treatments-xml/data/03/7B/E0/037BE0492B2AFFA0FF7ED2DD539690B6.xml

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<mods:title id="980B17BF1C83E9A1B9B21F7F3E0DC307">Two new genera and five new species of Mugadina - like small grass cicadas (Hemiptera: Cicadidae: Cicadettini) from Central and Eastern Australia: comparative morphology, songs, behaviour and distributions</mods:title>
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<treatment id="037BE0492B2AFFA0FF7ED2DD539690B6" ID-DOI="http://doi.org/10.5281/zenodo.5974120" ID-GBIF-Taxon="143619668" ID-Zenodo-Dep="5974120" LSID="urn:lsid:plazi:treatment:037BE0492B2AFFA0FF7ED2DD539690B6" httpUri="http://treatment.plazi.org/id/037BE0492B2AFFA0FF7ED2DD539690B6" lastPageId="15" lastPageNumber="15" pageId="3" pageNumber="4">
<subSubSection id="C3C802D42B2AFFACFF7ED2DD55AB929D" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8B6D515F2B2AFFACFF7ED2DD552392FC" blockId="3.[151,644,932,991]" box="[151,524,932,958]" pageId="3" pageNumber="4">
<heading id="D025E6332B2AFFACFF7ED2DD552392FC" bold="true" box="[151,524,932,958]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="B9A68D4D2B2AFFACFF7ED2DD552392FC" bold="true" box="[151,524,932,958]" pageId="3" pageNumber="4">
Genus 
<taxonomicName id="4CD22ADC2B2AFFACFF07D2DD569C92FC" ID-CoL="8JL9T" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[238,435,932,958]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B2AFFACFF07D2DD569C92FC" bold="true" box="[238,435,932,958]" italics="true" pageId="3" pageNumber="4">HEREMUSINA</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2AFFACFE53D2DD552392FC" box="[442,524,932,958]" pageId="3" pageNumber="4" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B6D515F2B2AFFACFF7ED2BF55AB929D" blockId="3.[151,644,932,991]" box="[151,644,966,991]" pageId="3" pageNumber="4">
<figureCitation id="13E94DDA2B2AFFACFF7ED2BF57D7929D" box="[151,248,966,991]" captionStart="FIGURE2" captionStartId="5.[151,250,1814,1836]" captionTargetBox="[211,1376,193,1783]" captionTargetId="figure@5.[211,1376,193,1793]" captionTargetPageId="5" captionText="FIGURE2. Lateralabdomen views of the five new species described in thispaper;(A), (B), Heremusina udeoecetes n. sp.and H. pipatio n. sp.; (C) to (E), Xeropsalta thomsoni n. sp., X. aridula n. sp., X.rattrayi n. sp., respectively." httpUri="https://zenodo.org/record/1221892/files/figure.png" pageId="3" pageNumber="4">Figs. 2A</figureCitation>
– 
<figureCitation id="13E94DDA2B2AFFACFEE1D2BE5605929D" box="[264,298,967,991]" captionStart="FIGURE7" captionStartId="10.[151,250,1814,1836]" captionTargetBox="[226,1361,193,1778]" captionTargetId="figure@10.[226,1361,193,1793]" captionTargetPageId="10" captionText="FIGURE7. Pygofer and male genitalia,ventral views, respectively,of the five new species described in thispaper; (A), (B),H. udeoecetes n. sp. and H.pipatio n. sp.; (C) to (E), X. thomsoni n. sp. (see also Pl. 4), X. aridula n. sp., X. rattrayi n. sp. Sizes as in Fig. 6." httpUri="https://zenodo.org/record/1221902/files/figure.png" pageId="3" pageNumber="4">7A</figureCitation>
, 
<figureCitation id="13E94DDA2B2AFFACFEDFD2BE5678929D" box="[310,343,967,991]" captionStart="FIGURE2" captionStartId="5.[151,250,1814,1836]" captionTargetBox="[211,1376,193,1783]" captionTargetId="figure@5.[211,1376,193,1793]" captionTargetPageId="5" captionText="FIGURE2. Lateralabdomen views of the five new species described in thispaper;(A), (B), Heremusina udeoecetes n. sp.and H. pipatio n. sp.; (C) to (E), Xeropsalta thomsoni n. sp., X. aridula n. sp., X.rattrayi n. sp., respectively." httpUri="https://zenodo.org/record/1221892/files/figure.png" pageId="3" pageNumber="4">2B</figureCitation>
– 
<figureCitation id="13E94DDA2B2AFFACFE8ED2BE56A7929D" box="[359,392,967,991]" captionStart="FIGURE7" captionStartId="10.[151,250,1814,1836]" captionTargetBox="[226,1361,193,1778]" captionTargetId="figure@10.[226,1361,193,1793]" captionTargetPageId="10" captionText="FIGURE7. Pygofer and male genitalia,ventral views, respectively,of the five new species described in thispaper; (A), (B),H. udeoecetes n. sp. and H.pipatio n. sp.; (C) to (E), X. thomsoni n. sp. (see also Pl. 4), X. aridula n. sp., X. rattrayi n. sp. Sizes as in Fig. 6." httpUri="https://zenodo.org/record/1221902/files/figure.png" pageId="3" pageNumber="4">7B</figureCitation>
, 
<figureCitation id="13E94DDA2B2AFFACFE7AD2BE568D929D" box="[403,418,967,991]" captionStart="FIGURE 8" captionStartId="11.[151,250,1344,1366]" captionTargetBox="[151,1436,193,1323]" captionTargetId="figure@11.[151,1436,193,1323]" captionTargetPageId="11" captionText="FIGURE 8. Map of Central and Northeastern Australia, showing the currently known occurrences of Heremusina udeoecetes n. sp., H. pipatio n. sp., Xeropsalta thomsoni n. sp., X.aridula n. sp., and X. rattrayi n. sp. The heavy dot-dash line shows the approximate northern and eastern boundaries of the ecoregion which defines the dominance of deserts and xeric shrublands within inland Australia (based on National Reserve System map, published by the Australian Government). The fine dotted lines are major roads, the names of some prominent towns listed. Lake Eyre and associated ephemoral lakes are shown by the darker shading. The approximate outlines of the modern dunefields comprising much of the Simpson and Strzelecki deserts are shown by the lighter shading, these based on 'Google Earth' images." httpUri="https://zenodo.org/record/1221904/files/figure.png" pageId="3" pageNumber="4">8</figureCitation>
, 
