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<document id="8D46F0DC16C7700A5A1115234B8658DA" ID-DOI="10.11646/zootaxa.3919.1.6" ID-GBIF-Dataset="4c40552e-e778-4d8e-a6b7-9662436cadc2" ID-ISSN="1175-5326" ID-Zenodo-Dep="244587" ID-ZooBank="F711CA99-1B2C-4E18-9F4B-7521D38D2303" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460989857372" checkinUser="plazi" docAuthor="Smith, Ian M., Cook, David R. &amp; Gerecke, Reinhard" docDate="2015" docId="03DA87C4FF8E1761FF4C6AF9AE98F99D" docLanguage="en" docName="zt03919p156.pdf" docOrigin="Zootaxa 3919 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Madawaska Habeeb 1954" docType="treatment" docVersion="9" lastPageNumber="117" masterDocId="FFE3FFBCFF881767FFDB6A61AA1CFF8D" masterDocTitle="Revision of the status of some genus-level water mite taxa in the families Pionidae Thor, 1900, Aturidae Thor, 1900, and Nudomideopsidae Smith, 1990 (Acari: Hydrachnidiae)" masterLastPageNumber="156" masterPageNumber="111" pageNumber="117" updateTime="1698561557693" updateUser="plazi">
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<mods:title id="5A4894A65D98DE0CB907F3FA0EAD22C8">Revision of the status of some genus-level water mite taxa in the families Pionidae Thor, 1900, Aturidae Thor, 1900, and Nudomideopsidae Smith, 1990 (Acari: Hydrachnidiae)</mods:title>
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<mods:namePart id="77680394019B4411F8C6089E3057B4FA">Smith, Ian M.</mods:namePart>
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<treatment id="03DA87C4FF8E1761FF4C6AF9AE98F99D" ID-DOI="http://doi.org/10.5281/zenodo.6109824" ID-GBIF-Taxon="119596547" ID-Zenodo-Dep="6109824" LSID="urn:lsid:plazi:treatment:03DA87C4FF8E1761FF4C6AF9AE98F99D" httpUri="http://treatment.plazi.org/id/03DA87C4FF8E1761FF4C6AF9AE98F99D" lastPageNumber="117" pageId="6" pageNumber="117">
<subSubSection id="C3696559FF8E1761FF4C6AF9A8BEFF3F" box="[151,674,152,178]" pageId="6" pageNumber="117" type="nomenclature">
<paragraph id="8BCC36D2FF8E1761FF4C6AF9A8BEFF3F" blockId="6.[151,674,152,178]" box="[151,674,152,178]" pageId="6" pageNumber="117">
<heading id="D08481BEFF8E1761FF4C6AF9A8BEFF3F" bold="true" box="[151,674,152,178]" fontSize="11" level="1" pageId="6" pageNumber="117" reason="1">
<emphasis id="B907EAC0FF8E1761FF4C6AF9A8BEFF3F" bold="true" box="[151,674,152,178]" pageId="6" pageNumber="117">
Genus
<taxonomicName id="4C734D51FF8E1761FF356AF9A82CFF3F" authority="Habeeb, 1954" authorityName="Habeeb" authorityYear="1954" box="[238,560,152,178]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FF356AF9AB63FF3F" bold="true" box="[238,383,152,178]" italics="true" pageId="6" pageNumber="117">Madawaska</emphasis>
Habeeb, 1954
</taxonomicName>
<emphasis id="B907EAC0FF8E1761FDEC6AF9A8BEFF3F" bold="true" box="[567,674,152,178]" italics="true" pageId="6" pageNumber="117">stat. nov.</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8E1761FF4C6ABFA8DDFEE2" pageId="6" pageNumber="117" type="reference_group">
<paragraph id="8BCC36D2FF8E1761FF4C6ABFA8C3FEBF" blockId="6.[151,838,222,367]" pageId="6" pageNumber="117">
<taxonomicName id="4C734D51FF8E1761FF4C6ABFAB6FFF79" ID-CoL="4KRJ" authority="(Madawaska)" baseAuthorityName="Madawaska" box="[151,371,222,244]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FF4C6ABFAAFEFF79" box="[151,226,222,244]" italics="true" pageId="6" pageNumber="117">Forelia</emphasis>
(
<emphasis id="B907EAC0FF8E1761FF2A6ABFAB6FFF79" box="[241,371,222,244]" italics="true" pageId="6" pageNumber="117">Madawaska)</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFE24B23FF8E1761FE5A6ABFA819FF79" author="Cook" box="[385,517,222,244]" pageId="6" pageNumber="117" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook, 1974a</bibRefCitation>
, pp. 287288, figs.
