treatments-xml/data/03/A4/8E/03A48E76DC68FFE1F56236EBFCA3FEF9.xml
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<document id="BC07A2BDC0014E441B40C6D40E516EE1" ID-DOI="10.5281/zenodo.10114370" ID-ISSN="0253-116X" ID-Zenodo-Dep="10114370" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630359721084" checkinUser="valdenar" docAuthor="Assing, V." docDate="2007" docId="03A48E76DC68FFE1F56236EBFCA3FEF9" docLanguage="en" docName="LinzerbiolBeitr.2007.39.2.777-790.pdf" docOrigin="Linzer biologische Beiträge 39 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Geostiba (Sipalotricha) atrioculata Assing, 2007, sp.nov." docType="treatment" docVersion="4" lastPageNumber="785" masterDocId="FF9DF60EDC6DFFE9F439306EFF90FF83" masterDocTitle="Four new species and additional records of Geostiba from Turkey and Crete, and a new synonymy (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="790" masterPageNumber="777" pageNumber="782" updateTime="1700278669180" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7941FB61EDFED2BAA83FECCBC00622A5">Four new species and additional records of Geostiba from Turkey and Crete, and a new synonymy (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<heading id="D0FA880CDC68FFECF56236EBFC7BF918" box="[347,1003,1669,1691]" fontSize="8" level="2" pageId="5" pageNumber="782" reason="2">
<emphasis id="B979E372DC68FFECF56236EBFC89F918" bold="true" box="[347,793,1669,1691]" pageId="5" pageNumber="782">
<taxonomicName id="4C0D44E3DC68FFECF56236EBFD56F918" ID-CoL="3FSPL" authorityName="Assing" authorityYear="2007" box="[347,710,1669,1691]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="782" phylum="Arthropoda" rank="species" species="atrioculata" status="sp. nov." subGenus="Sipalotricha">
<emphasis id="B979E372DC68FFECF56236EBFE26F918" bold="true" box="[347,438,1669,1691]" italics="true" pageId="5" pageNumber="782">Geostiba</emphasis>
(
<emphasis id="B979E372DC68FFECF5FC36EBFDD5F918" bold="true" box="[453,581,1669,1691]" italics="true" pageId="5" pageNumber="782">Sipalotricha</emphasis>
)
<emphasis id="B979E372DC68FFECF66D36EBFD56F918" bold="true" box="[596,710,1669,1691]" italics="true" pageId="5" pageNumber="782">atrioculata</emphasis>
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<taxonomicNameLabel id="A24A5E09DC68FFECF6F536EBFC89F918" box="[716,793,1669,1691]" pageId="5" pageNumber="782" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="133623E5DC68FFECF71F36EBFC06F918" box="[806,918,1669,1691]" captionStart-0="Figs 10-20" captionStart-1="Fig" captionStart-2="Figs 22-34" captionStartId-0="7.[347,387,1524,1544]" captionStartId-1="8.[347,380,1122,1142]" captionStartId-2="9.[347,387,1682,1702]" captionTargetBox-0="[347,1250,425,1506]" captionTargetBox-1="[347,1262,424,1122]" captionTargetBox-2="[348,1247,497,1652]" captionTargetId-0="figure-10@7.[347,1260,559,1037]" captionTargetId-1="figure-7@8.[348,1262,425,1110]" captionTargetId-2="figure-42@9.[347,1260,718,1197]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." captionText-1="Fig. 21: Type locality of Geostiba atrioculata nov.sp." captionText-2="Figs 22-34: Geostiba atrioculata nov.sp. (22-23) and G. albimontis nov.sp. (24-34): (22-23, 33-34) spermatheca; (23) male tergite VIII; (25) male sternite VIII; (26-28) median lobe of aedeagus in lateral and in ventral view; (29-30) internal spines of median lobe in lateral and in ventral view; (31) female tergite VIII; (32) female sternite VIII. Scale bars: 24-25, 31-32: 0.2 mm; 22-23, 26-30, 33-34: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10114378" figureDoi-1="http://doi.org/10.5281/zenodo.10114380" figureDoi-2="http://doi.org/10.5281/zenodo.10114382" httpUri-0="https://zenodo.org/record/10114378/files/figure.png" httpUri-1="https://zenodo.org/record/10114380/files/figure.png" httpUri-2="https://zenodo.org/record/10114382/files/figure.png" pageId="5" pageNumber="782">Figs 10-23</figureCitation>
