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<document id="D10DAD1A8F4CB3D14762AEAC03A8F69F" ID-CLB-Dataset="25918" ID-DOI="10.11646/zootaxa.4659.1.1" ID-GBIF-Dataset="0cbe0b8e-2019-4f5a-8336-2d4d9fcd8bf6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3377239" ID-ZooBank="57309E14-0330-4ED7-BCDA-355EE6618215" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566808762347" checkinUser="plazi" docAuthor="Whitworth, Terry L. &amp; Yusseff-Vanegas, Sohath" docDate="2019" docId="03DA87E60F1DFFF2FF19B97E35EBF822" docLanguage="en" docName="zootaxa.4659.1.1.pdf" docOrigin="Zootaxa 4659 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Laneella patriciae" docType="treatment" docVersion="20" lastPageNumber="40" masterDocId="FFE3FF9E0F3AFFD5FF8EBA27342CFFB3" masterDocTitle="A revision of the genera and species of the Neotropical family Mesembrinellidae (Diptera: Oestroidea)" masterLastPageNumber="146" masterPageNumber="1" pageNumber="40" updateTime="1698752698920" updateUser="ExternalLinkService">
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<mods:title id="EF7A91767BC78FC660AA2FB54C9B588F">A revision of the genera and species of the Neotropical family Mesembrinellidae (Diptera: Oestroidea)</mods:title>
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<mods:namePart id="8A2F9966E1349905C62D34105CB4C406">Whitworth, Terry L.</mods:namePart>
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<treatment id="03DA87E60F1DFFF2FF19B97E35EBF822" ID-DOI="http://doi.org/10.5281/zenodo.5586993" ID-GBIF-Taxon="159243674" ID-Zenodo-Dep="5586993" LSID="urn:lsid:plazi:treatment:03DA87E60F1DFFF2FF19B97E35EBF822" httpUri="http://treatment.plazi.org/id/03DA87E60F1DFFF2FF19B97E35EBF822" lastPageNumber="40" pageId="39" pageNumber="40">
<subSubSection id="C369657B0F1DFFF2FF19B97E3631FCC0" box="[151,541,857,884]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8BCC36F00F1DFFF2FF19B97E3631FCC0" blockId="39.[151,541,857,884]" box="[151,541,857,884]" pageId="39" pageNumber="40">
<heading id="D084819C0F1DFFF2FF19B97E3631FCC0" bold="true" box="[151,541,857,884]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<taxonomicName id="4C734D730F1DFFF2FF19B97E3631FCC0" authority="(Wolff, 2013)" baseAuthorityName="Wolff" baseAuthorityYear="2013" box="[151,541,857,884]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="patriciae">
<emphasis id="B907EAE20F1DFFF2FF19B97E3631FCC0" bold="true" box="[151,541,857,884]" pageId="39" pageNumber="40">
<emphasis id="B907EAE20F1DFFF2FF19B97E355EFCC0" bold="true" box="[151,370,857,884]" italics="true" pageId="39" pageNumber="40">Laneella patriciae</emphasis>
(Wolff, 2013)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369657B0F1DFFF2FF19B9B93697FB83" pageId="39" pageNumber="40" type="reference_group">
<paragraph id="8BCC36F00F1DFFF2FF19B9B93013FC40" blockId="39.[151,1437,925,1073]" pageId="39" pageNumber="40">
<taxonomicName id="4C734D730F1DFFF2FF19B9B9366AFC05" authority="Wolff, 2013: 121" authorityName="Wolff" authorityPageNumber="121" authorityYear="2013" box="[151,582,925,950]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="patriciae">
<emphasis id="B907EAE20F1DFFF2FF19B9B935BCFC06" box="[151,400,926,950]" italics="true" pageId="39" pageNumber="40">Mesembrinella patriciae</emphasis>
Wolff, 2013: 121
</taxonomicName>
.