<date id="FFCD1012FF8E1761FD176ABFA95DFF79" box="[716,833,222,244]" pageId="6" pageNumber="117" value="1214" valueMax="1218">12141218</date>
.
<taxonomicName id="4C734D51FF8E1761FF4C6A9CAAFEFE9E" ID-CoL="4KRJ" box="[151,226,253,275]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FF4C6A9CAAFEFE9E" box="[151,226,253,275]" italics="true" pageId="6" pageNumber="117">Forelia</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8E1761FE936A9CABDAFE9E" author="Smith" box="[328,454,253,275]" pageId="6" pageNumber="117" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith, 1976</bibRefCitation>
, pp. 2829, 79, 81, 92, figs. 21, 22.
<taxonomicName id="4C734D51FF8E1761FF4C6B7DAAFEFEBF" ID-CoL="4KRJ" box="[151,226,284,306]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FF4C6B7DAAFEFEBF" box="[151,226,284,306]" italics="true" pageId="6" pageNumber="117">Forelia</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8E1761FE9C6B7DA801FEBF" author="Smith" box="[327,541,284,306]" pageId="6" pageNumber="117" refString="Smith, I. M. &amp; Cook, D. R. (1991) Water mites. Chapter 16. In: Thorp, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. Academic Press, San Diego, CA, pp. 523 - 592. [total page number 911 pp.]" type="book chapter" year="1991">Smith &amp; Cook, 1991</bibRefCitation>
, pp. 554, 576, 578.
</paragraph>
<paragraph id="8BCC36D2FF8E1761FF4C6B5BA8DDFEDD" blockId="6.[151,838,222,367]" box="[151,705,314,336]" pageId="6" pageNumber="117">
<taxonomicName id="4C734D51FF8E1761FF4C6B5BAAFEFEDD" ID-CoL="4KRJ" box="[151,226,314,336]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FF4C6B5BAAFEFEDD" box="[151,226,314,336]" italics="true" pageId="6" pageNumber="117">Forelia</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8E1761FE9C6B5BABE3FEDD" author="Smith" box="[327,511,314,336]" pageId="6" pageNumber="117" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2001) Water mites and other arachnids. Chapter 16. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 2 nd Edition. Academic Press, Waltham, Massachusetts, pp. 551 - 659. [1056 pp.] http: // dx. doi. org / 10.1016 / B 978 - 012690647 - 9 / 50017 - X" type="book chapter" year="2001">
Smith
<emphasis id="B907EAC0FF8E1761FE506B5AABABFEDD" box="[395,439,314,336]" italics="true" pageId="6" pageNumber="117">et al</emphasis>
., 2001
</bibRefCitation>
, pp. 581, 612, 615.
</paragraph>
<paragraph id="8BCC36D2FF8E1761FF4C6B38A8DDFEE2" blockId="6.[151,838,222,367]" box="[151,705,345,367]" pageId="6" pageNumber="117">
<taxonomicName id="4C734D51FF8E1761FF4C6B38AAFEFEE2" ID-CoL="4KRJ" box="[151,226,345,367]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FF4C6B38AAFEFEE2" box="[151,226,345,367]" italics="true" pageId="6" pageNumber="117">Forelia</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8E1761FE9C6B38ABE3FEE2" author="Smith" box="[327,511,345,367]" pageId="6" pageNumber="117" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2010) Water mites and other arachnids. Chapter 15. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 3 rd Edition. Academic Press, Waltham, Massachusetts, pp. 485 - 586. [total page number: 1021 pp.] http: // dx. doi. org / 10.1016 / B 978 - 0 - 12 - 374855 - 3.00015 - 7" type="book chapter" year="2010">
Smith
<emphasis id="B907EAC0FF8E1761FE506B3BABABFEE2" box="[395,439,345,367]" italics="true" pageId="6" pageNumber="117">et al</emphasis>
., 2010
</bibRefCitation>
, pp. 524, 553, 556.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8E1761FF4C6BFDAAC2FBB0" pageId="6" pageNumber="117" type="diagnosis">
<paragraph id="8BCC36D2FF8E1761FF4C6BFDA919FDC9" blockId="6.[151,1437,412,1553]" pageId="6" pageNumber="117">
<emphasis id="B907EAC0FF8E1761FF4C6BFDAB7FFE3E" bold="true" box="[151,355,412,437]" pageId="6" pageNumber="117">
Diagnosis.
