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<mods:title id="1F7BAD56E2F20B0836F6158AF2ED79AC">A revision of the genera and species of the Neotropical family Mesembrinellidae (Diptera: Oestroidea)</mods:title>
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<heading id="D084819C0F00FFEFFF19BFD637D8F9B8" bold="true" box="[151,1012,1521,1548]" fontSize="11" level="1" pageId="58" pageNumber="59" reason="1">
<taxonomicName id="4C734D730F00FFEFFF19BFD637D8F9B8" authority="(Wolff, Ramos-Pastrana &amp; Pujol-Luz, 2013)" baseAuthorityName="Wolff, Ramos-Pastrana &amp; Pujol-Luz" baseAuthorityYear="2013" box="[151,1012,1521,1548]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="carvalhoi">
<emphasis id="B907EAE20F00FFEFFF19BFD637D8F9B8" bold="true" box="[151,1012,1521,1548]" pageId="58" pageNumber="59">
<emphasis id="B907EAE20F00FFEFFF19BFD635E9F9BF" bold="true" box="[151,453,1521,1548]" italics="true" pageId="58" pageNumber="59">Mesembrinella carvalhoi</emphasis>
(Wolff, Ramos-Pastrana &amp; Pujol-Luz, 2013)
</emphasis>
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<treatmentCitationGroup id="AB6311DE0F00FFEFFF19BC123539F9DF" pageId="58" pageNumber="59">
<taxonomicName id="4C734D730F00FFEFFF19BC12365DF9FE" authority="Wolff et al., 2013 b: 130" authorityName="Wolff et al." authorityPageNumber="130" authorityYear="2013" box="[151,625,1588,1613]" class="Insecta" family="Calliphoridae" genus="Giovanella" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="carvalhoi">
<emphasis id="B907EAE20F00FFEFFF19BC12355CF9FE" box="[151,368,1589,1613]" italics="true" pageId="58" pageNumber="59">Giovanella carvalhoi</emphasis>
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<treatmentCitation id="0AD210E10F00FFEFFEF9BC13365DF9FE" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R." box="[375,625,1588,1613]" page="130" pageId="58" pageNumber="59" year="2013">
<bibRefCitation id="EFE24B010F00FFEFFEF9BC13365DF9FE" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R." box="[375,625,1588,1613]" pageId="58" pageNumber="59" pagination="129 - 132" refId="ref110824" refString="Wolff, M., Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R. (2013 b) A new species of Giovanella Bonatto (Diptera, Calliphoridae, Mesembrinellinae) from Colombia. Revista Brasileira de Entomologia, 57 (2), 129 - 132. https: // doi. org / 10.1590 / S 0085 - 5626201300500001 3" type="journal article" year="2013">
Wolff
<emphasis id="B907EAE20F00FFEFFE37BC1135C1F9FE" box="[441,493,1589,1613]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
, 2013b: 130
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.
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male (CEUA), not examined. Type locality:
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,
<collectingRegion id="49B7F8120F00FFEFFA98BC123140F9FE" box="[1302,1388,1589,1613]" country="Colombia" name="Caqueta" pageId="58" pageNumber="59">Caquetá</collectingRegion>
,
<collectingCountry id="F36476600F00FFEFFAF8BC12353DF9DF" name="Colombia" pageId="58" pageNumber="59">Colombia</collectingCountry>
.