<typeStatus id="54C888520F1DFFF2FDC1B9BA3681FC05" box="[591,685,925,950]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
male (CEUA), not examined. Type locality: Filandia,
<collectingRegion id="49B7F8120F1DFFF2FB5DB9BA3105FC05" box="[1235,1321,925,950]" country="Colombia" name="Quindio" pageId="39" pageNumber="40">Quindio</collectingRegion>
,
<collectingCountry id="F36476600F1DFFF2FABCB9BA31B5FC05" box="[1330,1433,925,950]" name="Colombia" pageId="39" pageNumber="40">Colombia</collectingCountry>
. [NB: originally placed under
<taxonomicName id="4C734D730F1DFFF2FE7EB99B36A4FC67" baseAuthorityName="Marinho et al." baseAuthorityYear="2017" box="[496,648,956,980]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F1DFFF2FE7EB99B36A4FC67" box="[496,648,956,980]" italics="true" pageId="39" pageNumber="40">Mesembrinella</emphasis>
</taxonomicName>
, later moved to
<taxonomicName id="4C734D730F1DFFF2FCA2B99B37A9FC67" authorityName="Mello" authorityYear="1967" box="[812,901,956,980]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F1DFFF2FCA2B99B37A9FC67" box="[812,901,956,980]" italics="true" pageId="39" pageNumber="40">Laneella</emphasis>
</taxonomicName>
based on DNA analysis and the examination of spermathecae which revealed they are tuberform like other
<taxonomicName id="4C734D730F1DFFF2FD79B9FC301AFC40" authority="(Marinho et al. 2017)" authorityName="Marinho" authorityYear="2017" baseAuthorityName="Marinho et al." baseAuthorityYear="2017" box="[759,1078,986,1011]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F1DFFF2FD79B9FC377CFC40" box="[759,848,987,1011]" italics="true" pageId="39" pageNumber="40">Laneella</emphasis>
(
<bibRefCitation id="EFE24B010F1DFFF2FCD3B9FD3002FC40" author="Marinho, M. A. T. &amp; Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Azeredo-Espin, A. M. L. de &amp; Amorim, D." box="[861,1070,986,1011]" pageId="39" pageNumber="40" pagination="134 - 152" refId="ref108821" refString="Marinho, M. A. T., Wolff, M., Ramos-Pastrana, Y., Azeredo-Espin, A. M. L. de &amp; Amorim, D. d. S. (2017) The first phylogenetic study of Mesembrinellidae (Diptera: Oestroidea) based on molecular data: clades and congruence with morphological characters. Cladistics, 33 (2), 134 - 152. https: // doi. org / 10.1111 / cla. 1215 7" type="journal article" year="2017">
Marinho
<emphasis id="B907EAE20F1DFFF2FC32B9FB37C3FC40" box="[956,1007,987,1011]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
2017
</bibRefCitation>
)
</taxonomicName>
.]
</paragraph>
<paragraph id="8BCC36F00F1DFFF2FF19B9DD37E7FBA1" blockId="39.[151,1437,925,1073]" box="[151,971,1017,1042]" pageId="39" pageNumber="40">
<treatmentCitationGroup id="AB6311DE0F1DFFF2FF19B9DD37E7FBA1" box="[151,971,1017,1042]" pageId="39" pageNumber="40">
<taxonomicName id="4C734D730F1DFFF2FF19B9DD36B7FBA1" authority=": Wolff &amp; Kosmann (2016: 870)" authorityName="Wolff &amp; Kosmann" authorityPageNumber="870" authorityYear="2016" box="[151,667,1017,1042]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="patriciae">
<emphasis id="B907EAE20F1DFFF2FF19B9DD357DFBA2" box="[151,337,1018,1042]" italics="true" pageId="39" pageNumber="40">Laneella patriciae</emphasis>
:
<treatmentCitation id="0AD210E10F1DFFF2FED1B9DE36B7FBA1" author="Wolff, M. &amp; Kosmann, C." box="[351,667,1017,1042]" httpUri="http://treatment.plazi.org/id/039F87BCFF80C403FF377E08FB204C0E" page="870" pageId="39" pageNumber="40" year="2016">