<figureCitation id="13E94DDA2B2AFFACFE46D2BE56D7929D" box="[431,504,967,991]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStart-3="FIGURE15" captionStart-4="FIGURE 16" captionStart-5="FIGURE 17" captionStartId-0="16.[151,250,1798,1820]" captionStartId-1="17.[151,250,1732,1754]" captionStartId-2="18.[151,250,1760,1782]" captionStartId-3="19.[151,250,1872,1894]" captionStartId-4="22.[151,250,1708,1730]" captionStartId-5="23.[151,250,1641,1663]" captionTargetBox-0="[240,1347,185,1785]" captionTargetBox-1="[295,1289,193,1717]" captionTargetBox-2="[259,1328,193,1741]" captionTargetBox-3="[152,1436,193,1851]" captionTargetBox-4="[274,1318,185,1697]" captionTargetBox-5="[227,1360,153,1619]" captionTargetId-0="figure@16.[239,1347,185,1785]" captionTargetId-1="figure@17.[294,1289,193,1717]" captionTargetId-2="figure@18.[257,1328,193,1741]" captionTargetId-3="figure@19.[151,1436,193,1851]" captionTargetId-4="figure@22.[274,1318,185,1697]" captionTargetId-5="figure@23.[226,1360,153,1619]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="22" captionTargetPageId-5="23" captionText-0="FIGURE 12. Heremusina udeoecetes n. sp. Waveform plots, based on field recordings at 11 km N.W. along Tanami Desert Road, 20 km north of Alice Springs, in low grassland. Figs. A, D show the characteristic repeated echemes, these having durations ranging between 272–332 ms, each commencing with 2–9 isolated pulses, followed by a sequence of closely spaced pulses; an isolatedpulse isemitted at end of each echeme.Figs.B, Dare time expandedwaveform plots showingmore detail of the pulses. The initial pulses (Fig. B) are isolated pulses, these changing to doublet pulses within the continuous echeme segment.Pulses (including doublet pulses) have repetition rates varying between 160–193 Hz. (C), shows the final segment of an echeme in which the pulses alternate between single and double pulses, followed by the single isolated final pulse. Filtered to 3.5 kHz." captionText-1="FIGURE 13. Heremusina udeoecetes n. sp. Time expanded details of selected pulses shown in Fig. 12, shown as waveform and companion envelope plots. In 13A, B, the initial single pulse is markedly of longer duration compared to each of the following double pulses, suggesting that it results from a partial overlapping of asynchronous pulses from the two timbals; the double clicks are therefore interpreted as temporally more clearly separated pulses emitted by the two timbals.Figs 13C, D are strongly developed doublet pulses, emitted asynchronously by the two timbals. The following smaller peaks which are almost coalesced with the main peak are believed to represent the emission from timbal ribs 2–5. The peaks labeled 'S' are interpreted to be secondary ('OUT') clicks of the timbals, not seen following first peak most likely as they overlap the emission of the second primary peak of each doublet. Fig. 13E, F, time expanded single pulses emitted at start of echeme, these showing merged weaker peaks (probably emissionsof ribs 2–5).Secondary peaks, marked 'S', occurring 1.7ms after the primary peaks, are inferred to represent be 'OUT' clicks of the timbals.Filtered to3.5 kHz." captionText-2="FIGURE 14. Heremusina udeoecetes n. sp., waveform (A, B, D) and envelope (C, E) plots from parabola recordings made 3 km east of Hermannsburg, N.T., ~112 km WSW of Alice Springs; 14A shows an echeme segment with two initial isolated pulses, followed by rapidly repeated doublet pulses, the PRR measured at 192 Hz. Figs.14B, C, are time expanded plots of the pulse doubletsshowing both thepeaks tobe stronglydeveloped, witha weak secondary peak1.5 ms followingthe second peak, evidently not so clearly developed after the first peak (the doublet inferred to be emitted asynchronously from both timbals); Fig.14D, E illustratessingle pulses occurring at startof echeme showing only weakly developed secondary peaks(marked 'S'). These plots also shows small but increasingly stronger peaks (arrowed) almost joined to the preceding primary peaks, these thought to represent the initial development of doublet peaks (i.e. increasing asynchrony of timbal emissions). Filtered to 7.5 kHz (A), and 9 kHz (B to E)." captionText-3="FIGURE15. Comparison of Heremusina udeoecetes n. sp. song in terms of, (A), waveformplot (temporalstructures of song), (B),spectrogram showing timbal emissionfrequencies, these concentrated inrange ~11–17 kHz, but exhibiting no indication of systematic changes of frequencies during song emission, although minor frequency disturbances may occur at the start of each echeme; and (C), amplitude spectrum showing the dominant frequency envelope between 11.5–16.8 kHz, with the estimated dominant frequency at 13.9 kHz.Filtered to 3.5 kHz (A), and 7.5 kHz (C)." captionText-4="FIGURE 16. Heremusina pipatio n. sp. song, field recording from ~15 km N. of Cloncurry, N.W. Queensland. Fig. 16A is a waveform overview showing the structure of the echemes which comprise sets of closely spaced single pulse emissions, without any isolated single pulses at the start of each echeme (except at the beginning of anew songsequence, as shown in this segment). Each echeme is followed with an isolated single pulse. The echeme durations (52–63 ms) are clearly shorter than those of H. udeoecetes n. sp., and the resulting echeme repetition rates therefore higher (9.2–10.7 Hz). Pulse repetition rates withinthe echemes range between 180–225 (mean207) Hz.Fig.16B, C, time expanded waveform plot of a single echeme, and the following isolated pulse, showing the pulses to be single pulses, presumably emitted by the alternation of the timbal contractions; Fig. 16C, D, higher resolution, time expanded waveform and envelope plots of three echeme pulses showing the initial high amplitude peaks immediately followed, without a gap, by weaker peaks, inferred as ribs 2–5 emissions. Well developed, isolated secondary peaks (marked as 'S') occur 1.7 ms following the preceding main peaks, these inferred to be secondary, relaxation ('OUT') pulses of the timbals. Filtered to 1 kHz." captionText-5="FIGURE 