,
<figureCitation id="133623E5DC68FFECF79836E8FC70F918" box="[929,992,1669,1691]" captionStart="Map 1" captionStartId="5.[347,392,897,917]" captionTargetBox="[347,1254,432,897]" captionTargetId="figure-7@5.[347,1253,433,889]" captionTargetPageId="5" captionText="Map 1: Distributions of Geostiba janbellini nov.sp. (filled circle), G. atrioculata nov.sp. (open circle), and G. spinosula nov.sp. (square) in southern Anatolia." figureDoi="http://doi.org/10.5281/zenodo.10114376" httpUri="https://zenodo.org/record/10114376/files/figure.png" pageId="5" pageNumber="782">Map 1</figureCitation>
)
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<paragraph id="8BB23F60DC68FFECF56236DFFC48F970" blockId="5.[347,1255,1713,1779]" pageId="5" pageNumber="782">
<materialsCitation id="3B65353DDC68FFECF56236DFFD1EF95F" collectingDate="2006-12-23" collectionCode="V" elevation="1565" latitude="36.483334" location="Alanya" longLatPrecision="1187" longitude="32.383335" pageId="5" pageNumber="782" specimenCount="1" stateProvince="Antalya" typeStatus="holotype">
<typeStatus id="54B681C2DC68FFECF56236DFFE20F946" box="[347,432,1713,1733]" pageId="5" pageNumber="782" type="holotype">Holotype</typeStatus>
: TR [8]
<collectingRegion id="49C9F182DC68FFECF62236DFFDF5F946" box="[539,613,1713,1733]" country="Turkey" name="Antalya" pageId="5" pageNumber="782">Antalya</collectingRegion>
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E
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,
<quantity id="4CF59285DC68FFECF72A36DFFC00F946" box="[787,912,1713,1733]" metricMagnitude="3" metricUnit="m" metricValue="1.565" metricValueMax="1.6" metricValueMin="1.53" pageId="5" pageNumber="782" unit="m" value="1565.0" valueMax="1600.0" valueMin="1530.0">
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, pine litter and grass sifted,
<geoCoordinate id="EE3959A7DC68FFECF0AF36DFFB72F946" box="[1174,1250,1713,1733]" degrees="36" direction="north" minutes="29" orientation="latitude" pageId="5" pageNumber="782" precision="925" value="36.483334">36°29'N</geoCoordinate>
,
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,
<date id="FFB319A0DC68FFECF58B36A6FDB4F95F" box="[434,548,1736,1756]" pageId="5" pageNumber="782" value="2006-12-23">
<collectingDate id="EFF7E048DC68FFECF58B36A6FDB4F95F" box="[434,548,1736,1756]" pageId="5" pageNumber="782" value="2006-12-23">23.XII.2006</collectingDate>
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,
<collectionCode id="ED1CA7A5DC68FFECF60936A6FDD3F95F" box="[560,579,1736,1756]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="782" type="Museum">V</collectionCode>
. Assing
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/
<typeStatus id="54B681C2DC68FFECF69E36A6FC96F95F" box="[679,774,1736,1756]" pageId="5" pageNumber="782" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C0D44E3DC68FFECF71336A6FC7AF95F" authorityName="Assing" authorityYear="2007" box="[810,1002,1736,1756]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="782" phylum="Arthropoda" rank="species" species="atrioculata" status="sp. nov.">
<emphasis id="B979E372DC68FFECF71336A6FC7AF95F" box="[810,1002,1736,1756]" italics="true" pageId="5" pageNumber="782">Geostiba atrioculata</emphasis>
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<taxonomicNameLabel id="A24A5E09DC68FFECF7CA36A6FB8FF95F" box="[1011,1055,1736,1756]" pageId="5" pageNumber="782" rank="species">sp.n.</taxonomicNameLabel>
det.