<bibRefCitation id="EFE24B010F1DFFF2FED1B9DE36B7FBA1" author="Wolff, M. &amp; Kosmann, C." box="[351,667,1017,1042]" pageId="39" pageNumber="40" pagination="856 - 875" refId="ref110604" refString="Wolff, M. &amp; Kosmann, C. (2016) Families Calliphoridae and Mesembrinellidae. In: Wolff, M., Nihei, S. S. &amp; Carvalho, C. J. B. de (Eds.), Catalogue of Diptera of Colombia. Zootaxa, 4122 (1), 856 - 875. https: // doi. org / 10.11646 / zootaxa. 4122.1.7 2" type="journal article" year="2016">Wolff &amp; Kosmann (2016: 870)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<bibRefCitation id="EFE24B010F1DFFF2FD2BB9DE3757FBA1" author="Marinho, M. A. T. &amp; Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Azeredo-Espin, A. M. L. de &amp; Amorim, D." box="[677,891,1017,1042]" pageId="39" pageNumber="40" pagination="134 - 152" refId="ref108821" refString="Marinho, M. A. T., Wolff, M., Ramos-Pastrana, Y., Azeredo-Espin, A. M. L. de &amp; Amorim, D. d. S. (2017) The first phylogenetic study of Mesembrinellidae (Diptera: Oestroidea) based on molecular data: clades and congruence with morphological characters. Cladistics, 33 (2), 134 - 152. https: // doi. org / 10.1111 / cla. 1215 7" type="journal article" year="2017">
Marinho
<emphasis id="B907EAE20F1DFFF2FC8CB9DD3703FBA1" box="[770,815,1018,1042]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
. (2017
</bibRefCitation>
: tab. 1).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCC36F00F1DFFF2FF19BE3F3697FB83" blockId="39.[151,1437,925,1073]" box="[151,699,1048,1073]" pageId="39" pageNumber="40">
<taxonomicName id="4C734D730F1DFFF2FF19BE3F3646FB83" authority=": Cerretti et al. (2017" authorityName=": Cerretti et al." authorityYear="2017" box="[151,618,1048,1073]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="patriciae">
<emphasis id="B907EAE20F1DFFF2FF19BE3F35BCFB83" box="[151,400,1048,1072]" italics="true" pageId="39" pageNumber="40">Mesembrinella patriciae</emphasis>
:
<bibRefCitation id="EFE24B010F1DFFF2FE10BE3F3646FB83" author="Cerretti, P. &amp; Stireman, J. O. III. &amp; Pape, T. &amp; O'Hara, J. E. &amp; Marinho, M. A. T. &amp; Rognes, K. &amp; Grimaldi, D. A." box="[414,618,1048,1073]" pageId="39" pageNumber="40" pagination="0182101" refId="ref107732" refString="Cerretti, P., Stireman, J. O. III., Pape, T., O'Hara, J. E., Marinho, M. A. T., Rognes, K. &amp; Grimaldi, D. A. (2017) First fossil of an oestroid fly (Diptera: Calyptratae: Oestroidea) and the dating of oestroid divergences. PLoS ONE, 12 (8), e 0182101. https: // doi. org / 10.1371 / journal. pone. 018210 1" type="journal article" year="2017">
Cerretti
<emphasis id="B907EAE20F1DFFF2FE7CBE3E3609FB83" box="[498,549,1048,1072]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
(2017
</bibRefCitation>
</taxonomicName>
: tab. 2).
</paragraph>
</subSubSection>
<subSubSection id="C369657B0F1DFFF2FF19BE7935EBF822" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8BCC36F00F1DFFF2FF19BE793118FB56" blockId="39.[151,1437,1118,1937]" pageId="39" pageNumber="40">
<emphasis id="B907EAE20F1DFFF2FF19BE79353EFBCA" bold="true" box="[151,274,1118,1145]" pageId="39" pageNumber="40">Diagnosis.</emphasis>
A medium-sized fly averaging
<quantity id="4C8B9B150F1DFFF2FDF3BE7836F9FBCA" box="[637,725,1119,1145]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="39" pageNumber="40" unit="mm" value="9.4">9.4 mm</quantity>
(9–10/5) in length. Abdomen bluish to purple-green vs. shiny brown in
<taxonomicName id="4C734D730F1DFFF2FE8EBEA335BBFB2E" authority="Whitworth &amp; Yusseff-Vanegas, 2019" authorityName="Whitworth &amp; Yusseff-Vanegas" authorityYear="2019" box="[256,407,1155,1181]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="fusconitida" status="sp. nov.">
<emphasis id="B907EAE20F1DFFF2FE8EBEA335BBFB2E" box="[256,407,1155,1181]" italics="true" pageId="39" pageNumber="40">L. fusconitida</emphasis>
</taxonomicName>
<emphasis id="B907EAE20F1DFFF2FE13BEA435D9FB2E" bold="true" box="[413,501,1155,1181]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="A23457990F1DFFF2FE13BEA435D9FB2E" box="[413,501,1155,1181]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; mid and hind femora with apices dark brown in