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<treatmentCitationGroup id="AB6311DE0F00FFEFFF19BC5537C5F938" box="[151,1001,1650,1675]" pageId="58" pageNumber="59">
<taxonomicName id="4C734D730F00FFEFFF19BC553694F938" authority=": Wolff &amp; Kosmann (2016: 866)" authorityName="Wolff &amp; Kosmann" authorityPageNumber="866" authorityYear="2016" box="[151,696,1650,1675]" class="Insecta" family="Calliphoridae" genus="Giovanella" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="carvalhoi">
<emphasis id="B907EAE20F00FFEFFF19BC553543F939" box="[151,367,1650,1674]" italics="true" pageId="58" pageNumber="59">Giovanella carvalhoi</emphasis>
:
<treatmentCitation id="0AD210E10F00FFEFFEF2BC553694F938" ID-DOI="http://doi.org/10.5281/zenodo.6083259" ID-Zenodo-Dep="6083259" author="Wolff, M. &amp; Kosmann, C." box="[380,696,1650,1675]" httpUri="http://treatment.plazi.org/id/039F87BCFF81C402FF377FFDFB4E4CC7" page="866" pageId="58" pageNumber="59" year="2016">
<bibRefCitation id="EFE24B010F00FFEFFEF2BC553694F938" author="Wolff, M. &amp; Kosmann, C." box="[380,696,1650,1675]" pageId="58" pageNumber="59" pagination="856 - 875" refId="ref110604" refString="Wolff, M. &amp; Kosmann, C. (2016) Families Calliphoridae and Mesembrinellidae. In: Wolff, M., Nihei, S. S. &amp; Carvalho, C. J. B. de (Eds.), Catalogue of Diptera of Colombia. Zootaxa, 4122 (1), 856 - 875. https: // doi. org / 10.11646 / zootaxa. 4122.1.7 2" type="journal article" year="2016">Wolff &amp; Kosmann (2016: 866)</bibRefCitation>
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;
<bibRefCitation id="EFE24B010F00FFEFFD4CBC5537B4F938" author="Marinho, M. A. T. &amp; Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Azeredo-Espin, A. M. L. de &amp; Amorim, D." box="[706,920,1650,1675]" pageId="58" pageNumber="59" pagination="134 - 152" refId="ref108821" refString="Marinho, M. A. T., Wolff, M., Ramos-Pastrana, Y., Azeredo-Espin, A. M. L. de &amp; Amorim, D. d. S. (2017) The first phylogenetic study of Mesembrinellidae (Diptera: Oestroidea) based on molecular data: clades and congruence with morphological characters. Cladistics, 33 (2), 134 - 152. https: // doi. org / 10.1111 / cla. 1215 7" type="journal article" year="2017">
Marinho
<emphasis id="B907EAE20F00FFEFFCAEBC54377FF939" box="[800,851,1650,1674]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
(2017
</bibRefCitation>
: tab. 1).
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</paragraph>
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<treatmentCitationGroup id="AB6311DE0F00FFEFFF19BCB5374FF91A" box="[151,867,1680,1705]" pageId="58" pageNumber="59">
<taxonomicName id="4C734D730F00FFEFFF19BCB5368EF91A" authority=": Wolff et al. (2017: 253)" authorityName="Wolff" authorityPageNumber="253" authorityYear="2017" box="[151,674,1680,1705]" class="Insecta" family="Calliphoridae" genus="Huascaromusca" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="carvalhoi" status="comb. nov.">
<emphasis id="B907EAE20F00FFEFFF19BCB5358EF91A" box="[151,418,1681,1705]" italics="true" pageId="58" pageNumber="59">Huascaromusca carvalhoi</emphasis>
:
<treatmentCitation id="0AD210E10F00FFEFFE21BCB7368EF91A" ID-DOI="http://doi.org/10.5281/zenodo.6039417" ID-Zenodo-Dep="6039417" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Marinho, M. A. T. &amp; Amorim, D." box="[431,674,1680,1705]" httpUri="http://treatment.plazi.org/id/03F06820FFB4FFA9F8F631BEFE57FE90" page="253" pageId="58" pageNumber="59" year="2017">
<bibRefCitation id="EFE24B010F00FFEFFE21BCB7368EF91A" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Marinho, M. A. T. &amp; Amorim, D." box="[431,674,1680,1705]" pageId="58" pageNumber="59" pagination="251 - 263" refId="ref110679" refString="Wolff, M., Ramos-Pastrana, Y., Marinho, M. A. T. &amp; Amorim, D. d. S. (2017) Two new species of Huascaromusca Townsend from Colombia, and a new combination for Giovanella carvalhoi (Diptera: Mesembrinellidae). Zootaxa, 4231 (2), 251 - 263. https: // doi. org / 10.11646 / zootaxa. 4231.2. 7" type="journal article" year="2017">
Wolff
<emphasis id="B907EAE20F00FFEFFE7EBCB5360FF91A" box="[496,547,1681,1705]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
(2017: 253)
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</taxonomicName>
,
<taxonomicNameLabel id="A23457990F00FFEFFD22BCB53773F91A" box="[684,863,1680,1705]" pageId="58" pageNumber="59" rank="species">new combination</taxonomicNameLabel>
.
