164 lines
19 KiB
XML
164 lines
19 KiB
XML
<document id="2B2BCAAB4344353B774F96BA301F95B5" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio & Müller, Patrick" docDate="2022" docId="03DAD52BFFACFFE0FFEBBFA45BD2A03C" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Hukawngichthyurus barsevskisi Fanti & Müller, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="341" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="340" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:titleInfo id="52F48DA837B864D95E1DBD76F663DE0C">
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<mods:title id="F34500931C73FDAB89E7B6E4BD8C355D">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="1FAD1A7529DBE88C1C0890A33A04D63C">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="9949C7F4C2E70200D0EBFC4CB8ED0815">Müller, Patrick</mods:namePart>
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<mods:typeOfResource id="EF05BBC133618F758103093EA11F6DD0">text</mods:typeOfResource>
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<mods:titleInfo id="32698044CD5F400DEE7AA3C042E71292">
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<mods:title id="109CDB8BE1E9054A5265FD764694BC30">Baltic Journal of Coleopterology</mods:title>
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<mods:part id="E6C3F5F97E1D578614D41F28CDFBE3DB">
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<mods:date id="B165F94B96476A1FCACCDAA50F932C22">2022</mods:date>
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<mods:detail id="8E259EBA530A52AA1CFD87531E64D180" type="volume">
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<mods:number id="3FA1E68D7EBFE12B669C3FDA43B08E0D">22</mods:number>
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<mods:detail id="95C401E239733F50B0F5EBB77BE00E2D" type="issue">
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<mods:number id="4E1F6FD02E6D74E6065EBE73DDC97E15">2</mods:number>
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<mods:classification id="B6698A72455BB0BCAB601125E3A0D727">journal article</mods:classification>
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<mods:identifier id="E75C12165D82650BF59240DE7D594538" type="DOI">10.5281/zenodo.13203160</mods:identifier>
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<mods:identifier id="D8E39AD6F9FD621F4D22C406AA9CA0CD" type="ISSN">1407-8619</mods:identifier>
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<treatment id="03DAD52BFFACFFE0FFEBBFA45BD2A03C" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C" httpUri="http://treatment.plazi.org/id/03DAD52BFFACFFE0FFEBBFA45BD2A03C" lastPageId="10" lastPageNumber="341" pageId="9" pageNumber="340">
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<subSubSection id="C36937B6FFACFFE3FFEBBFA45D68A3DE" box="[113,558,662,686]" pageId="9" pageNumber="340" type="nomenclature">
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<paragraph id="8BCC643DFFACFFE3FFEBBFA45D68A3DE" blockId="9.[113,558,662,718]" box="[113,558,662,686]" pageId="9" pageNumber="340">
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<heading id="D084D351FFACFFE3FFEBBFA45D68A3DE" bold="true" box="[113,558,662,686]" fontSize="10" level="2" pageId="9" pageNumber="340" reason="4">
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<emphasis id="B907B82FFFACFFE3FFEBBFA45D68A3DE" bold="true" box="[113,558,662,686]" pageId="9" pageNumber="340">
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<taxonomicName id="4C731FBEFFACFFE3FFEBBFA45E95A3DE" authorityName="Fanti & Müller" authorityYear="2022" box="[113,467,662,686]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="barsevskisi" status="sp. nov.">
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<emphasis id="B907B82FFFACFFE3FFEBBFA45E95A3DE" bold="true" box="[113,467,662,686]" italics="true" pageId="9" pageNumber="340">Hukawngichthyurus barsevskisi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFACFFE3FE43BFA45D68A3DE" box="[473,558,662,686]" pageId="9" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FFACFFE3FFEBBF855F87A3BE" box="[113,193,695,718]" pageId="9" pageNumber="340" type="description">
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<paragraph id="8BCC643DFFACFFE3FFEBBF855F87A3BE" blockId="9.[113,558,662,718]" box="[113,193,695,718]" pageId="9" pageNumber="340">
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(
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<figureCitation id="134878B8FFACFFE3FFE3BF855FFFA3BE" box="[121,185,695,718]" captionStart="Fig" captionStartId="10.[151,190,1639,1662]" captionTargetBox="[151,1172,386,1611]" captionTargetId="figure-18@10.[151,1172,386,1611]" captionTargetPageId="10" captionText="Fig. 5. Hukawngichthyurus barsevskisi sp. nov. in Burmese amber. A–B: Holotype, dorsolateral views." figureDoi="http://doi.org/10.5281/zenodo.13203179" httpUri="https://zenodo.org/record/13203179/files/figure.png" pageId="9" pageNumber="340">Fig. 5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFACFFE3FFEBBFDC5ED6A549" pageId="9" pageNumber="340" type="materials_examined">
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<paragraph id="8BCC643DFFACFFE3FFEBBFDC5D58A2F9" blockId="9.[113,605,750,905]" pageId="9" pageNumber="340">