<taxonomicName id="4C734D730F1DFFF2FB9CBEA330BEFB2E" authorityName=": Cerretti et al." authorityYear="2017" box="[1042,1170,1156,1181]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="patriciae">
<emphasis id="B907EAE20F1DFFF2FB9CBEA330BEFB2E" box="[1042,1170,1156,1181]" italics="true" pageId="39" pageNumber="40">L. patriciae</emphasis>
</taxonomicName>
vs. entirely orange in
<taxonomicName id="4C734D730F1DFFF2FA08BEA3353CFB72" authorityName="Whitworth &amp; Yusseff-Vanegas" authorityYear="2019" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="fusconitida">
<emphasis id="B907EAE20F1DFFF2FA08BEA3353CFB72" italics="true" pageId="39" pageNumber="40">L. fusconitida</emphasis>
</taxonomicName>
; male with surstylus straight in
<taxonomicName id="4C734D730F1DFFF2FDE9BE8F36CAFB72" authorityName=": Cerretti et al." authorityYear="2017" box="[615,742,1192,1217]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="patriciae">
<emphasis id="B907EAE20F1DFFF2FDE9BE8F36CAFB72" box="[615,742,1192,1217]" italics="true" pageId="39" pageNumber="40">L. patriciae</emphasis>
</taxonomicName>
vs. curved forward in
<taxonomicName id="4C734D730F1DFFF2FC59BE8F3047FB72" authorityName="Whitworth &amp; Yusseff-Vanegas" authorityYear="2019" box="[983,1131,1191,1217]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="fusconitida">
<emphasis id="B907EAE20F1DFFF2FC59BE8F3047FB72" box="[983,1131,1191,1217]" italics="true" pageId="39" pageNumber="40">L. fusconitida</emphasis>
</taxonomicName>
. This species is very similar to
<taxonomicName id="4C734D730F1DFFF2FF3DBEEB3565FB56" authorityName="Whitworth &amp; Yusseff-Vanegas" authorityYear="2019" box="[179,329,1227,1253]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="fusconitida">
<emphasis id="B907EAE20F1DFFF2FF3DBEEB3565FB56" box="[179,329,1227,1253]" italics="true" pageId="39" pageNumber="40">L. fusconitida</emphasis>
</taxonomicName>
, both sharing the distinctive tuberform spermathecae and brown spot around crossvein r-m.
</paragraph>
<paragraph id="8BCC36F00F1DFFF2FF49BEC937F4F926" blockId="39.[151,1437,1118,1937]" pageId="39" pageNumber="40">
<emphasis id="B907EAE20F1DFFF2FF49BEC93541FABA" bold="true" box="[199,365,1262,1289]" pageId="39" pageNumber="40">Redescription</emphasis>
. [Summarized from Wolff (2013).]
<emphasis id="B907EAE20F1DFFF2FD70BEC83714FABA" box="[766,824,1263,1289]" italics="true" pageId="39" pageNumber="40">Male</emphasis>
. Head. Frons narrow (holoptic); fronto-orbital, parafacial and gena yellow with silvery tomentum; frontal vitta reddish-brown, obliterated above; postgena and occiput with white tomentum and fine long pale setae; antenna orange and palpus typical; postocular setae extending to gena. Dorsum of thorax reddish-brown with white tomentum and testaceous stripes; ppn and pleura orange with fine white tomentum; spiracles pale yellow. Coxae, trochanters and femora orange; mid and hind femora with dark reddish-brown apices; chaetotaxy: ac 2:3, dc 2:3, ia 1:2, kat 2:1, ppn
<date id="FFCD10300F1DFFF2FCE2BF8437A6FA0E" box="[876,906,1443,1469]" pageId="39" pageNumber="40">3x</date>
3, meral setae in form of inverted L; ap scutellar setae 1:1. Wing hyaline with brown spot around crossvein r-m; subcosta bare; section IV about 0.30 of III (estimated from Wolff 2013: fig. 1); calypters with dark rims. Abdomen with dorsum bluish to greenish-purple except T1+2 yellow; T1+2 and T3 with lateral marginal setae; T4 and T5 with rows of marginal setae; disc of T5 without row of stout setae, whole segment with short fine setae. T6 partially fused with T7+8; pregonite apically truncated; tip of paraphallus with dorsal line of denticles; hypophallus shorter than paraphallus; epiphallus long and thin; cercus and surstylus slightly curved backward in lateral view (see Wolff 2013: fig. 2).