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</paragraph>
<paragraph id="8BCC36F00F00FFEFFF19BC9736ECF97B" blockId="58.[151,1436,1588,1736]" box="[151,704,1711,1736]" pageId="58" pageNumber="59">
<taxonomicName id="4C734D730F00FFEFFF19BC97365CF97B" authority=": Cerretti et al. (2017" authorityName=": Cerretti et al." authorityYear="2017" box="[151,624,1711,1736]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="carvalhoi">
<emphasis id="B907EAE20F00FFEFFF19BC9735BAF97B" box="[151,406,1712,1736]" italics="true" pageId="58" pageNumber="59">Mesembrinella carvalhoi</emphasis>
:
<bibRefCitation id="EFE24B010F00FFEFFE2ABC88365CF97B" author="Cerretti, P. &amp; Stireman, J. O. III. &amp; Pape, T. &amp; O'Hara, J. E. &amp; Marinho, M. A. T. &amp; Rognes, K. &amp; Grimaldi, D. A." box="[420,624,1711,1736]" pageId="58" pageNumber="59" pagination="0182101" refId="ref107732" refString="Cerretti, P., Stireman, J. O. III., Pape, T., O'Hara, J. E., Marinho, M. A. T., Rognes, K. &amp; Grimaldi, D. A. (2017) First fossil of an oestroid fly (Diptera: Calyptratae: Oestroidea) and the dating of oestroid divergences. PLoS ONE, 12 (8), e 0182101. https: // doi. org / 10.1371 / journal. pone. 018210 1" type="journal article" year="2017">
Cerretti
<emphasis id="B907EAE20F00FFEFFE76BC963607F97B" box="[504,555,1712,1736]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
(2017
</bibRefCitation>
</taxonomicName>
: tab. 2).
</paragraph>
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<subSubSection id="C369657B0F00FFEEFF19BCD23607FE1C" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="8BCC36F00F00FFEFFF19BCD236E9F8EB" blockId="58.[151,1437,1781,2025]" pageId="58" pageNumber="59">
<emphasis id="B907EAE20F00FFEFFF19BCD2353EF8A3" bold="true" box="[151,274,1781,1808]" pageId="58" pageNumber="59">Diagnosis.</emphasis>
A medium-sized dark blue fly with pale tomentum, T1+2 and anterior portion of T3 yellowish. Anterior spiracle yellow-orange, posterior spiracle dark brown; legs including femora entirely dark brown; epandrium, cerci and surstyli as in
<bibRefCitation id="EFE24B010F00FFEFFED5BD193619F8EB" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R." box="[347,565,1854,1880]" pageId="58" pageNumber="59" pagination="129 - 132" refId="ref110824" refString="Wolff, M., Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R. (2013 b) A new species of Giovanella Bonatto (Diptera, Calliphoridae, Mesembrinellinae) from Colombia. Revista Brasileira de Entomologia, 57 (2), 129 - 132. https: // doi. org / 10.1590 / S 0085 - 5626201300500001 3" type="journal article" year="2013">
Wolff
<emphasis id="B907EAE20F00FFEFFE2CBD1835F6F8EB" box="[418,474,1854,1880]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
(2013b
</bibRefCitation>
: figs 1011).