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<materialsCitation id="3B1B6E60FFACFFE3FFEBBFDC5D58A2F9" collectingDate="2021-12-19" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="9" pageNumber="340" specimenCode="BUB4328" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="B907B82FFFACFFE3FFEBBFDC5FA6A276" bold="true" box="[113,224,750,774]" pageId="9" pageNumber="340">
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<typeStatus id="54C8DA9FFFACFFE3FFEBBFDC5F9DA276" box="[113,219,750,774]" pageId="9" pageNumber="340" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62AD1CB1FFACFFE3FF7DBFDC5E66A275" box="[231,288,750,773]" pageId="9" pageNumber="340">Male</collectingCounty>
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, adult specimen in a
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<collectingMunicipality id="6BA8FE47FFACFFE3FE64BFDC5D1BA275" box="[510,605,750,773]" pageId="9" pageNumber="340">Burmese</collectingMunicipality>
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||
(
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<collectingRegion id="49B7AADFFFACFFE3FFE3BE3D5F8FA256" box="[121,201,783,806]" country="Myanmar" name="Kachin" pageId="9" pageNumber="340">Kachin</collectingRegion>
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) amber piece:
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<location id="8EAC32E6FFACFFE3FEF8BE3D5EF3A256" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FEF8BE3D5EF3A256" box="[354,437,783,806]" county="Male" municipality="Burmese" name="Catalog" pageId="9" pageNumber="340" stateProvince="Kachin">Catalog</location>
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number SNSBBSPG
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<date id="FFCD42FDFFACFFE3FF27BE025E03A237" box="[189,325,816,839]" pageId="9" pageNumber="340" value="2021-12-19">
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<collectingDate id="EF89BB15FFACFFE3FF27BE025E03A237" box="[189,325,816,839]" pageId="9" pageNumber="340" value="2021-12-19">
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2021 XII
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<quantity id="4C8BC9D8FFACFFE3FEB0BE025E03A237" box="[298,325,816,839]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="9" pageNumber="340" unit="in" value="19.0">19</quantity>
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</collectingDate>
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</date>
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in the
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<location id="8EAC32E6FFACFFE3FE0FBE025FA6A218" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FE0FBE025FA6A218" county="Male" municipality="Burmese" name="Bayerische Staatssammlung" pageId="9" pageNumber="340" stateProvince="Kachin">Bayerische Staatssammlung</location>
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für
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<location id="8EAC32E6FFACFFE3FE96BE635ED9A218" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FE96BE635ED9A218" box="[268,415,849,872]" county="Male" municipality="Burmese" name="Palaontologie" pageId="9" pageNumber="340" stateProvince="Kachin">Paläontologie</location>
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und
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<location id="8EAC32E6FFACFFE3FE49BE635D73A218" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FE49BE635D73A218" box="[467,565,849,872]" county="Male" municipality="Burmese" name="Geologie" pageId="9" pageNumber="340" stateProvince="Kachin">Geologie</location>
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(ex
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<location id="8EAC32E6FFACFFE3FFEBBE405F8BA2F9" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FFEBBE405F8BA2F9" box="[113,205,882,905]" county="Male" municipality="Burmese" name="Muller's" pageId="9" pageNumber="340" stateProvince="Kachin">Müller’s</location>
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collection number:
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<specimenCode id="DBD5CC46FFACFFE3FE38BE405D55A2F9" box="[418,531,882,905]" collectionCode="BUB" pageId="9" pageNumber="340">BUB4328</specimenCode>
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).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BCC643DFFACFFE3FFEBBE9B5E82A291" blockId="9.[113,605,937,993]" pageId="9" pageNumber="340">
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<emphasis id="B907B82FFFACFFE3FFEBBE9B5E4CA2B1" bold="true" box="[113,266,937,961]" pageId="9" pageNumber="340">Type locality.</emphasis>
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||
<collectingCountry id="F36424ADFFACFFE3FE89BE9B5E39A2B0" box="[275,383,937,960]" name="Myanmar" pageId="9" pageNumber="340">Myanmar</collectingCountry>
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:
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<collectingRegion id="49B7AADFFFACFFE3FE10BE9B5D52A2B0" box="[394,532,937,960]" country="Myanmar" name="Kachin" pageId="9" pageNumber="340">Kachin state</collectingRegion>
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, Myitkyina District, Hukawng Valley.