</paragraph>
<paragraph id="8BCC36F00F1DFFF2FF49BCB8372BF90A" blockId="39.[151,1437,1118,1937]" box="[199,775,1695,1721]" pageId="39" pageNumber="40">
<emphasis id="B907EAE20F1DFFF2FF49BCB83537F90A" box="[199,283,1695,1721]" italics="true" pageId="39" pageNumber="40">Female</emphasis>
. Dichoptic, frons broad and reddish-brown.
</paragraph>
<paragraph id="8BCC36F00F1DFFF2FF49BCE53018F96D" blockId="39.[151,1437,1118,1937]" box="[199,1076,1730,1758]" pageId="39" pageNumber="40">
<emphasis id="B907EAE20F1DFFF2FF49BCE5359DF96E" bold="true" box="[199,433,1730,1757]" pageId="39" pageNumber="40">Material examined.</emphasis>
None (no specimens could be obtained for examination).
</paragraph>
<paragraph id="8BCC36F00F1DFFF2FF49BCC135F2F8B2" blockId="39.[151,1437,1118,1937]" box="[199,478,1766,1793]" pageId="39" pageNumber="40">
<emphasis id="B907EAE20F1DFFF2FF49BCC1354DF8B2" bold="true" box="[199,353,1766,1793]" pageId="39" pageNumber="40">Distribution.</emphasis>
<collectingCountry id="F36476600F1DFFF2FEE9BCC035F5F8B2" box="[359,473,1767,1793]" name="Colombia" pageId="39" pageNumber="40">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="8BCC36F00F1DFFF2FF49BD2C35EBF822" blockId="39.[151,1437,1118,1937]" pageId="39" pageNumber="40">
<emphasis id="B907EAE20F1DFFF2FF49BD2C3517F896" bold="true" box="[199,315,1803,1829]" pageId="39" pageNumber="40">Remarks.</emphasis>
A single sequence of this species from GenBank (
<accessionNumber id="9420AB130F1DFFF2FCFEBD2C37C3F896" box="[880,1007,1803,1829]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KR820718" pageId="39" pageNumber="40">KR820718</accessionNumber>
) was added to our molecular analysis. It was distinct from other species and was recovered close to
<taxonomicName id="4C734D730F1DFFF2FCCFBD1737F6F8FA" authority="Whitworth &amp; Yusseff-Vanegas, 2019" authorityName="Whitworth &amp; Yusseff-Vanegas" authorityYear="2019" box="[833,986,1839,1865]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="fusconitida" status="sp. nov.">
<emphasis id="B907EAE20F1DFFF2FCCFBD1737F6F8FA" box="[833,986,1839,1865]" italics="true" pageId="39" pageNumber="40">L. fusconitida</emphasis>
</taxonomicName>
<emphasis id="B907EAE20F1DFFF2FC6FBD083016F8FA" bold="true" box="[993,1082,1839,1865]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="A23457990F1DFFF2FC6FBD083016F8FA" box="[993,1082,1839,1865]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13482A750F1DFFF2FBC7BD083085F8FA" box="[1097,1193,1839,1865]" captionStart="FIGURE 488" captionStartId="104.[151,250,1948,1973]" captionTargetBox="[179,1381,165,1919]" captionTargetId="figure@104.[179,1409,165,1919]" captionTargetPageId="104" captionText="FIGURE 488. Neighbor-joining tree of COI DNA barcode sequences of specimens and species of Mesembrinellidae, using K2P distances and generated in BOLD; species names on the tree are followed by BOLD accession number, voucher number, country of origin, number of sequenced base pairs and number of ambiguous nucleotides." httpUri="https://zenodo.org/record/3377367/files/figure.png" pageId="39" pageNumber="40">Fig. 488</figureCitation>
) with a genetic divergence of 3.84%, which supports them as two separate species; this is supported also by the several morphological differences as listed above.
</paragraph>
</subSubSection>
</treatment>
</document>
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