</paragraph>
<paragraph id="8BCC36F00F00FFEEFF49BD46371DFEAC" blockId="58.[151,1437,1781,2025]" lastBlockId="59.[151,1437,153,432]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
<emphasis id="B907EAE20F00FFEFFF49BD463559F8CF" bold="true" box="[199,373,1889,1916]" pageId="58" pageNumber="59">Redescription.</emphasis>
[Summarized from
<bibRefCitation id="EFE24B010F00FFEFFDD0BD453713F8CF" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R." box="[606,831,1890,1916]" pageId="58" pageNumber="59" pagination="129 - 132" refId="ref110824" refString="Wolff, M., Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R. (2013 b) A new species of Giovanella Bonatto (Diptera, Calliphoridae, Mesembrinellinae) from Colombia. Revista Brasileira de Entomologia, 57 (2), 129 - 132. https: // doi. org / 10.1590 / S 0085 - 5626201300500001 3" type="journal article" year="2013">
Wolff
<emphasis id="B907EAE20F00FFEFFD29BD4436CEF8CF" box="[679,738,1890,1916]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
(2013b
</bibRefCitation>
: figs 111); this species is similar to
<taxonomicName id="4C734D730F00FFEFFB61BD443144F8CF" authorityName=": Kosmann et al." authorityYear="2013" box="[1263,1384,1890,1916]" class="Insecta" family="Calliphoridae" genus="Giovanella" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="bolivar">
<emphasis id="B907EAE20F00FFEFFB61BD443144F8CF" box="[1263,1384,1890,1916]" italics="true" pageId="58" pageNumber="59">M. bolivar</emphasis>
</taxonomicName>
, see comparison under that species.] Thorax with black [blue-black according to figs
<quantity id="4C8B9B150F00FFEFFBA2BDA13059F813" box="[1068,1141,1926,1952]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="58" pageNumber="59" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">13 in</quantity>
<bibRefCitation id="EFE24B010F00FFEFFBF2BDA1314EF813" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R." box="[1148,1378,1925,1952]" pageId="58" pageNumber="59" pagination="129 - 132" refId="ref110824" refString="Wolff, M., Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R. (2013 b) A new species of Giovanella Bonatto (Diptera, Calliphoridae, Mesembrinellinae) from Colombia. Revista Brasileira de Entomologia, 57 (2), 129 - 132. https: // doi. org / 10.1590 / S 0085 - 5626201300500001 3" type="journal article" year="2013">
Wolff
<emphasis id="B907EAE20F00FFEFFB4BBDA030D2F813" box="[1221,1278,1926,1952]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
(2013b)
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] mesonotum and white tomentum with well-defined stripes. Chaetotaxy: ac 0:0, dc 1:2, ia 0:1, ph 1, ppn
<date id="FFCD10300F00FFEFFABABD8D317EF877" box="[1332,1362,1962,1988]" pageId="58" pageNumber="59">2x</date>
2, disc absent; anterior spiracle yellow-orange, posterior spiracle dark brown; T5 without discal setae; T4 with marginal setae. Wing mostly hyaline, veins faintly infuscated. Epandrium, cerci and surstyli in left lateral view with surstylus curved backward, cercus with apical hook (
<bibRefCitation id="EFE24B010F01FFEEFD0ABA9A3776FF64" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R." box="[644,858,189,215]" pageId="59" pageNumber="60" pagination="129 - 132" refId="ref110824" refString="Wolff, M., Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R. (2013 b) A new species of Giovanella Bonatto (Diptera, Calliphoridae, Mesembrinellinae) from Colombia. Revista Brasileira de Entomologia, 57 (2), 129 - 132. https: // doi. org / 10.1590 / S 0085 - 5626201300500001 3" type="journal article" year="2013">