17. Comparison of Heremusina pipatio n. sp. songs in relation to; (A) waveform plot showing temporal pulses structures; (B), spectrogram, showing a dominant frequency band between ~10 to 18 kHz, without significant frequency shift during the song; and (C), amplitude spectra showing the clearly developed frequency envelope between 10.7–18 kHz, with the dominant frequency estimated at 14.0 kHz, located at the low amplitude centre of the frequency envelope.Filtered to 1 kHz." httpUri-0="https://zenodo.org/record/1221912/files/figure.png" httpUri-1="https://zenodo.org/record/1221914/files/figure.png" httpUri-2="https://zenodo.org/record/1221916/files/figure.png" httpUri-3="https://zenodo.org/record/1221918/files/figure.png" httpUri-4="https://zenodo.org/record/1221920/files/figure.png" httpUri-5="https://zenodo.org/record/1221922/files/figure.png" pageId="3" pageNumber="4">12–17</figureCitation>
, 
<figureCitation id="13E94DDA2B2AFFACFDEAD2BE550D929D" box="[515,546,967,991]" captionStart="FIGURE 30" captionStartId="46.[151,250,1538,1560]" captionTargetBox="[166,1420,299,1518]" captionTargetId="figure@46.[166,1420,299,1518]" captionTargetPageId="46" captionText="FIGURE 30. Generalised map of central and northeastern Australia showing the distributions of the described Heremusina n. gen. andXeropsalta n. gen. species, as in Fig. 8, compared to the geography of selected river systems of inland Australia.The more heavily dotted lines show the systems draining towards the Lake Eyre Basin; the finely dotted lines of the rivers systems of the southeastern area of the Northern Territory are internally draining, not reaching beyond the State boundaries, while the river systems of northern and eastern Queensland, northern Northern Territory and NSW drain into the surrounding oceans." httpUri="https://zenodo.org/record/1221948/files/figure.png" pageId="3" pageNumber="4">30</figureCitation>
; Pl. 1, 2.
</paragraph>
</subSubSection>
<subSubSection id="C3C802D42B2AFFACFF7ED570534C9507" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8B6D515F2B2AFFACFF7ED570534C9507" blockId="3.[151,1437,1033,1954]" pageId="3" pageNumber="4">
<emphasis id="B9A68D4D2B2AFFACFF7ED57056379560" bold="true" box="[151,280,1033,1058]" pageId="3" pageNumber="4">Etymology</emphasis>
: Derived from Latin, a feminine construct of the masculine Heremus, referring to wilderness/desert/ wasteland, which broadly describe the environments of the two new species described.
</paragraph>
</subSubSection>
<subSubSection id="C3C802D42B2AFFACFF2ED529521595CF" pageId="3" pageNumber="4" type="distribution">
<paragraph id="8B6D515F2B2AFFACFF2ED529521595CF" blockId="3.[151,1437,1033,1954]" pageId="3" pageNumber="4">
<emphasis id="B9A68D4D2B2AFFACFF2ED5295676952B" bold="true" box="[199,345,1104,1129]" pageId="3" pageNumber="4">Distribution</emphasis>
(
<figureCitation id="13E94DDA2B2AFFACFE99D5295694952B" box="[368,443,1104,1129]" captionStart="FIGURE 8" captionStartId="11.[151,250,1344,1366]" captionTargetBox="[151,1436,193,1323]" captionTargetId="figure@11.[151,1436,193,1323]" captionTargetPageId="11" captionText="FIGURE 8. Map of Central and Northeastern Australia, showing the currently known occurrences of Heremusina udeoecetes n. sp., H. pipatio n. sp., Xeropsalta thomsoni n. sp., X.aridula n. sp., and X. rattrayi n. sp. The heavy dot-dash line shows the approximate northern and eastern boundaries of the ecoregion which defines the dominance of deserts and xeric shrublands within inland Australia (based on National Reserve System map, published by the Australian Government). The fine dotted lines are major roads, the names of some prominent towns listed. Lake Eyre and associated ephemoral lakes are shown by the darker shading. The approximate outlines of the modern dunefields comprising much of the Simpson and Strzelecki deserts are shown by the lighter shading, these based on 'Google Earth' images." httpUri="https://zenodo.org/record/1221904/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
): 
<collectingRegion id="49169FBD2B2AFFACFE30D5295580952B" box="[473,687,1104,1129]" country="Australia" name="Northern Territory" pageId="3" pageNumber="4">Northern Territory</collectingRegion>
in the regions around Alice Springs, extending westwards to Hermannsburg, and northern 
<collectingRegion id="49169FBD2B2AFFACFE09D50D554B95CF" box="[480,612,1140,1165]" country="Australia" name="Queensland" pageId="3" pageNumber="4">Queensland</collectingRegion>
in the Cloncurry region, north and south of Cloncurry township.
</paragraph>
</subSubSection>
<subSubSection id="C3C802D42B2AFFA4FF2ED5E15574966E" lastPageId="11" lastPageNumber="12" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8B6D515F2B2AFFA4FF2ED5E15574966E" blockId="3.[151,1437,1033,1954]" lastBlockId="11.[151,1436,1595,2014]" lastPageId="11" lastPageNumber="12" pageId="3" pageNumber="4">
<emphasis id="B9A68D4D2B2AFFACFF2ED5E1561595F3" bold="true" box="[199,314,1176,1201]" pageId="3" pageNumber="4">Diagnosis</emphasis>
: 
<emphasis id="B9A68D4D2B2AFFACFEA3D5E156A895F3" box="[330,391,1176,1201]" italics="true" pageId="3" pageNumber="4">Head</emphasis>
: Outer margins of eyes approximately as wide as mesonotum; supra-antennal plate meeting eye; postclypeus outline rounded across ventral midline; in lateral profile gently rounded between 'top' and 'sides'. 
<emphasis id="B9A68D4D2B2AFFACFF7ED5A657C795BA" box="[151,232,1247,1272]" italics="true" pageId="3" pageNumber="4">Thorax</emphasis>
: Pronotum in dorsal view parallel-sided; pronotal collar width at dorsal midline much less than diameter of eyes; paranota confluent with adjoining pronotal sclerites, no mid lateral tooth; cruciform elevation wider than long; epimeral lobe not reaching operculum; metanotum clearly visible at dorsal midline. 
<emphasis id="B9A68D4D2B2AFFACFB6CD451522D947D" box="[1157,1282,1320,1343]" italics="true" pageId="3" pageNumber="4">Fore wings</emphasis>
hyaline, with 8 apical cells; subapical cells absent; ulnar cell 3 angled to radial cell; basal cell clearly longer than narrow; costal vein higher than R+Sc; costa nearly parallel-sided to node, very slight widening at node; costa almost straight with only minimal curvature; pterostigma with pale brown infuscation; vein CuA with gentle curvature, with the cubital cell larger than medial cell; veins M and CuA meeting basal cell with their stems fused as one; vein RA1 not completely aligned with Sc for its full length, slightly diverging subapically; vein CuA1 divided by cross-vein mcu, the two segments of similar length; veins CuP and 1A fused for most of their length; distance between cross veins 
<taxonomicName id="4CD22ADC2B2AFFACFF31D75C563F977F" box="[216,272,1572,1597]" pageId="3" pageNumber="4" rank="subSpecies" subSpecies="and">r and</taxonomicName>
r-m approximately equal to distance between r-m and m; apical cells 3 to 6 more or less equal to lengths of ulnar cells, some slightly shorter, some slightly longer; radial cell shorter than distance from its apex to wing tip; wing outer margin clearly developed for its total length. 
<emphasis id="B9A68D4D2B2AFFACFCADD71254E697C6" box="[836,969,1643,1668]" italics="true" pageId="3" pageNumber="4">Hind wings</emphasis>
with 3 to 4 apical cells, mostly 4; no infuscation on ambient vein; width of 1 
<taxonomicName id="4CD22ADC2B2AFFACFDA3D7F6559C97EA" box="[586,691,1679,1704]" pageId="3" pageNumber="4" rank="subSpecies" subSpecies="cubital">st cubital</taxonomicName>
cell at distal end more than twice that of 2nd cubital cell; anal lobe narrow with vein 3A curved, separated from wing margin for most of its length. 
<emphasis id="B9A68D4D2B2AFFACFBC4D7CA53FE978E" box="[1069,1233,1715,1740]" italics="true" pageId="3" pageNumber="4">Male opercula</emphasis>
: Relatively broad and bulbous in ventral view, mostly reaching margin of timbal cavity, excepting between lateral margin and crest; distal margin very broadly rounded; margins raised above level of tympanal cavity; basal plate strongly domed and relatively large; opercula not reaching across abdomen. 
<emphasis id="B9A68D4D2B2AFFACFCEFD66754829675" box="[774,941,1822,1847]" italics="true" pageId="3" pageNumber="4">Male abdomen</emphasis>
generally bulbous, most strongly developed around tergites 2 to 4 and sternites II to IV, clearly wider than thorax; tergites in cross-section somewhat tent-like, dorsally straight, curved ventrally; epipleurites reflexed ventrally from junction with tergites; tergite 1 narrowed along dorsal midline; tergite 2 slightly wider than tergite 3 along dorsal midline; sternites in cross-section convex, relatively swollen so that in lateral view are mostly at least partially visible. 
<emphasis id="B9A68D4D2B22FFA4FBEBD74253769716" box="[1026,1113,1595,1620]" italics="true" pageId="11" pageNumber="12">Timbals</emphasis>
with 3 continuous long ribs with long ribs 1 to 3 unbroken, relatively similar in length, each unfused ventrally, and spanning the full height of timbal; long ribs 1 to 3 fused dorsally to basal spur; rib 4 very thin medially, and not always clearly seen to be continuous across timbal, ventrally and dorsally unfused; rib 5 (most posterior) short; 3 to 4 intercalary ribs, these very thin and not easily seen; basal dome bulbous, narrow, somewhat lens-like; the outer anterior area to basal dome broad and gently undulating; anterior area of timbal dominantly occupied by ribs; timbals not extended below wing bases; timbal covers absent.
</paragraph>
</subSubSection>
<caption id="DFAD01D72B2DFFABFF7ED631524E969B" httpUri="https://zenodo.org/record/1221890/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,250,1864,1886]" targetBox="[283,1306,215,1807]" targetPageId="4">
<paragraph id="8B6D515F2B2DFFABFF7ED631524E969B" blockId="4.[151,1436,1864,2009]" pageId="4" pageNumber="5">
<emphasis id="B9A68D4D2B2DFFABFF7ED631563A961C" bold="true" box="[151,277,1864,1886]" pageId="4" pageNumber="5">FIGURE 1.</emphasis>
<taxonomicName id="4CD22ADC2B2DFFABFEF4D63156E5961D" box="[285,458,1864,1887]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="emma">
<emphasis id="B9A68D4D2B2DFFABFEF4D63156E5961D" box="[285,458,1864,1887]" italics="true" pageId="4" pageNumber="5">Mugadina emma</emphasis>
</taxonomicName>
and 
<taxonomicName id="4CD22ADC2B2DFFABFDE9D63355AA961C" baseAuthorityName="Distant" baseAuthorityYear="1911" box="[512,645,1864,1887]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B9A68D4D2B2DFFABFDE9D63355AA961C" box="[512,645,1864,1887]" italics="true" pageId="4" pageNumber="5">M. marshalli</emphasis>
</taxonomicName>
, comparison of selected morphological characters of both species. (A), (B), lateral views of abdomen. (C), (D), lateral views of male genitalia, both lengths 1.9 mm. (E), (F), ventral views of same genitalia; (G, H), timbals; dorsal to right, and (I), (J) left opercula. Figs. C, D based on specimens from 4 km north of Amby, southern Queensland and Figs. D, F from Lake Broadwater, southeast Queensland. Figs. A, G, I, based on specimens from Boonah, southeastern Queensland and Figs. B, H, J from Drillham, southern Queensland. The scale bars represent 1 mm.
</paragraph>
</caption>
<caption id="DFAD01D72B2CFFAAFF7ED66F53919609" httpUri="https://zenodo.org/record/1221892/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,250,1814,1836]" targetBox="[211,1376,193,1783]" targetPageId="5">
<paragraph id="8B6D515F2B2CFFAAFF7ED66F53919609" blockId="5.[151,1435,1814,1867]" pageId="5" pageNumber="6">
<emphasis id="B9A68D4D2B2CFFAAFF7ED66F563D966E" bold="true" box="[151,274,1814,1836]" pageId="5" pageNumber="6">FIGURE 2.</emphasis>
Lateral abdomen views of the five new species described in this paper; (A), (B), 
<taxonomicName id="4CD22ADC2B2CFFAAFBAAD661521C966E" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1091,1331,1814,1837]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="udeoecetes" status="sp. nov.">
<emphasis id="B9A68D4D2B2CFFAAFBAAD661521C966E" box="[1091,1331,1814,1837]" italics="true" pageId="5" pageNumber="6">Heremusina udeoecetes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2CFFAAFAD0D66F525D966E" box="[1337,1394,1814,1836]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B9A68D4D2B2CFFAAFAD0D66F525D966E" bold="true" box="[1337,1394,1814,1836]" pageId="5" pageNumber="6">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4CD22ADC2B2CFFAAFF7ED64F57D19609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[151,254,1846,1867]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pipatio" status="sp. nov.">
<emphasis id="B9A68D4D2B2CFFAAFF7ED64F57D19609" box="[151,254,1846,1867]" italics="true" pageId="5" pageNumber="6">H. pipatio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2CFFAAFEEDD64C56129609" box="[260,317,1845,1867]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B9A68D4D2B2CFFAAFEEDD64C56129609" bold="true" box="[260,317,1845,1867]" pageId="5" pageNumber="6">n. sp.</emphasis>
</taxonomicNameLabel>
; (C) to (E), 
<taxonomicName id="4CD22ADC2B2CFFAAFE50D64C55A59609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[441,650,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="thomsoni" status="sp. nov.">
<emphasis id="B9A68D4D2B2CFFAAFE50D64C55A59609" box="[441,650,1845,1867]" italics="true" pageId="5" pageNumber="6">Xeropsalta thomsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2CFFAAFD78D64C55E59609" box="[657,714,1845,1867]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B9A68D4D2B2CFFAAFD78D64C55E59609" bold="true" box="[657,714,1845,1867]" pageId="5" pageNumber="6">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B2CFFAAFD3FD64F54119609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[726,830,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aridula" status="sp. nov.">
<emphasis id="B9A68D4D2B2CFFAAFD3FD64F54119609" box="[726,830,1845,1867]" italics="true" pageId="5" pageNumber="6">X. aridula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2CFFAAFCADD64C54529609" box="[836,893,1845,1867]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B9A68D4D2B2CFFAAFCADD64C54529609" bold="true" box="[836,893,1845,1867]" pageId="5" pageNumber="6">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B2CFFAAFC60D64F54DD9609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[905,1010,1846,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rattrayi" status="sp. nov.">
<emphasis id="B9A68D4D2B2CFFAAFC60D64F54DD9609" box="[905,1010,1846,1867]" italics="true" pageId="5" pageNumber="6">X. rattrayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2CFFAAFC10D64C531D9609" box="[1017,1074,1845,1867]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B9A68D4D2B2CFFAAFC10D64C531D9609" bold="true" box="[1017,1074,1845,1867]" pageId="5" pageNumber="6">n. sp.</emphasis>
</taxonomicNameLabel>
, respectively.
</paragraph>
</caption>
<caption id="DFAD01D72B2FFFA9FF7ED66F569F9628" httpUri="https://zenodo.org/record/1221894/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1814,1836]" targetBox="[217,1369,193,1775]" targetPageId="6">
<paragraph id="8B6D515F2B2FFFA9FF7ED66F569F9628" blockId="6.[151,1436,1814,1898]" pageId="6" pageNumber="7">
<emphasis id="B9A68D4D2B2FFFA9FF7ED66F563C966E" bold="true" box="[151,275,1814,1836]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
Male opercula, all left opercula, of the five new species described in this paper; (A), (B), 
<taxonomicName id="4CD22ADC2B2FFFA9FB42D66152B3966E" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1195,1436,1814,1837]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="udeoecetes" status="sp. nov.">
<emphasis id="B9A68D4D2B2FFFA9FB42D66152B3966E" box="[1195,1436,1814,1837]" italics="true" pageId="6" pageNumber="7">Heremusina udeoecetes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2FFFA9FF7ED64C57FE9609" box="[151,209,1845,1867]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B9A68D4D2B2FFFA9FF7ED64C57FE9609" bold="true" box="[151,209,1845,1867]" pageId="6" pageNumber="7">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4CD22ADC2B2FFFA9FEEAD64F56449609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[259,363,1846,1867]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pipatio" status="sp. nov.">
<emphasis id="B9A68D4D2B2FFFA9FEEAD64F56449609" box="[259,363,1846,1867]" italics="true" pageId="6" pageNumber="7">H. pipatio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2FFFA9FE9BD64C56849609" box="[370,427,1845,1867]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B9A68D4D2B2FFFA9FE9BD64C56849609" bold="true" box="[370,427,1845,1867]" pageId="6" pageNumber="7">n. sp.</emphasis>
</taxonomicNameLabel>
; (C) to (E), 
<taxonomicName id="4CD22ADC2B2FFFA9FDC2D64C55D29609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[555,765,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="thomsoni" status="sp. nov.">
<emphasis id="B9A68D4D2B2FFFA9FDC2D64C55D29609" box="[555,765,1845,1867]" italics="true" pageId="6" pageNumber="7">Xeropsalta thomsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2FFFA9FCEDD64C54119609" box="[772,830,1845,1867]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B9A68D4D2B2FFFA9FCEDD64C54119609" bold="true" box="[772,830,1845,1867]" pageId="6" pageNumber="7">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B2FFFA9FCA2D64F549C9609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[843,947,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aridula" status="sp. nov.">
<emphasis id="B9A68D4D2B2FFFA9FCA2D64F549C9609" box="[843,947,1845,1867]" italics="true" pageId="6" pageNumber="7">X. aridula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2FFFA9FC50D64C54DC9609" box="[953,1011,1845,1867]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B9A68D4D2B2FFFA9FC50D64C54DC9609" bold="true" box="[953,1011,1845,1867]" pageId="6" pageNumber="7">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B2FFFA9FBE8D64F53459609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1025,1130,1846,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rattrayi" status="sp. nov.">
<emphasis id="B9A68D4D2B2FFFA9FBE8D64F53459609" box="[1025,1130,1846,1867]" italics="true" pageId="6" pageNumber="7">X. rattrayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2FFFA9FB9BD64C53839609" box="[1138,1196,1845,1867]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B9A68D4D2B2FFFA9FB9BD64C53839609" bold="true" box="[1138,1196,1845,1867]" pageId="6" pageNumber="7">n. sp.</emphasis>
</taxonomicNameLabel>
, respectively. Opercula nomenclature shown in (E).
</paragraph>
</caption>
<caption id="DFAD01D72B2EFFA8FF7ED606540A96B0" httpUri="https://zenodo.org/record/1221896/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1919,1941]" targetBox="[163,1417,193,1911]" targetPageId="7">
<paragraph id="8B6D515F2B2EFFA8FF7ED606540A96B0" blockId="7.[151,1436,1919,2034]" pageId="7" pageNumber="8">
<emphasis id="B9A68D4D2B2EFFA8FF7ED606563B96D7" bold="true" box="[151,276,1919,1941]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
Left timbal, posterior margin to right, dorsal margin at top, of the five new species described in this paper; (A), (B), 
<taxonomicName id="4CD22ADC2B2EFFA8FF2CD6E6569A96F6" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[197,437,1950,1972]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="udeoecetes" status="sp. nov.">
<emphasis id="B9A68D4D2B2EFFA8FF2CD6E6569A96F6" box="[197,437,1950,1972]" italics="true" pageId="7" pageNumber="8">Heremusina udeoecetes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2EFFA8FE52D6E756DB96F6" box="[443,500,1950,1972]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B9A68D4D2B2EFFA8FE52D6E756DB96F6" bold="true" box="[443,500,1950,1972]" pageId="7" pageNumber="8">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4CD22ADC2B2EFFA8FDCFD6E655A396F6" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[550,652,1951,1972]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pipatio" status="sp. nov.">
<emphasis id="B9A68D4D2B2EFFA8FDCFD6E655A396F6" box="[550,652,1951,1972]" italics="true" pageId="7" pageNumber="8">H. pipatio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2EFFA8FD7BD6E755E396F6" box="[658,716,1950,1972]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B9A68D4D2B2EFFA8FD7BD6E755E396F6" bold="true" box="[658,716,1950,1972]" pageId="7" pageNumber="8">n. sp.</emphasis>
</taxonomicNameLabel>
; (C) to (E), 
<taxonomicName id="4CD22ADC2B2EFFA8FCA1D6E7533596F6" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[840,1050,1950,1972]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="thomsoni" status="sp. nov.">
<emphasis id="B9A68D4D2B2EFFA8FCA1D6E7533596F6" box="[840,1050,1950,1972]" italics="true" pageId="7" pageNumber="8">Xeropsalta thomsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2EFFA8FBC9D6E7537696F6" box="[1056,1113,1950,1972]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B9A68D4D2B2EFFA8FBC9D6E7537696F6" bold="true" box="[1056,1113,1950,1972]" pageId="7" pageNumber="8">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B2EFFA8FB8CD6E653E396F6" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1125,1228,1950,1972]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="aridula" status="sp. nov.">
<emphasis id="B9A68D4D2B2EFFA8FB8CD6E653E396F6" box="[1125,1228,1950,1972]" italics="true" pageId="7" pageNumber="8">X. aridula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2EFFA8FB3BD6E7522496F6" box="[1234,1291,1950,1972]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B9A68D4D2B2EFFA8FB3BD6E7522496F6" bold="true" box="[1234,1291,1950,1972]" pageId="7" pageNumber="8">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B2EFFA8FAF1D6E652AE96F6" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1304,1409,1951,1972]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rattrayi" status="sp. nov.">
<emphasis id="B9A68D4D2B2EFFA8FAF1D6E652AE96F6" box="[1304,1409,1951,1972]" italics="true" pageId="7" pageNumber="8">X. rattrayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B2EFFA8FA6ED6E7579A9691" pageId="7" pageNumber="8" rank="species">
<emphasis id="B9A68D4D2B2EFFA8FA6ED6E7579A9691" bold="true" pageId="7" pageNumber="8">n. sp.</emphasis>
</taxonomicNameLabel>
, respectively. For clarity, only selected long ribs and one short (intercalary) rib are labelled, as are the timbal plate and apodeme. 5 long ribs are present, only numbers 1 and 5 labelled.
</paragraph>
</caption>
<caption id="DFAD01D72B21FFA7FF7ED66F53C69609" httpUri="https://zenodo.org/record/1221898/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1814,1836]" targetBox="[228,1360,193,1792]" targetPageId="8">
<paragraph id="8B6D515F2B21FFA7FF7ED66F53C69609" blockId="8.[151,1436,1814,1867]" pageId="8" pageNumber="9">
<emphasis id="B9A68D4D2B21FFA7FF7ED66F563D966E" bold="true" box="[151,274,1814,1836]" pageId="8" pageNumber="9">FIGURE 5.</emphasis>
Right fore and hind wings of the five new species described in this paper; (A), (B), 
<emphasis id="B9A68D4D2B21FFA7FB8FD6615273966E" box="[1126,1372,1814,1837]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4CD22ADC2B21FFA7FB8FD66153CB966F" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1126,1252,1816,1837]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="genus" status="gen. nov.">Heremusina</taxonomicName>
. 
<taxonomicName id="4CD22ADC2B21FFA7FB04D66F5273966E" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1261,1372,1814,1836]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="udeoecetes" status="sp. nov.">udeoecetes</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A29530362B21FFA7FA8AD66F52B3966E" box="[1379,1436,1814,1836]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B9A68D4D2B21FFA7FA8AD66F52B3966E" bold="true" box="[1379,1436,1814,1836]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4CD22ADC2B21FFA7FF2BD64F56069609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[194,297,1846,1867]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pipatio" status="sp. nov.">
<emphasis id="B9A68D4D2B21FFA7FF2BD64F56069609" box="[194,297,1846,1867]" italics="true" pageId="8" pageNumber="9">H. pipatio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B21FFA7FEC6D64C56479609" box="[303,360,1845,1867]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B9A68D4D2B21FFA7FEC6D64C56479609" bold="true" box="[303,360,1845,1867]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
; (C) to (E), 
<taxonomicName id="4CD22ADC2B21FFA7FE0DD64C559A9609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[484,693,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="thomsoni" status="sp. nov.">
<emphasis id="B9A68D4D2B21FFA7FE0DD64C559A9609" box="[484,693,1845,1867]" italics="true" pageId="8" pageNumber="9">Xeropsalta thomsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B21FFA7FD55D64C55DA9609" box="[700,757,1845,1867]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B9A68D4D2B21FFA7FD55D64C55DA9609" bold="true" box="[700,757,1845,1867]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B21FFA7FCE8D64F54479609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[769,872,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aridula" status="sp. nov.">
<emphasis id="B9A68D4D2B21FFA7FCE8D64F54479609" box="[769,872,1845,1867]" italics="true" pageId="8" pageNumber="9">X. aridula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B21FFA7FC87D64C54889609" box="[878,935,1845,1867]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B9A68D4D2B21FFA7FC87D64C54889609" bold="true" box="[878,935,1845,1867]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B21FFA7FC5DD64F53329609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[948,1053,1846,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rattrayi" status="sp. nov.">
<emphasis id="B9A68D4D2B21FFA7FC5DD64F53329609" box="[948,1053,1846,1867]" italics="true" pageId="8" pageNumber="9">X. rattrayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B21FFA7FBCDD64C53729609" box="[1060,1117,1845,1867]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B9A68D4D2B21FFA7FBCDD64C53729609" bold="true" box="[1060,1117,1845,1867]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
, respectively.
</paragraph>
</caption>
<caption id="DFAD01D72B20FFA6FF7ED66F54CB9628" httpUri="https://zenodo.org/record/1221900/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1814,1836]" targetBox="[219,1367,193,1790]" targetPageId="9">
<paragraph id="8B6D515F2B20FFA6FF7ED66F54CB9628" blockId="9.[151,1437,1814,1898]" pageId="9" pageNumber="10">
<emphasis id="B9A68D4D2B20FFA6FF7ED66F563B966E" bold="true" box="[151,276,1814,1836]" pageId="9" pageNumber="10">FIGURE 6.</emphasis>
Pygofer and male genitalia, lateral views, respectively, of the five new species described in this paper; (A), (B), 
<taxonomicName id="4CD22ADC2B20FFA6FF7ED64F56A89609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[151,391,1845,1867]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="udeoecetes" status="sp. nov.">
<emphasis id="B9A68D4D2B20FFA6FF7ED64F56A89609" box="[151,391,1845,1867]" italics="true" pageId="9" pageNumber="10">Heremusina udeoecetes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B20FFA6FE64D64C56E89609" box="[397,455,1845,1867]" pageId="9" pageNumber="10" rank="species">
<emphasis id="B9A68D4D2B20FFA6FE64D64C56E89609" bold="true" box="[397,455,1845,1867]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4CD22ADC2B20FFA6FE11D64F55709609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[504,607,1846,1867]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pipatio" status="sp. nov.">
<emphasis id="B9A68D4D2B20FFA6FE11D64F55709609" box="[504,607,1846,1867]" italics="true" pageId="9" pageNumber="10">H. pipatio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B20FFA6FD8CD64C55B19609" box="[613,670,1845,1867]" pageId="9" pageNumber="10" rank="species">
<emphasis id="B9A68D4D2B20FFA6FD8CD64C55B19609" bold="true" box="[613,670,1845,1867]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
; (C) to (E), 
<taxonomicName id="4CD22ADC2B20FFA6FCF3D64C54C39609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[794,1004,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="thomsoni" status="sp. nov.">
<emphasis id="B9A68D4D2B20FFA6FCF3D64C54C39609" box="[794,1004,1845,1867]" italics="true" pageId="9" pageNumber="10">Xeropsalta thomsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B20FFA6FC1BD64C53049609" box="[1010,1067,1845,1867]" pageId="9" pageNumber="10" rank="species">
<emphasis id="B9A68D4D2B20FFA6FC1BD64C53049609" bold="true" box="[1010,1067,1845,1867]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
(see also Pl. 4), 
<taxonomicName id="4CD22ADC2B20FFA6FB3AD64F52159609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1235,1338,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aridula" status="sp. nov.">
<emphasis id="B9A68D4D2B20FFA6FB3AD64F52159609" box="[1235,1338,1845,1867]" italics="true" pageId="9" pageNumber="10">X. aridula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B20FFA6FAA9D64C52569609" box="[1344,1401,1845,1867]" pageId="9" pageNumber="10" rank="species">
<emphasis id="B9A68D4D2B20FFA6FAA9D64C52569609" bold="true" box="[1344,1401,1845,1867]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B20FFA6FA6FD64F57CB9628" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rattrayi" status="sp. nov.">
<emphasis id="B9A68D4D2B20FFA6FA6FD64F57CB9628" italics="true" pageId="9" pageNumber="10">X. rattrayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B20FFA6FF02D62D560B9628" box="[235,292,1876,1898]" pageId="9" pageNumber="10" rank="species">
<emphasis id="B9A68D4D2B20FFA6FF02D62D560B9628" bold="true" box="[235,292,1876,1898]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
Pygofer lengths are, respectively, 1.60, 1.70, 1.86, 1.91, and 1.9 mm.