<collectionCode id="ED1CA7A5DC68FFECF06836A6FBF4F95F" box="[1105,1124,1736,1756]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="782" type="Museum">V</collectionCode>
.
<bibRefCitation id="EF9C4291DC68FFECF04836A6FB76F95F" author="ASSING V" box="[1137,1254,1736,1756]" pageId="5" pageNumber="782" pagination="1179 - 1215" refId="ref6721" refString="ASSING V. (2006): Thirteen new species and additional records of Eastern Mediterranean Geostiba THOMSON (Coleoptera: Staphylinidae, Aleocharinae). - Linzer biologische Beitrage 38 (2): 1179 - 1215." type="journal article" year="2006">Assing 2006</bibRefCitation>
(cAss).
<typeStatus id="54B681C2DC68FFECF59B36B1FE6EF970" box="[418,510,1759,1779]" pageId="5" pageNumber="782" type="paratype">Paratypes</typeStatus>
: 3, 6: same data as
<typeStatus id="54B681C2DC68FFECF6C036B1FCD9F970" box="[761,841,1759,1779]" pageId="5" pageNumber="782" type="holotype">holotype</typeStatus>
(cAss,
<collectionCode id="ED1CA7A5DC68FFECF7B436B1FC5EF970" box="[909,974,1759,1779]" pageId="5" pageNumber="782">OÖLL</collectionCode>
).
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF736310BFCA3FEF9" blockId="6.[783,819,357,378]" box="[783,819,357,378]" pageId="6" pageNumber="783">783</paragraph>
<paragraph id="8BB23F60DC6BFFEFF56231DFFCDCFE60" blockId="6.[347,1254,432,483]" pageId="6" pageNumber="783">
D e s c r i p t i o n: 2.0-
<quantity id="4CF59285DC6BFFEFF65231DEFD50FE45" box="[619,704,432,454]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="783" unit="mm" value="2.7">2.7 mm</quantity>
. Habitus as in
<figureCitation id="133623E5DC6BFFEFF75E31DEFC26FE45" box="[871,950,432,454]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 10</figureCitation>
. Coloration: body uniformly yellowish, except for the blackish eye rudiments.
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF562319CFD19FDFE" blockId="6.[347,1255,498,637]" pageId="6" pageNumber="783">
Head 1.05-1.10 times as wide as long; puncturation extremely fine, barely noticeable; surface with shallow microreticulation and with some shine (
<figureCitation id="133623E5DC6BFFEFF7D73261FBAEFDA6" box="[1006,1086,527,549]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 11</figureCitation>
). Eyes strongly reduced (
<figureCitation id="133623E5DC6BFFEFF5833243FD96FDC0" box="[442,518,557,579]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 12</figureCitation>
), slightly smaller than antennomere III in cross-section, with pigmentation, but without ommatidia. Antennae distinctly incrassate apically; antennomeres V-X 2.0-2.5 times as wide as long.
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF56232E3FDFBFD5E" blockId="6.[347,1254,653,733]" pageId="6" pageNumber="783">
Pronotum approximately 1.1 times as wide as head and 1.05-1.10 times as wide as long; puncturation extremely fine, barely noticeable; microreticulation somewhat more distinct than that of head (
<figureCitation id="133623E5DC6BFFEFF62A32A9FDCDFD5E" box="[531,605,711,733]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF5623282FB0CFCBE" blockId="6.[347,1255,748,829]" pageId="6" pageNumber="783">
Elytra approximately as wide as pronotum or slightly wider, and about 0.55 times as long as pronotum (
<figureCitation id="133623E5DC6BFFEFF5DE3364FDA2FCA3" box="[487,562,778,800]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 11</figureCitation>
); puncturation very fine, but slightly more distinct than that of head and pronotum; microreticulation shallow; sexual dimorphism weakly pronounced.
