174C02F577FB68FD5AFB15" box="[575,708,1159,1184]" italics="true" pageId="38" pageNumber="39">L. stueningi</emphasis>
</taxonomicName>
; and the basal edge of the uncus that is rounded and less bent at the middle in
<taxonomicName id="4C75A6E3CC174C02F67AFB44FD8FFB70" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[306,529,1195,1221]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="utaheidenreichae">
<emphasis id="B9010172CC174C02F67AFB44FD8FFB70" box="[306,529,1195,1221]" italics="true" pageId="38" pageNumber="39">L. utaheidenreichae</emphasis>
</taxonomicName>
, but strongly bent at the middle and triangular in
<taxonomicName id="4C75A6E3CC174C02F375FB44FB5FFB71" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1085,1217,1195,1220]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC174C02F375FB44FB5FFB71" box="[1085,1217,1195,1220]" italics="true" pageId="38" pageNumber="39">L. stueningi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC174C02F78FFB21FE77FA74" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BCADD60CC174C02F78FFB21FDB5FA28" blockId="38.[151,1437,151,1581]" pageId="38" pageNumber="39">
<emphasis id="B9010172CC174C02F78FFB21FEFFFB5D" bold="true" box="[199,353,1230,1256]" pageId="38" pageNumber="39">Distribution.</emphasis>
<taxonomicName id="4C75A6E3CC174C02F620FB20FDD3FB5D" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[360,589,1231,1257]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC174C02F620FB20FDD3FB5D" box="[360,589,1231,1257]" italics="true" pageId="38" pageNumber="39">Lukeniana stueningi</emphasis>
</taxonomicName>
is known only from Manyenjere Forest (elevation
<quantity id="4C8D7085CC174C02F3C5FB20FAA7FB5C" box="[1165,1337,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.9945" metricValueMax="2.099" metricValueMin="1.89" pageId="38" pageNumber="39" unit="m" value="1994.5" valueMax="2099.0" valueMin="1890.0">
<geoCoordinate id="EE41BBA7CC174C02F3C5FB20FB55FB5C" box="[1165,1227,1231,1257]" degrees="1,890" direction="north" orientation="latitude" pageId="38" pageNumber="39" precision="55" value="1.89">1,890</geoCoordinate>
<geoCoordinate id="EE41BBA7CC174C02F383FB20FA87FB5C" box="[1227,1305,1231,1257]" degrees="2,099" direction="west" orientation="longitude" pageId="38" pageNumber="39" precision="55" value="-2.099">‒2,099</geoCoordinate>
m
</quantity>
, average annual rainfall
<emphasis id="B9010172CC174C02F60EFB1CFEF6FAB9" box="[326,360,1267,1292]" italics="true" pageId="38" pageNumber="39">ca.</emphasis>
<quantity id="4C8D7085CC174C02F63AFB1CFDB4FAB8" box="[370,554,1267,1293]" metricMagnitude="0" metricUnit="m" metricValue="1.285" metricValueMax="1.3" metricValueMin="1.27" pageId="38" pageNumber="39" unit="mm" value="1285.0" valueMax="1300.0" valueMin="1270.0">1270‒1300 mm</quantity>
), located on the Zambian side of the Nyika Plateau that borders the
<collectingCountry id="F3629DF0CC174C02F20FFB1CFA03FAB8" box="[1351,1437,1267,1293]" name="Malawi" pageId="38" pageNumber="39">Malawi</collectingCountry>
Rift,
<emphasis id="B9010172CC174C02F799FAF8FF6DFA85" box="[209,243,1303,1328]" italics="true" pageId="38" pageNumber="39">ca.</emphasis>
<quantity id="4C8D7085CC174C02F7B2FAF9FEDFFA84" box="[250,321,1302,1329]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="38" pageNumber="39" unit="km" value="45.0">45 km</quantity>
east from Manyenjere Forest, and extends westward almost to the eastern edge of the Luangwa rift graben (
<bibRefCitation id="EFE4A091CC174C02F7BBFAD4FE4AFAE0" author="Roberts, E. M. &amp; Stevens, N. J. &amp; O'Connor, P. M. &amp; Dirks, P. H. G. M. &amp; Gottfried, M. D. &amp; Clyde, W. C. &amp; Armstrong, R. A. &amp; Kemp, A. I. S. &amp; Hemming, S." box="[243,468,1339,1365]" pageId="38" pageNumber="39" pagination="289 - 294" refId="ref82238" refString="Roberts, E. M., Stevens, N. J., O'Connor, P. M., Dirks, P. H. G. M., Gottfried, M. D., Clyde, W. C., Armstrong, R. A., Kemp, A. I. S. &amp; Hemming, S. (2012) Initiation of the western branch of the East African Rift coeval with the eastern branch. Nature Geoscience, 5, 289 - 294. https: // doi. org / 10.1038 / NGEO 1432" type="journal article" year="2012">
Roberts
<emphasis id="B9010172CC174C02F61DFAD4FE11FAE0" box="[341,399,1339,1365]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2012
</bibRefCitation>
). The Nyika Plateau is the largest isolated upland area in
<collectingCountry id="F3629DF0CC174C02F32EFAD4FB20FAE0" box="[1126,1214,1339,1365]" name="Malawi" pageId="38" pageNumber="39">Malawi</collectingCountry>
, and belongs to the Afromontane archipelago-like regional centre of endemism (
<emphasis id="B9010172CC174C02F400FAB0FC18FACD" box="[840,902,1375,1400]" italics="true" pageId="38" pageNumber="39">sensu</emphasis>
<bibRefCitation id="EFE4A091CC174C02F4C6FAB0FB89FACC" author="White, F." box="[910,1047,1375,1401]" pageId="38" pageNumber="39" refId="ref83858" refString="White, F. (1983) The Vegetation of Africa: a Descriptive Memoir to Accompany the Unesco / AETFAT / UNSO Vegetation Map of Africa. Natural Resources Research No. XX. Unesco, Paris, 356 pp." type="book" year="1983">White 1983</bibRefCitation>
). This species can be classified an Afromontane near-endemic species.
</paragraph>
<paragraph id="8BCADD60CC174C02F78FFA49FE77FA74" blockId="38.[151,1437,151,1581]" box="[199,489,1446,1473]" pageId="38" pageNumber="39">
<emphasis id="B9010172CC174C02F78FFA49FEB4FA75" bold="true" box="[199,298,1446,1472]" pageId="38" pageNumber="39">Habitat.</emphasis>
See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC174C02F78FFA24FC66F998" pageId="38" pageNumber="39" type="etymology">
<paragraph id="8BCADD60CC174C02F78FFA24FC66F998" blockId="38.[151,1437,151,1581]" pageId="38" pageNumber="39">
<emphasis id="B9010172CC174C02F78FFA24FED0FA50" bold="true" box="[199,334,1483,1509]" pageId="38" pageNumber="39">Etymology.</emphasis>
<taxonomicName id="4C75A6E3CC174C02F61EFA24FDA5FA51" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[342,571,1483,1509]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC174C02F61EFA24FDA5FA51" box="[342,571,1483,1509]" italics="true" pageId="38" pageNumber="39">Lukeniana stueningi</emphasis>
</taxonomicName>
is named in honour of Dr. Dieter Stüning for his constant, valuable, and patient support of the first author’s studies on the
<taxonomicName id="4C75A6E3CC174C02F5F9FA00FCDBF9BC" baseAuthorityName="Lehmann" baseAuthorityYear="2019" box="[689,837,1519,1545]" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
at the ZFMK in 2009–2014, and in particular for his help on the publication of the new genus
<taxonomicName id="4C75A6E3CC174C02F52DF9FCFC6BF998" authority="Lehmann &amp; Rajaei, 2013" authorityName="Lehmann &amp; Rajaei" authorityYear="2013" box="[613,1013,1555,1581]" class="Insecta" family="Metarbelidae" genus="Shimonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B9010172CC174C02F52DF9FCFD51F998" box="[613,719,1555,1581]" italics="true" pageId="38" pageNumber="39">Shimonia</emphasis>
<bibRefCitation id="EFE4A091CC174C02F59EF9FCFC6BF998" author="Lehmann, I. &amp; Rajaei, H." box="[726,1013,1555,1581]" pageId="38" pageNumber="39" pagination="100 - 110" refId="ref80199" refString="Lehmann, I. &amp; Rajaei, H. (2013) Description of a new genus and three new species of Metarbelidae (Lepidoptera: Cossoidea) from East and Central Africa, with notes on biogeography. Bonn zoological Bulletin, 62 (1), 100 - 110. https: // doi. org / 10.5962 / bhl. title. 79421" type="journal article" year="2013">Lehmann &amp; Rajaei, 2013</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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