<emphasis id="B907EAC0FF8E1761FEC46BFDAB7FFE3E" bold="true" box="[287,355,412,435]" italics="true" pageId="6" pageNumber="117">Larva</emphasis>
</emphasis>
(modified from
<bibRefCitation id="EFE24B23FF8E1761FDEF6BFDA8A2FE38" author="Smith" box="[564,702,412,437]" pageId="6" pageNumber="117" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith 1976</bibRefCitation>
): Character states of the subfamily Foreliinae. Lateral coxal apodeme nearly transverse. Third coxal plate with medial coxal apodeme present and transverse muscle attachment scar very small or absent. Excretory pore plate much larger than excretory pore and roughly circular in shape with setae
<emphasis id="B907EAC0FF8E1761FF0D6869AAE1FDAD" box="[214,253,520,544]" italics="true" pageId="6" pageNumber="117">ps1</emphasis>
slightly anterior to setae
<emphasis id="B907EAC0FF8E1761FDC76869A85FFDAD" box="[540,579,520,544]" italics="true" pageId="6" pageNumber="117">ps2</emphasis>
. Numbers of setae on leg segments as follows: IITi 11 (setae Ti12 and Ti13 present); IIITr 1; IIITi 11 (setae Ti12 and Ti13 present).
</paragraph>
<paragraph id="8BCC36D2FF8E1761FF1C682EAAC2FBB0" blockId="6.[151,1437,412,1553]" pageId="6" pageNumber="117">
<emphasis id="B907EAC0FF8E1761FF1C682EAB0EFDE5" bold="true" box="[199,274,591,616]" italics="true" pageId="6" pageNumber="117">Adults</emphasis>
(modified from
<bibRefCitation id="EFE24B23FF8E1761FE31682EA86BFDE5" author="Smith" box="[490,631,591,616]" pageId="6" pageNumber="117" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith 1976</bibRefCitation>
): Character states of subfamily Foreliinae. Idiosoma with dorsal integument mostly soft and bearing only tiny platelets associated with glandularia and idiosomal setae. Venter with coxal plates and genital field incorporated into a ventral shield in males, but surrounded by soft integument bearing only tiny sclerites associated with glandularia and idiosomal setae in females. First coxal group with apodemes elongated. Fourth coxal plate with medial edge short but not reduced to angle in males, reduced to highly acute angle in females. Genital field with five to seven relatively large acetabula per side borne on tongueshaped acetabular plates that surround gonopore in males and flank gonopore in females, and that extend laterally to but not beyond posterolateral angles of fourth coxal plates. Pedipalp segments relatively short and stocky, with tibia bearing two slender setae on a small projection ventrally and a sessile, peg-like seta distomedially. Male third leg with tarsus bearing reduced claw sockets that are terminal in position and small but only slightly modified claws; fourth leg with genu short, slightly expanded, and bearing patches of thick, blade-like setae both at the base of and on a prominent distoventral projection, tibia relatively long and slender, tarsus curved dorsally, bearing a patch of thick, peg-like setae only in region proximal to deepest part of curved portion, and bearing thick, highly modified claws.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8E1761FF1C6E26AE8EFB08" pageId="6" pageNumber="117" type="materials_examined">
<paragraph id="8BCC36D2FF8E1761FF1C6E26A8AAFBED" blockId="6.[151,1437,412,1553]" box="[199,694,1095,1120]" pageId="6" pageNumber="117">
<emphasis id="B907EAC0FF8E1761FF1C6E26AB7EFBED" bold="true" box="[199,354,1095,1120]" pageId="6" pageNumber="117">
<typeStatus id="54C88870FF8E1761FF1C6E26AB1EFBED" box="[199,258,1095,1120]" pageId="6" pageNumber="117">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C734D51FF8E1761FEB26E26A8AAFBED" authority="Habeeb." authorityName="Habeeb." box="[361,694,1095,1120]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B907EAC0FF8E1761FEB26E26A84EFBED" box="[361,594,1095,1120]" italics="true" pageId="6" pageNumber="117">Madawaska borealis</emphasis>
Habeeb.
</taxonomicName>
</paragraph>
<paragraph id="8BCC36D2FF8E1761FF1C6E0DAE8EFB08" blockId="6.[151,1437,412,1553]" box="[199,1170,1132,1157]" pageId="6" pageNumber="117">
<emphasis id="B907EAC0FF8E1761FF1C6E0DAB8FFB08" bold="true" box="[199,403,1132,1157]" pageId="6" pageNumber="117">Species included.</emphasis>
<taxonomicName id="4C734D51FF8E1761FE416E0DA8C3FB08" authority="Habeeb" authorityName="Habeeb" box="[410,735,1132,1157]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B907EAC0FF8E1761FE416E0DA89EFB08" box="[410,642,1132,1157]" italics="true" pageId="6" pageNumber="117">Madawaska borealis</emphasis>
Habeeb
</taxonomicName>
(eastern North
<collectingCountry id="F3647642FF8E1761FC556E0DA9ECFB08" box="[910,1008,1132,1157]" name="United States of America" pageId="6" pageNumber="117">America</collectingCountry>
) (monobasic).
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8E1761FF1C6EEEA86EFB25" box="[199,626,1167,1192]" pageId="6" pageNumber="117" type="distribution">
<paragraph id="8BCC36D2FF8E1761FF1C6EEEA86EFB25" blockId="6.[151,1437,412,1553]" box="[199,626,1167,1192]" pageId="6" pageNumber="117">
<emphasis id="B907EAC0FF8E1761FF1C6EEEAB7CFB25" bold="true" box="[199,352,1167,1192]" pageId="6" pageNumber="117">Distribution.</emphasis>
Eastern North
<collectingCountry id="F3647642FF8E1761FDD16EEEA872FB25" box="[522,622,1167,1192]" name="United States of America" pageId="6" pageNumber="117">America</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8E1761FF1C6ED5AE98F99D" pageId="6" pageNumber="117" type="discussion">
<paragraph id="8BCC36D2FF8E1761FF1C6ED5AE98F99D" blockId="6.[151,1437,412,1553]" pageId="6" pageNumber="117">
<emphasis id="B907EAC0FF8E1761FF1C6ED5AB50FB40" bold="true" box="[199,332,1204,1229]" pageId="6" pageNumber="117">Discussion.</emphasis>
<bibRefCitation id="EFE24B23FF8E1761FE886ED5ABF2FB41" author="Cook" box="[339,494,1204,1229]" pageId="6" pageNumber="117" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook (1974a)</bibRefCitation>
treated
<taxonomicName id="4C734D51FF8E1761FD926ED5A8CDFB40" box="[585,721,1204,1229]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FD926ED5A8CDFB40" box="[585,721,1204,1229]" italics="true" pageId="6" pageNumber="117">Madawaska</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName id="4C734D51FF8E1761FC436ED5A9F7FB40" box="[920,1003,1204,1229]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FC436ED5A9F7FB40" box="[920,1003,1204,1229]" italics="true" pageId="6" pageNumber="117">Forelia</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE24B23FF8E1761FC226ED5AE92FB40" author="Smith" box="[1017,1166,1204,1229]" pageId="6" pageNumber="117" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith (1976)</bibRefCitation>
recognized the
<taxonomicName id="4C734D51FF8E1761FA9A6ED5AF87FB40" box="[1345,1435,1204,1229]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B907EAC0FF8E1761FA9A6ED5AF87FB40" box="[1345,1435,1204,1229]" italics="true" pageId="6" pageNumber="117">borealis</emphasis>
</taxonomicName>
group as a distinct clade, but at that time questioned the merit of retaining
<taxonomicName id="4C734D51FF8E1761FC3B6EB6AE7BFB7D" box="[992,1127,1239,1264]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FC3B6EB6AE7BFB7D" box="[992,1127,1239,1264]" italics="true" pageId="6" pageNumber="117">Madawaska</emphasis>
</taxonomicName>
as a subgenus of the genus
<taxonomicName id="4C734D51FF8E1761FF4C6E9DAAF7FA98" box="[151,235,1276,1301]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FF4C6E9DAAF7FA98" box="[151,235,1276,1301]" italics="true" pageId="6" pageNumber="117">Forelia</emphasis>
</taxonomicName>
noting that the nominate subgenus was apparently not monophyletic and in need of further revision. Here we propose elevation of
<taxonomicName id="4C734D51FF8E1761FE646F7EA85AFAB5" box="[447,582,1311,1336]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FE646F7EA85AFAB5" box="[447,582,1311,1336]" italics="true" pageId="6" pageNumber="117">Madawaska</emphasis>
</taxonomicName>
to full generic rank based upon the apomorphic condition of the distal segments of the fourth leg in males and the unique configuration of the genital field in both sexes. Larvae of
<taxonomicName id="4C734D51FF8E1761FF4C6F06AB9DFA0D" box="[151,385,1383,1408]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B907EAC0FF8E1761FF4C6F06AB9DFA0D" box="[151,385,1383,1408]" italics="true" pageId="6" pageNumber="117">Madawaska borealis</emphasis>
</taxonomicName>
exhibit some suggestive apomorphic similarities to those of some, but not all, species of the genus
<taxonomicName id="4C734D51FF8E1761FF3B6FEDABCEFA28" authority="Smith 1976" authorityName="Smith" authorityYear="1976" box="[224,466,1420,1445]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FF3B6FEDAB2FFA28" box="[224,307,1420,1445]" italics="true" pageId="6" pageNumber="117">Forelia</emphasis>
(
<bibRefCitation id="EFE24B23FF8E1761FE996FEDABD6FA28" author="Smith" box="[322,458,1420,1445]" pageId="6" pageNumber="117" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith 1976</bibRefCitation>
)
</taxonomicName>
. The modifications to the distal segments of the fourth leg of males of
<taxonomicName id="4C734D51FF8E1761FB296FECAF6EFA28" box="[1266,1394,1420,1445]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B907EAC0FF8E1761FB296FECAF6EFA28" box="[1266,1394,1420,1445]" italics="true" pageId="6" pageNumber="117">M. borealis</emphasis>
</taxonomicName>
are more similar to those found in males of the genus
<taxonomicName id="4C734D51FF8E1761FD326FD0A989FA45" box="[745,917,1457,1480]" class="Arachnida" family="Pionidae" genus="Pionacercopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FD326FD0A989FA45" box="[745,917,1457,1480]" italics="true" pageId="6" pageNumber="117">Pionacercopsis</emphasis>
</taxonomicName>
(see below) than to those found in males of
<taxonomicName id="4C734D51FF8E1761FF4C6FB5AAF6FA60" box="[151,234,1492,1517]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FF4C6FB5AAF6FA60" box="[151,234,1492,1517]" italics="true" pageId="6" pageNumber="117">Forelia</emphasis>
</taxonomicName>
. Further resolution of the phylogenetic relationships between
<taxonomicName id="4C734D51FF8E1761FC776FB5AE2FFA60" box="[940,1075,1492,1517]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8E1761FC776FB5AE2FFA60" box="[940,1075,1492,1517]" italics="true" pageId="6" pageNumber="117">Madawaska</emphasis>
</taxonomicName>
and members of other foreliine genera may be possible when adults and larvae of additional species have been described.
</paragraph>
</subSubSection>
</treatment>
</document>