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF5623322FD8FFC1E" blockId="6.[347,1255,844,925]" pageId="6" pageNumber="783">
Abdomen 1.05-1.10 times as wide as elytra (
<figureCitation id="133623E5DC6BFFEFF7113322FCE4FCE1" box="[808,884,844,866]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 10</figureCitation>
); puncturation very fine and sparse; microreticulation present everywhere, but shallow; posterior margin of tergite VII without palisade fringe.
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF54933C3FC10FBD7" blockId="6.[347,1255,940,1108]" pageId="6" pageNumber="783">
: elytra in postero-lateral region usually with shallow, ill-defined, somewhat diagonal impression; tergite VIII weakly transverse, posterior margin weakly convex, in the middle truncate (
<figureCitation id="133623E5DC6BFFEFF5DB3389FDBFFC7E" box="[482,559,999,1021]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 13</figureCitation>
); sternite VIII weakly transverse and with broadly convex posterior margin (
<figureCitation id="133623E5DC6BFFEFF58D346AFD94FB99" box="[436,516,1028,1050]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 14</figureCitation>
); median lobe of aedeagus as in
<figureCitation id="133623E5DC6BFFEFF753346AFC4DFB99" box="[874,989,1028,1050]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Figs 15-17</figureCitation>
; apical lobe of paramere moderately slender, with a long median seta, two moderately short setae in submedian and apical position, and a minute apical seta (
<figureCitation id="133623E5DC6BFFEFF7113450FCE4FBD7" box="[808,884,1086,1108]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF549340BFD7FFB37" blockId="6.[347,1255,1124,1204]" pageId="6" pageNumber="783">
: elytra unmodified; tergite VIII as in, but more transverse (
<figureCitation id="133623E5DC6BFFEFF03A340AFBC1FBF9" box="[1027,1105,1124,1146]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 19</figureCitation>
); sternite VIII distinctly transverse, posterior margin broadly convex and without central concavity (
<figureCitation id="133623E5DC6BFFEFF55A34F0FE3FFB37" box="[355,431,1182,1204]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Fig. 20</figureCitation>
); spermatheca as in
<figureCitation id="133623E5DC6BFFEFF64234F0FD7FFB37" box="[635,751,1182,1204]" captionStart="Figs 22-34" captionStartId="9.[347,387,1682,1702]" captionTargetBox="[348,1247,497,1652]" captionTargetId="figure-42@9.[347,1260,718,1197]" captionTargetPageId="9" captionText="Figs 22-34: Geostiba atrioculata nov.sp. (22-23) and G. albimontis nov.sp. (24-34): (22-23, 33-34) spermatheca; (23) male tergite VIII; (25) male sternite VIII; (26-28) median lobe of aedeagus in lateral and in ventral view; (29-30) internal spines of median lobe in lateral and in ventral view; (31) female tergite VIII; (32) female sternite VIII. Scale bars: 24-25, 31-32: 0.2 mm; 22-23, 26-30, 33-34: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114382" httpUri="https://zenodo.org/record/10114382/files/figure.png" pageId="6" pageNumber="783">Figs 22-23.</figureCitation>
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF56234ABFDB4FA96" blockId="6.[347,1255,1220,1301]" pageId="6" pageNumber="783">
E t y m o l o g y: The name refers to the eyes, which, in contrast to those of most other
<taxonomicName id="4C0D44E3DC6BFFEFF562348FFD9AFB74" box="[347,522,1249,1271]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="783" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B979E372DC6BFFEFF562348FFE26FB74" box="[347,438,1249,1271]" italics="true" pageId="6" pageNumber="783">Geostiba</emphasis>
species
</taxonomicName>
with minute eye rudiments without ommatidia, are of conspicuously blackish coloration.
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF562354AFD7BFA11" blockId="6.[347,1255,1316,1426]" pageId="6" pageNumber="783">
C o m p a r a t i v e n o t e s: From other Eastern Mediterranean congeners with similarly reduced eyes (minute size, ommatidia absent),
<taxonomicName id="4C0D44E3DC6BFFEFF793352CFBACFAD4" authorityName="Assing" authorityYear="2007" box="[938,1084,1345,1367]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="783" phylum="Arthropoda" rank="species" species="atrioculata">
<emphasis id="B979E372DC6BFFEFF793352CFBACFAD4" box="[938,1084,1345,1367]" italics="true" pageId="6" pageNumber="783">G. atrioculata</emphasis>
</taxonomicName>
is distinguished by the blackish coloration of the eyes. In order to account for the new species, the key in
<bibRefCitation id="EF9C4291DC6BFFEFF5623512FE6CFA11" author="ASSING V" box="[347,508,1404,1426]" pageId="6" pageNumber="783" pagination="903 - 1006" refId="ref6631" refString="ASSING V. (2005 a): A revision of the species of Geostiba THOMSON and Tropimenelytron PACE of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae, Aleocharinae). - Linzer biologische Beitrage 37 (2): 903 - 1006." type="journal article" year="2005">ASSING (2005a)</bibRefCitation>
is modified as follows:
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF56235CBFB5DFA74" blockId="6.[347,1255,1445,1581]" pageId="6" pageNumber="783">
3 Eyes reduced to minute rudiments without ommatidia .....................................................3a - Eyes sometimes small, but always with ommatidia..........................................................19 3a Eyes with pigmentation, blackish. Aedeagus and spermatheca as in
<figureCitation id="133623E5DC6BFFEFF021358CFB12FA74" box="[1048,1154,1506,1527]" captionStart="Figs 10-20" captionStartId="7.[347,387,1524,1544]" captionTargetBox="[347,1250,425,1506]" captionTargetId="figure-10@7.[347,1260,559,1037]" captionTargetPageId="7" captionText="Figs 10-20: Geostiba atrioculata nov.sp.: (10) habitus; (11) forebody; (12) head in lateral view; (13) male tergite VIII; (14) male sternite VIII; (15-17) median lobe of aedeagus in lateral and in ventral view; (18) apical lobe of paramere; (19) female tergite VIII; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="6" pageNumber="783">Figs 15-17</figureCitation>
,
<figureCitation id="133623E5DC6BFFEFF0B7358CFB59FA74" box="[1166,1225,1506,1527]" captionStart="Figs 22-34" captionStartId="9.[347,387,1682,1702]" captionTargetBox="[348,1247,497,1652]" captionTargetId="figure-42@9.[347,1260,718,1197]" captionTargetPageId="9" captionText="Figs 22-34: Geostiba atrioculata nov.sp. (22-23) and G. albimontis nov.sp. (24-34): (22-23, 33-34) spermatheca; (23) male tergite VIII; (25) male sternite VIII; (26-28) median lobe of aedeagus in lateral and in ventral view; (29-30) internal spines of median lobe in lateral and in ventral view; (31) female tergite VIII; (32) female sternite VIII. Scale bars: 24-25, 31-32: 0.2 mm; 22-23, 26-30, 33-34: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114382" httpUri="https://zenodo.org/record/10114382/files/figure.png" pageId="6" pageNumber="783">22-23</figureCitation>
.