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</paragraph>
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<paragraph id="8BCC643DFFACFFE3FFEBB9335ED6A549" blockId="9.[113,605,1025,1081]" pageId="9" pageNumber="340">
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<emphasis id="B907B82FFFACFFE3FFEBB9335E43A569" bold="true" box="[113,261,1025,1049]" pageId="9" pageNumber="340">Type horizon.</emphasis>
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Lowermost Cenomanian (98.79 ± 0.62 Ma), midCretaceous.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C36937B6FFACFFE3FFEBB96B5E11A76B" pageId="9" pageNumber="340" type="diagnosis">
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<paragraph id="8BCC643DFFACFFE3FFEBB96B5E11A76B" blockId="9.[113,605,1113,1563]" pageId="9" pageNumber="340">
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||
<emphasis id="B907B82FFFACFFE3FFEBB96B5E3EA501" bold="true" box="[113,376,1113,1137]" pageId="9" pageNumber="340">Differential diagnosis.</emphasis>
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The new species is distinguishable from other
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<taxonomicName id="4C731FBEFFACFFE3FE12B9495D1AA5E2" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[392,604,1147,1170]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="genus">
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<emphasis id="B907B82FFFACFFE3FE12B9495D1AA5E2" box="[392,604,1147,1170]" italics="true" pageId="9" pageNumber="340">Hukawngichthyurus</emphasis>
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</taxonomicName>
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by its pectinate antennae. We attribute this new species to this genus on the basis of the very short and irregular antennomeres II–III, for the pronotum with depressions and for the last abdominal segments in a kind of “fish tail.” The antennae are often a variable character. For example, in the fossil genus
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<emphasis id="B907B82FFFACFFE3FEB3B8525EF2A407" box="[297,436,1376,1399]" italics="true" pageId="9" pageNumber="340">
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<taxonomicName id="4C731FBEFFACFFE3FEB3B8525EF6A407" authorityName="Fanti, Damgaard & Ellenberger" authorityYear="2018" box="[297,432,1376,1399]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="genus">Sanaungulus</taxonomicName>
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,
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</emphasis>
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there are species with pectinate or filiform antennae. Unlike what
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<bibRefCitation id="EFE219CCFFACFFE3FFEBB8905E6AA4C9" author="Yang Y. & Zhao W. & Geiser M. F. & Zhang W. & Bai M. & Liu H." box="[113,300,1442,1465]" pageId="9" pageNumber="340" refId="ref21251" refString="Yang Y., Zhao W., Geiser M. F., Zhang W., Bai M., Liu H. 2021. Review of the genus Burmomiles Fanti, Damgaard & Ellenberger from mid-Cretaceous Burmese amber (Coleoptera, Cantharidae). Cretaceous Research 128: 104961 [9 pp.]." type="book" year="2021">Yang et al. (2021)</bibRefCitation>
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say about
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<taxonomicName id="4C731FBEFFACFFE3FE00B8905EE1A4AA" authority="(Hsiao & Huang, 2018)" baseAuthorityName="Hsiao & Huang" baseAuthorityYear="2018" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="ruicheni">
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<emphasis id="B907B82FFFACFFE3FE00B8905FD9A4AA" italics="true" pageId="9" pageNumber="340">Sanaungulus ruicheni</emphasis>
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(
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<bibRefCitation id="EFE219CCFFACFFE3FF34B8F15ED9A4AA" author="Hsiao Y. & Huang C. - L." box="[174,415,1475,1498]" pageId="9" pageNumber="340" pagination="257 - 263" refId="ref20597" refString="Hsiao Y., Huang C. - L. 2018. Taxonomic revision on the genus Ornatomalthinus Poinar and Fanti (Coleoptera: Cantharidae), with description of a new species from the Cretaceous Burmese amber. Cretaceous Research 92: 257 - 263." type="journal article" year="2018">Hsiao & Huang, 2018</bibRefCitation>
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)
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</taxonomicName>
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, this is not a distinguishing character, being extremely variable even in living genera.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FFACFFE0FFEBBB095EF6A1A3" lastPageId="10" lastPageNumber="341" pageId="9" pageNumber="340" type="description">
|
||
<paragraph id="8BCC643DFFACFFE3FFEBBB095EA6A7E4" blockId="9.[113,605,1595,1684]" pageId="9" pageNumber="340">
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||
<emphasis id="B907B82FFFACFFE3FFEBBB095FBCA723" bold="true" box="[113,250,1595,1619]" pageId="9" pageNumber="340">Description.</emphasis>
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Adult, alate, slender. Male, for the shape of the last abdominal segment. Entirely dark brown. Body length:
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<quantity id="4C8BC9D8FFACFFE3FE12BB4F5E9AA7E4" box="[392,476,1661,1684]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="340" unit="mm" value="3.4">3.4 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BCC643DFFACFFE0FD19BDA95EF6A1A3" blockId="9.[643,1134,155,1682]" lastBlockId="10.[151,643,155,211]" lastPageId="10" lastPageNumber="341" pageId="9" pageNumber="340">Head rounded, convex, slightly restricted behind the eyes, surface with shallow punctation and short setae. Eyes large and prominent, subelliptical, inserted laterally to the head. Rostrum elongated. Mandibles falciform. Maxillary palps 4 segmented, last palpomere securiform with rounded apex. Labial palps 3 segmented, last palpomere stout and slightly securiform.Antennae 11 segmented, relatively short, pectinate, surpassing the half of elytra and reaching the half of abdomen, insertion of the antennae close to the eyes and in their lower part; scape robust, elongated, clubshaped; antennomere II very short, moniliformglobular, asymmetrical; antennomere III short, sturdier and longer than second, slightly irregulardistorted; antennomeres IV–X pectinate, equipped with a lobe which is very long (longer than antennomeres), rounded at apex and inserted in the basal part of antennomeres; antennomere XI filiform, rounded at apex; all antennomeres punctate with not erected and very short setae. Pronotum transverse, slightly wider than head, anterior margin undulate, posterior margin undulate and bordered, sides irregular (almost straight), surface irregular with draft and depressions (particularly near the anterior margin) and equipped with short setae. Scutellum triangular. Elytra wider than pronotum, parallelsided, rounded and slightly dehiscent at apex, rather short and revealing the last three abdominal segments, surface smooth without punctation, equipped with few little and thin striae. Metathoracic wings transparent, folded, very slightly surpassing the elytra. Metasternum subquadrate; sternites transverse. Last urite with an apical lobe in each side, which is extremely long, thin, with rounded apex. Legs slender, rather long, with pubescence; coxae elongated and robust; trochanters globularshaped, irregular, with apex rounded; femora robust and not curved, almost flat, sturdier and longer than tibiae; tibiae cylindrical, thin, without spurs at apex. Tarsal formula 5 5 5; first tarsomere elongated, about 1.2 times longer than second; second tarsomere thin at base and slightly enlarged apically; third tarsomere triangular and shorter than second; fourth tarsomere bilobed with the lobes very long and not particularly wide; fifth tarsomere very long; claws simple without denticles or lobes.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FFAFFFE0FF0DBDC15C96A1A3" pageId="10" pageNumber="341" type="etymology">
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<paragraph id="8BCC643DFFAFFFE0FF0DBDC15EDAA03C" blockId="10.[151,643,237,332]" pageId="10" pageNumber="341">
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||
<emphasis id="B907B82FFFAFFFE0FF0DBDC15E5FA07B" bold="true" box="[151,281,243,267]" pageId="10" pageNumber="341">Etymology.</emphasis>
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||
Named in honor of ArvÓds Barševskis (
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<collectingRegion id="49B7AADFFFAFFFE0FF6FBC265E36A05B" box="[245,368,276,299]" country="Latvia" name="Daugavpils" pageId="10" pageNumber="341">Daugavpils</collectingRegion>
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University,
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<collectingCountry id="F36424ADFFAFFFE0FE62BC265D06A05B" box="[504,576,276,299]" name="Latvia" pageId="10" pageNumber="341">Latvia</collectingCountry>
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), dear friend of the first author.
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</paragraph>
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<paragraph id="8BCC643DFFAFFFE0FD32BDA95C96A1A3" blockId="10.[680,1172,155,211]" pageId="10" pageNumber="341">
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<emphasis id="B907B82FFFAFFFE0FD32BDA95C14A1C3" bold="true" box="[680,850,155,179]" pageId="10" pageNumber="341">Syninclusions.</emphasis>
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Remains of another soldier beetle, air bubbles, detritus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFAFFFE0FD32BDC15BD2A03C" pageId="10" pageNumber="341" type="discussion">
|
||
<paragraph id="8BCC643DFFAFFFE0FD32BDC15BD2A03C" blockId="10.[680,1172,243,332]" pageId="10" pageNumber="341">
|
||
<emphasis id="B907B82FFFAFFFE0FD32BDC15C52A07B" bold="true" box="[680,788,243,267]" pageId="10" pageNumber="341">Remarks.</emphasis>
|
||
Piece of clear, golden amber that measures 11x9x
|
||
<quantity id="4C8BC9D8FFAFFFE0FC87BC265C10A05B" box="[797,854,276,299]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="341" unit="mm" value="3.0">3 mm</quantity>
|
||
.The inclusion is complete except for the left anterior leg preserved until the tibia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |