treatments-xml/data/03/DA/D5/03DAD52BFFB0FFFDFD19BDA85D18A2C0.xml

<document id="3573BF5EA7B1668482BA49096A46483B" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Müller, Patrick" docDate="2022" docId="03DAD52BFFB0FFFDFD19BDA85D18A2C0" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus kachinensis Fanti &amp; Müller, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="354" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="352" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="5F94A522D8EDF62B01FBAFE9EA2E9AD8">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="BCB696352C9A1B0B97CDB9891061E111">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="2A4308EDA494FAD8A298452F792C26DF">Müller, Patrick</mods:namePart>
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<mods:date id="917BB20F1619E4961D2C95694178814D">2022</mods:date>
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<treatment id="03DAD52BFFB0FFFDFD19BDA85D18A2C0" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0" httpUri="http://treatment.plazi.org/id/03DAD52BFFB0FFFDFD19BDA85D18A2C0" lastPageId="23" lastPageNumber="354" pageId="21" pageNumber="352">
<subSubSection id="C36937B6FFB0FFFFFD19BDA85CBFA1C2" box="[643,1017,154,178]" pageId="21" pageNumber="352" type="nomenclature">
<paragraph id="8BCC643DFFB0FFFFFD19BDA85CBFA1C2" blockId="21.[643,1017,154,210]" box="[643,1017,154,178]" pageId="21" pageNumber="352">
<heading id="D084D351FFB0FFFFFD19BDA85CBFA1C2" bold="true" box="[643,1017,154,178]" fontSize="10" level="2" pageId="21" pageNumber="352" reason="4">
<emphasis id="B907B82FFFB0FFFFFD19BDA85CBFA1C2" bold="true" box="[643,1017,154,178]" pageId="21" pageNumber="352">
<taxonomicName id="4C731FBEFFB0FFFFFD19BDA85CD8A1C2" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[643,926,154,178]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="352" phylum="Arthropoda" rank="species" species="kachinensis" status="sp. nov.">
<emphasis id="B907B82FFFB0FFFFFD19BDA85CD8A1C2" bold="true" box="[643,926,154,178]" italics="true" pageId="21" pageNumber="352">Sanaungulus kachinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB0FFFFFC3EBDA85CBFA1C2" box="[932,1017,154,178]" pageId="21" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB0FFFFFD19BD895DA6A1A2" box="[643,736,187,210]" pageId="21" pageNumber="352" type="description">
<paragraph id="8BCC643DFFB0FFFFFD19BD895DA6A1A2" blockId="21.[643,1017,154,210]" box="[643,736,187,210]" pageId="21" pageNumber="352">
(
<figureCitation id="134878B8FFB0FFFFFD11BD895D91A1A2" box="[651,727,187,210]" captionStart="Fig" captionStartId="22.[151,190,1665,1688]" captionTargetBox="[151,1171,412,1637]" captionTargetId="figure-115@22.[151,1171,412,1637]" captionTargetPageId="22" captionText="Fig. 12. Sanaungulus kachinensis sp. nov. in Burmese amber. A–B: Holotype, lateral views." figureDoi="http://doi.org/10.5281/zenodo.13203201" httpUri="https://zenodo.org/record/13203201/files/figure.png" pageId="21" pageNumber="352">Fig. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB0FFFFFD19BDC35CE4A346" pageId="21" pageNumber="352" type="materials_examined">
<paragraph id="8BCC643DFFB0FFFFFD19BDC35B69A0F8" blockId="21.[643,1134,241,392]" pageId="21" pageNumber="352">
<materialsCitation id="3B1B6E60FFB0FFFFFD19BDC35B69A0F8" collectingDate="2021-12-14" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="21" pageNumber="352" specimenCode="BUB4312" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFB0FFFFFD19BDC35DB7A079" bold="true" box="[643,753,241,265]" pageId="21" pageNumber="352">
<typeStatus id="54C8DA9FFFB0FFFFFD19BDC35DABA079" box="[643,749,241,265]" pageId="21" pageNumber="352" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFB0FFFFFD6DBDC35C76A078" box="[759,816,241,264]" pageId="21" pageNumber="352">Male</collectingCounty>
, adult specimen in a 
<collectingMunicipality id="6BA8FE47FFB0FFFFFB95BDC35B28A078" box="[1039,1134,241,264]" pageId="21" pageNumber="352">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFB0FFFFFD10BC235D9CA058" box="[650,730,273,296]" country="Myanmar" name="Kachin" pageId="21" pageNumber="352">Kachin</collectingRegion>
) amber piece: 
<location id="8EAC32E6FFB0FFFFFCE9BC235C80A058" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFCE9BC235C80A058" box="[883,966,273,296]" county="Male" municipality="Burmese" name="Catalog" pageId="21" pageNumber="352" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG 
<date id="FFCD42FDFFB0FFFFFD54BC035C10A038" box="[718,854,305,328]" pageId="21" pageNumber="352" value="2021-12-14">
<collectingDate id="EF89BB15FFB0FFFFFD54BC035C10A038" box="[718,854,305,328]" pageId="21" pageNumber="352" value="2021-12-14">
2021 XII 
<quantity id="4C8BC9D8FFB0FFFFFCA1BC035C10A038" box="[827,854,305,328]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="21" pageNumber="352" unit="in" value="14.0">14</quantity>
</collectingDate>
</date>
in the 
<location id="8EAC32E6FFB0FFFFFC3CBC035DB4A018" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFC3CBC035DB4A018" county="Male" municipality="Burmese" name="Bayerische Staatssammlung" pageId="21" pageNumber="352" stateProvince="Kachin">Bayerische Staatssammlung</location>
für 
<location id="8EAC32E6FFB0FFFFFC87BC635CF6A018" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFC87BC635CF6A018" box="[797,944,337,360]" county="Male" municipality="Burmese" name="Palaontologie" pageId="21" pageNumber="352" stateProvince="Kachin">Paläontologie</location>
und 
<location id="8EAC32E6FFB0FFFFFC7EBC635B00A018" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFC7EBC635B00A018" box="[996,1094,337,360]" county="Male" municipality="Burmese" name="Geologie" pageId="21" pageNumber="352" stateProvince="Kachin">Geologie</location>
(ex 
<location id="8EAC32E6FFB0FFFFFD19BC435D99A0F8" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFD19BC435D99A0F8" box="[643,735,369,392]" county="Male" municipality="Burmese" name="Muller's" pageId="21" pageNumber="352" stateProvince="Kachin">Müller’s</location>
collection number: 
<specimenCode id="DBD5CC46FFB0FFFFFC29BC435B62A0F8" box="[947,1060,369,392]" collectionCode="BUB" pageId="21" pageNumber="352">BUB4312</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFB0FFFFFD19BC9A5C93A0AF" blockId="21.[643,1134,424,479]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFD19BC9A5C5AA0B0" bold="true" box="[643,796,424,448]" pageId="21" pageNumber="352">Type locality.</emphasis>
<collectingCountry id="F36424ADFFB0FFFFFCBEBC9A5CD6A0CF" box="[804,912,424,447]" name="Myanmar" pageId="21" pageNumber="352">Myanmar</collectingCountry>
: 
<collectingRegion id="49B7AADFFFB0FFFFFC01BC9A5B63A0CF" box="[923,1061,424,447]" country="Myanmar" name="Kachin" pageId="21" pageNumber="352">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFB0FFFFFD19BCCC5CE4A346" blockId="21.[643,1134,510,566]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFD19BCCC5C50A366" bold="true" box="[643,790,510,534]" pageId="21" pageNumber="352">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB0FFFFFD19BF675CC8A2BC" pageId="21" pageNumber="352" type="diagnosis">
<paragraph id="8BCC643DFFB0FFFFFD19BF675CC8A2BC" blockId="21.[643,1134,597,972]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFD19BF675C31A31D" bold="true" box="[643,887,597,621]" pageId="21" pageNumber="352">Differential diagnosis.</emphasis>
The new species is characterized by antennae equipped with an antennal process near the base of antennomeres IV– X. It differs from 
<taxonomicName id="4C731FBEFFB0FFFFFCDCBF845B54A3BD" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[838,1042,694,717]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="352" phylum="Arthropoda" rank="species" species="havai" status="sp. nov.">
<emphasis id="B907B82FFFB0FFFFFCDCBF845B54A3BD" box="[838,1042,694,717]" italics="true" pageId="21" pageNumber="352">Sanaungulus havai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB0FFFFFB81BF875B28A3BC" box="[1051,1134,693,716]" pageId="21" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
based on the head that is less triangular behind the eyes, the pronotum with a median incision, and by antennae with longer rami. It differs from 
<taxonomicName id="4C731FBEFFB0FFFFFD19BE045B12A23C" authority="Y. Yang, Liu &amp; Zhao" authorityName="Y. Yang, Liu &amp; Zhao" box="[643,1108,821,845]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="352" phylum="Arthropoda" rank="species" species="ruficollis">
<emphasis id="B907B82FFFB0FFFFFD19BE045C34A23D" box="[643,882,822,845]" italics="true" pageId="21" pageNumber="352">Sanaungulus ruficollis</emphasis>
Y. Yang, Liu &amp; Zhao
</taxonomicName>
in Y. Yang, Zhao, Geiser, W. Zhang, Ren, Bai &amp; Liu, 2022 based on the antennal processes, the pronotum incised, and by longer metathoracic wings (
<bibRefCitation id="EFE219CCFFB0FFFFFD4ABE875CC5A2BC" author="Yang Y. &amp; Zhao W. &amp; Geiser M. F. &amp; Zhang W. &amp; Ren D. &amp; Bai M. &amp; Liu H." box="[720,899,949,972]" pageId="21" pageNumber="352" refId="ref21309" refString="Yang Y., Zhao W., Geiser M. F., Zhang W., Ren D., Bai M., Liu H. 2022. Taxonomic revision of the mid-Cretaceous genus Sanaungulus Fanti, Damgaard et Ellenberger, 2018 (Coleoptera, Cantharidae) from northern Myanmar based on a morphology-based phylogeny. Cretaceous Research 135: 105217 [14 pp. + supplementary data]." type="book" year="2022">Yang et al. 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB0FFFDFD19BEDE5D57A31E" lastPageId="23" lastPageNumber="354" pageId="21" pageNumber="352" type="description">
<paragraph id="8BCC643DFFB0FFFFFD19BEDE5B14A533" blockId="21.[643,1135,1004,1091]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFD19BEDE5C4EA574" bold="true" box="[643,776,1004,1028]" pageId="21" pageNumber="352">Description.</emphasis>
Adult, winged, slender. Male, defined on the shape of the last abdominal segments. Body length about 
<quantity id="4C8BC9D8FFB0FFFFFCD6B91E5CE7A533" box="[844,929,1068,1091]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="21" pageNumber="352" unit="mm" value="2.8">2.8 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFB0FFFDFD19B9515D57A31E" blockId="21.[643,1135,1123,1690]" lastBlockId="23.[113,605,155,622]" lastPageId="23" lastPageNumber="354" pageId="21" pageNumber="352">Head elongated, slightly constricted behind the eyes, fitted with small rugosity. Eyes very prominent, roundish, inserted in the upper and dorsolateral part of the head. Mandibles elongated, very robust, falciform.Maxillary palps 4­ segmented, with the last palpomere strongly securiform. Labial palps 3­ segmented, with the last palpomere securiform.Antennae 11­ segmented, pectinate, slender, long, surpassing the apex of the elytra; scape elongated, robust; antennomere II short; antennomere III filiform, about 1.9 times longer than second; antennomeres IV–X subequal in length, each equipped near their base with a very long, robust, almost straight antennal process (longer than antennomere), almost rounded apically; antennomere XI filiform, extremely long, rounded apically; all the antennomeres with short pubescence. Pronotum longer than wide, as wide as head, anterior margin rounded and bordered, posterior margin rather straight, corners obtuse, sides almost straight, pronotum disc with a transversal median and deep incision, equipped with short setae. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, as wide as pronotum, sub­parallel, rounded apexes, surface with shallow punctuation and several short setae. Metathoracic wings transparent, long, surpassing the elytra and reaching the last abdominal segments. Sternum elongated, almost straight posteriorly, sternites transverse equipped with short setae; last tergite elongated, wide and strongly globular (spatuliform); last sternite elongated, very thin. Legs thin, long, pubescent; coxae massive, elongated, with rounded apex; trochanters elongated, robust, with rounded apex; femora slightly curved, more robust than tibiae; tibiae cylindrical, thin, without spurs near the apex, pro­ and mesotibiae slightly shorter than pro­ and mesofemora, metatibiae slightly longer than metafemora. Tarsal formula 5­5­5; first tarsomere long; second tarsomere about 1.2 times shorter than first; third tarsomere triangular­shaped, short, straight apically; fourth tarsomere short, strongly bilobed; fifth tarsomere elongated, thin, flat and strongly curved; claws simple, apparently without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFB3FFFCFF0DBBB35B2EA7E8" ID-DOI="http://doi.org/10.5281/zenodo.13203201" ID-Zenodo-Dep="13203201" box="[151,1128,1665,1688]" httpUri="https://zenodo.org/record/13203201/files/figure.png" pageId="22" pageNumber="353" startId="22.[151,190,1665,1688]" targetBox="[151,1171,412,1637]" targetPageId="22" targetType="figure">
<paragraph id="8BCC643DFFB3FFFCFF0DBBB35B2EA7E8" blockId="22.[151,1128,1665,1688]" box="[151,1128,1665,1688]" pageId="22" pageNumber="353">
Fig. 12. 
<taxonomicName id="4C731FBEFFB3FFFCFF6ABBB35EBBA7E8" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[240,509,1665,1688]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="353" phylum="Arthropoda" rank="species" species="kachinensis" status="sp. nov.">
<emphasis id="B907B82FFFB3FFFCFF6ABBB35EBBA7E8" box="[240,509,1665,1688]" italics="true" pageId="22" pageNumber="353">Sanaungulus kachinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB3FFFCFD99BBB35D13A7E8" box="[515,597,1665,1688]" pageId="22" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A–B: Holotype, lateral views.
</paragraph>
</caption>
<subSubSection id="C36937B6FFB2FFFDFFEBBFBF5D18A26A" pageId="23" pageNumber="354" type="etymology">
<paragraph id="8BCC643DFFB2FFFDFFEBBFBF5E19A394" blockId="23.[113,605,653,741]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFFEBBFBF5FB7A3D5" bold="true" box="[113,241,653,677]" pageId="23" pageNumber="354">Etymology.</emphasis>
Derived from the toponym 
<collectingRegion id="49B7AADFFFB2FFFDFD8ABFBF5D1AA3D4" box="[528,604,653,676]" country="Myanmar" name="Kachin" pageId="23" pageNumber="354">Kachin</collectingRegion>
(the locality where the amber mines are located) + the Latin suffix ­Á 
<emphasis id="B907B82FFFB2FFFDFEB4BFFF5E1EA394" box="[302,344,717,740]" italics="true" pageId="23" pageNumber="354">nsis</emphasis>
.
</paragraph>
<paragraph id="8BCC643DFFB2FFFDFFEBBE315D18A26A" blockId="23.[113,606,771,795]" box="[113,606,771,795]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFFEBBE315E4CA26B" bold="true" box="[113,266,771,795]" pageId="23" pageNumber="354">Syninclusions.</emphasis>
Air bubbles, detritus, one 
<taxonomicName id="4C731FBEFFB2FFFDFD91BE315D1CA26A" box="[523,602,771,794]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="354" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB2FFFDFFEBBE0B5D18A2C0" pageId="23" pageNumber="354" type="discussion">
<paragraph id="8BCC643DFFB2FFFDFFEBBE0B5D18A2C0" blockId="23.[113,606,825,944]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFFEBBE0B5F9BA221" bold="true" box="[113,221,825,849]" pageId="23" pageNumber="354">Remarks.</emphasis>
Piece of clear (surface is slightly blurred), golden amber that measures 11x9x 
<quantity id="4C8BC9D8FFB2FFFDFD9DBE6B5D1FA200" box="[519,601,857,880]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="354" unit="mm" value="2.5">2.5 mm</quantity>
. The inclusion is complete except for the right antenna preserved until the tenth antennomere.
</paragraph>
</subSubSection>
</treatment>
</document>