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF5B83597FB76F9AE" blockId="6.[347,1255,1445,1581]" pageId="6" pageNumber="783">
Southern
<collectingCountry id="F31A7FF0DC6BFFEFF5E73597FDA5F98D" box="[478,565,1529,1550]" name="Turkey" pageId="6" pageNumber="783">Anatolia</collectingCountry>
:
<collectingRegion id="49C9F182DC6BFFEFF6073597FD1AF98D" box="[574,650,1529,1550]" country="Turkey" name="Antalya" pageId="6" pageNumber="783">Antalya</collectingRegion>
(
<figureCitation id="133623E5DC6BFFEFF6AE3597FD46F98D" box="[663,726,1529,1550]" captionStart="Map 1" captionStartId="5.[347,392,897,917]" captionTargetBox="[347,1254,432,897]" captionTargetId="figure-7@5.[347,1253,433,889]" captionTargetPageId="5" captionText="Map 1: Distributions of Geostiba janbellini nov.sp. (filled circle), G. atrioculata nov.sp. (open circle), and G. spinosula nov.sp. (square) in southern Anatolia." figureDoi="http://doi.org/10.5281/zenodo.10114376" httpUri="https://zenodo.org/record/10114376/files/figure.png" pageId="6" pageNumber="783">Map 1</figureCitation>
) ..................................................
<taxonomicName id="4C0D44E3DC6BFFEFF02A3597FB0BF98D" authorityName="Assing" authorityYear="2007" box="[1043,1179,1529,1550]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="783" phylum="Arthropoda" rank="species" species="atrioculata" status="sp. nov.">
<emphasis id="B979E372DC6BFFEFF02A3597FB0BF98D" box="[1043,1179,1529,1550]" italics="true" pageId="6" pageNumber="783">G. atrioculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A5E09DC6BFFEFF0983597FB77F98D" box="[1185,1255,1529,1550]" pageId="6" pageNumber="783" rank="species">nov.sp.</taxonomicNameLabel>
- Eyes without pigmentation..................................................................................................4
</paragraph>
<paragraph id="8BB23F60DC6BFFEFF562362AFB97F933" blockId="6.[347,1255,1603,1713]" pageId="6" pageNumber="783">
D i s t r i b u t i o n a n d b i o n o m i c s: The
<typeStatus id="54B681C2DC6BFFEFF7AC362AFC29F9DA" box="[917,953,1604,1625]" pageId="6" pageNumber="783">type</typeStatus>
locality is situated in the Taurus range, some
<quantity id="4CF59285DC6BFFEFF60C360EFDE9F9F5" box="[565,633,1632,1654]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="783" unit="km" value="40.0">40 km</quantity>
to the east of Alanya, eastern
<collectingRegion id="49C9F182DC6BFFEFF7F2360EFB11F9F5" box="[971,1153,1632,1654]" country="Turkey" name="Antalya" pageId="6" pageNumber="783">Antalya province</collectingRegion>
, southern
<collectingCountry id="F31A7FF0DC6BFFEFF5623613FE22F910" box="[347,434,1661,1683]" name="Turkey" pageId="6" pageNumber="783">Anatolia</collectingCountry>
(
<figureCitation id="133623E5DC6BFFEFF5F83613FD96F910" box="[449,518,1661,1683]" captionStart="Map 1" captionStartId="5.[347,392,897,917]" captionTargetBox="[347,1254,432,897]" captionTargetId="figure-7@5.[347,1253,433,889]" captionTargetPageId="5" captionText="Map 1: Distributions of Geostiba janbellini nov.sp. (filled circle), G. atrioculata nov.sp. (open circle), and G. spinosula nov.sp. (square) in southern Anatolia." figureDoi="http://doi.org/10.5281/zenodo.10114376" httpUri="https://zenodo.org/record/10114376/files/figure.png" pageId="6" pageNumber="783">Map 1</figureCitation>
). The
<typeStatus id="54B681C2DC6BFFEFF6743610FDE9F910" box="[589,633,1662,1683]" pageId="6" pageNumber="783">type</typeStatus>
specimens were collected at a pass below old pine trees by sifting litter and grass roots at an altitude of approximately
<quantity id="4CF59285DC6BFFEFF78D36F5FB93F933" box="[948,1027,1691,1713]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="783" unit="m" value="1600.0">1600 m</quantity>
.
</paragraph>
<paragraph id="8BB23F60DC6AFFEEF736310BFCA3FEF9" blockId="7.[783,819,357,378]" box="[783,819,357,378]" pageId="7" pageNumber="784">784</paragraph>
<caption id="DF726FE8DC6AFFEEF562359AFCF2F9CF" ID-DOI="http://doi.org/10.5281/zenodo.10114378" ID-Zenodo-Dep="10114378" httpUri="https://zenodo.org/record/10114378/files/figure.png" pageId="7" pageNumber="784" startId="7.[347,387,1524,1544]" targetBox="[347,1250,425,1506]" targetPageId="7">
<paragraph id="8BB23F60DC6AFFEEF562359AFCF2F9CF" blockId="7.[347,1255,1524,1612]" pageId="7" pageNumber="784">
<emphasis id="B979E372DC6AFFEEF562359AFE51F98B" bold="true" box="[347,449,1524,1544]" pageId="7" pageNumber="784">Figs 10-20</emphasis>
:
<taxonomicName id="4C0D44E3DC6AFFEEF5E9359AFD00F98B" authorityName="Assing" authorityYear="2007" box="[464,656,1524,1544]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="784" phylum="Arthropoda" rank="species" species="atrioculata" status="sp. nov.">
<emphasis id="B979E372DC6AFFEEF5E9359AFD00F98B" box="[464,656,1524,1544]" italics="true" pageId="7" pageNumber="784">Geostiba atrioculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A5E09DC6AFFEEF6A1359AFD4EF98B" box="[664,734,1524,1544]" pageId="7" pageNumber="784" rank="species">nov.sp.</taxonomicNameLabel>
: (
<emphasis id="B979E372DC6AFFEEF6C9359AFC97F98B" bold="true" box="[752,775,1524,1544]" pageId="7" pageNumber="784">10</emphasis>
) habitus; (
<emphasis id="B979E372DC6AFFEEF756359AFC16F98B" bold="true" box="[879,902,1524,1544]" pageId="7" pageNumber="784">11</emphasis>
) forebody; (
<emphasis id="B979E372DC6AFFEEF7C7359AFB85F98B" bold="true" box="[1022,1045,1524,1544]" pageId="7" pageNumber="784">12</emphasis>
) head in lateral view; (
<emphasis id="B979E372DC6AFFEEF55A3664FEEAF99D" bold="true" box="[355,378,1546,1566]" pageId="7" pageNumber="784">13</emphasis>
) male tergite VIII; (
<emphasis id="B979E372DC6AFFEEF6053664FDC3F99D" bold="true" box="[572,595,1546,1566]" pageId="7" pageNumber="784">14</emphasis>
) male sternite VIII; (
<emphasis id="B979E372DC6AFFEEF7243664FCC2F99D" bold="true" box="[797,850,1546,1566]" pageId="7" pageNumber="784">15-17</emphasis>
) median lobe of aedeagus in lateral and in ventral view; (
<emphasis id="B979E372DC6AFFEEF5D9364FFE67F9B6" bold="true" box="[480,503,1569,1589]" pageId="7" pageNumber="784">18</emphasis>
) apical lobe of paramere; (
<emphasis id="B979E372DC6AFFEEF6D6364FFC96F9B6" bold="true" box="[751,774,1569,1589]" pageId="7" pageNumber="784">19</emphasis>
) female tergite VIII; (
<emphasis id="B979E372DC6AFFEEF7EB364FFC79F9B6" bold="true" box="[978,1001,1569,1589]" pageId="7" pageNumber="784">20</emphasis>
) female sternite VIII. Scale bars: 10: 1.0 mm; 11-14, 19-20: 0.2 mm; 15-18: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BB23F60DC65FFE1F736310BFCA3FEF9" blockId="8.[783,819,357,378]" box="[783,819,357,378]" pageId="8" pageNumber="785">785</paragraph>
</subSubSection>
</treatment>
</document>