Wolff
<emphasis id="B907EAE20F01FFEEFD40BA993724FF64" box="[718,776,189,215]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
2013b
</bibRefCitation>
: fig. 10); in posterior view, cerci broad at base and tapering to a point, with sinuous curve midway (
<bibRefCitation id="EFE24B010F01FFEEFD39BAC637A7FF48" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R." box="[695,907,225,251]" pageId="59" pageNumber="60" pagination="129 - 132" refId="ref110824" refString="Wolff, M., Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R. (2013 b) A new species of Giovanella Bonatto (Diptera, Calliphoridae, Mesembrinellinae) from Colombia. Revista Brasileira de Entomologia, 57 (2), 129 - 132. https: // doi. org / 10.1590 / S 0085 - 5626201300500001 3" type="journal article" year="2013">
Wolff
<emphasis id="B907EAE20F01FFEEFC8FBAC53716FF48" box="[769,826,225,251]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
2013b
</bibRefCitation>
: fig. 11). Phallus in lateral with moderate sized epiphallus curving backward (
<bibRefCitation id="EFE24B010F01FFEEFE67BB223690FEAC" author="Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R." box="[489,700,261,287]" pageId="59" pageNumber="60" pagination="129 - 132" refId="ref110824" refString="Wolff, M., Ramos-Pastrana, Y. &amp; Pujol-Luz, J. R. (2013 b) A new species of Giovanella Bonatto (Diptera, Calliphoridae, Mesembrinellinae) from Colombia. Revista Brasileira de Entomologia, 57 (2), 129 - 132. https: // doi. org / 10.1590 / S 0085 - 5626201300500001 3" type="journal article" year="2013">
Wolff
<emphasis id="B907EAE20F01FFEEFDBCBB213646FEAC" box="[562,618,261,287]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
2013b
</bibRefCitation>
: figs 89).
</paragraph>
<paragraph id="8BCC36F00F01FFEEFF49BB0F3018FEF7" blockId="59.[151,1437,153,432]" box="[199,1076,296,324]" pageId="59" pageNumber="60">
<emphasis id="B907EAE20F01FFEEFF49BB0F359DFEF0" bold="true" box="[199,433,296,323]" pageId="59" pageNumber="60">Material examined.</emphasis>
None (no specimens could be obtained for examination).
</paragraph>
<paragraph id="8BCC36F00F01FFEEFF49BB6B35F2FED4" blockId="59.[151,1437,153,432]" box="[199,478,332,359]" pageId="59" pageNumber="60">
<emphasis id="B907EAE20F01FFEEFF49BB6B354DFED4" bold="true" box="[199,353,332,359]" pageId="59" pageNumber="60">Distribution.</emphasis>
<collectingCountry id="F36476600F01FFEEFEE9BB6A35F5FED4" box="[359,473,333,359]" name="Colombia" pageId="59" pageNumber="60">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="8BCC36F00F01FFEEFF49BB563607FE1C" blockId="59.[151,1437,153,432]" pageId="59" pageNumber="60">
<emphasis id="B907EAE20F01FFEEFF49BB563517FE38" bold="true" box="[199,315,369,395]" pageId="59" pageNumber="60">Remarks.</emphasis>
Morphologically, this species appears to be closely related to
<taxonomicName id="4C734D730F01FFEEFC7BBB553047FE38" authorityName=": Kosmann et al." authorityYear="2013" box="[1013,1131,369,395]" class="Insecta" family="Calliphoridae" genus="Giovanella" kingdom="Animalia" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="bolivar">
<emphasis id="B907EAE20F01FFEEFC7BBB553047FE38" box="[1013,1131,369,395]" italics="true" pageId="59" pageNumber="60">M. bolivar</emphasis>
</taxonomicName>
(see Diagnosis of that species). No specimens were barcoded.
</paragraph>
</subSubSection>
</treatment>
</document>