</paragraph>
</caption>
<caption id="DFAD01D72B23FFA5FF7ED66F56219628" httpUri="https://zenodo.org/record/1221902/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,1814,1836]" targetBox="[226,1361,193,1778]" targetPageId="10">
<paragraph id="8B6D515F2B23FFA5FF7ED66F56219628" blockId="10.[151,1437,1814,1898]" pageId="10" pageNumber="11">
<emphasis id="B9A68D4D2B23FFA5FF7ED66F563D966E" bold="true" box="[151,274,1814,1836]" pageId="10" pageNumber="11">FIGURE 7.</emphasis>
Pygofer and male genitalia, ventral views, respectively, of the five new species described in this paper; (A), (B), 
<taxonomicName id="4CD22ADC2B23FFA5FA6AD66156299609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="udeoecetes" status="sp. nov.">
<emphasis id="B9A68D4D2B23FFA5FA6AD66156299609" italics="true" pageId="10" pageNumber="11">H. udeoecetes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B23FFA5FEE5D64C56699609" box="[268,326,1845,1867]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B9A68D4D2B23FFA5FEE5D64C56699609" bold="true" box="[268,326,1845,1867]" pageId="10" pageNumber="11">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4CD22ADC2B23FFA5FE91D64F56F09609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[376,479,1846,1867]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pipatio" status="sp. nov.">
<emphasis id="B9A68D4D2B23FFA5FE91D64F56F09609" box="[376,479,1846,1867]" italics="true" pageId="10" pageNumber="11">H. pipatio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B23FFA5FE0FD64C550F9609" box="[486,544,1845,1867]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B9A68D4D2B23FFA5FE0FD64C550F9609" bold="true" box="[486,544,1845,1867]" pageId="10" pageNumber="11">n. sp.</emphasis>
</taxonomicNameLabel>
; (C) to (E), 
<taxonomicName id="4CD22ADC2B23FFA5FD74D64F54389609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[669,791,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="thomsoni" status="sp. nov.">
<emphasis id="B9A68D4D2B23FFA5FD74D64F54389609" box="[669,791,1845,1867]" italics="true" pageId="10" pageNumber="11">X. thomsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B23FFA5FCF4D64C54789609" box="[797,855,1845,1867]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B9A68D4D2B23FFA5FCF4D64C54789609" bold="true" box="[797,855,1845,1867]" pageId="10" pageNumber="11">n. sp.</emphasis>
</taxonomicNameLabel>
(see also Pl. 4), 
<taxonomicName id="4CD22ADC2B23FFA5FBE8D64F53469609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1025,1129,1845,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aridula" status="sp. nov.">
<emphasis id="B9A68D4D2B23FFA5FBE8D64F53469609" box="[1025,1129,1845,1867]" italics="true" pageId="10" pageNumber="11">X. aridula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B23FFA5FB86D64C53869609" box="[1135,1193,1845,1867]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B9A68D4D2B23FFA5FB86D64C53869609" bold="true" box="[1135,1193,1845,1867]" pageId="10" pageNumber="11">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B23FFA5FB5CD64F52309609" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1205,1311,1846,1867]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="rattrayi" status="sp. nov.">
<emphasis id="B9A68D4D2B23FFA5FB5CD64F52309609" box="[1205,1311,1846,1867]" italics="true" pageId="10" pageNumber="11">X. rattrayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B23FFA5FACFD64C524F9609" box="[1318,1376,1845,1867]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B9A68D4D2B23FFA5FACFD64C524F9609" bold="true" box="[1318,1376,1845,1867]" pageId="10" pageNumber="11">n. sp.</emphasis>
</taxonomicNameLabel>
Sizes as in Fig. 6.
</paragraph>
</caption>
<caption id="DFAD01D72B22FFA4FF7ED4395467974C" httpUri="https://zenodo.org/record/1221904/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1344,1366]" targetBox="[151,1436,193,1323]" targetPageId="11">
<paragraph id="8B6D515F2B22FFA4FF7ED4395467974C" blockId="11.[151,1436,1344,1550]" pageId="11" pageNumber="12">
<emphasis id="B9A68D4D2B22FFA4FF7ED439563C9414" bold="true" box="[151,275,1344,1366]" pageId="11" pageNumber="12">FIGURE 8.</emphasis>
Map of Central and Northeastern Australia, showing the currently known occurrences of 
<taxonomicName id="4CD22ADC2B22FFA4FB42D43852B39414" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1195,1436,1344,1366]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="udeoecetes" status="sp. nov.">
<emphasis id="B9A68D4D2B22FFA4FB42D43852B39414" box="[1195,1436,1344,1366]" italics="true" pageId="11" pageNumber="12">Heremusina udeoecetes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B22FFA4FF7ED42657FF9437" box="[151,208,1375,1397]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B9A68D4D2B22FFA4FF7ED42657FF9437" bold="true" box="[151,208,1375,1397]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B22FFA4FF34D419566B9437" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[221,324,1376,1397]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pipatio" status="sp. nov.">
<emphasis id="B9A68D4D2B22FFA4FF34D419566B9437" box="[221,324,1376,1397]" italics="true" pageId="11" pageNumber="12">H. pipatio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B22FFA4FEA3D42656AC9437" box="[330,387,1375,1397]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B9A68D4D2B22FFA4FEA3D42656AC9437" bold="true" box="[330,387,1375,1397]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B22FFA4FE79D426554D9437" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[400,610,1375,1397]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="thomsoni" status="sp. nov.">
<emphasis id="B9A68D4D2B22FFA4FE79D426554D9437" box="[400,610,1375,1397]" italics="true" pageId="11" pageNumber="12">Xeropsalta thomsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B22FFA4FD81D426558E9437" box="[616,673,1375,1397]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B9A68D4D2B22FFA4FD81D426558E9437" bold="true" box="[616,673,1375,1397]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4CD22ADC2B22FFA4FD47D419543A9437" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[686,789,1375,1397]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aridula" status="sp. nov.">
<emphasis id="B9A68D4D2B22FFA4FD47D419543A9437" box="[686,789,1375,1397]" italics="true" pageId="11" pageNumber="12">X. aridula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B22FFA4FCF5D426547A9437" box="[796,853,1375,1397]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B9A68D4D2B22FFA4FCF5D426547A9437" bold="true" box="[796,853,1375,1397]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
, and 
<taxonomicName id="4CD22ADC2B22FFA4FC65D41954D99437" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[908,1014,1376,1397]" class="Insecta" family="Cicadidae" genus="Xeropsalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="rattrayi" status="sp. nov.">
<emphasis id="B9A68D4D2B22FFA4FC65D41954D99437" box="[908,1014,1376,1397]" italics="true" pageId="11" pageNumber="12">X. rattrayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B22FFA4FC15D42653199437" box="[1020,1078,1375,1397]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B9A68D4D2B22FFA4FC15D42653199437" bold="true" box="[1020,1078,1375,1397]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
The heavy dot-dash line shows the approximate northern and eastern boundaries of the ecoregion which defines the dominance of deserts and xeric shrublands within inland Australia (based on National Reserve System map, published by the Australian Government). The fine dotted lines are major roads, the names of some prominent towns listed. Lake Eyre and associated ephemoral lakes are shown by the darker shading. The approximate outlines of the modern dunefields comprising much of the Simpson and Strzelecki deserts are shown by the lighter shading, these based on 'Google Earth' images.
</paragraph>
</caption>
<subSubSection id="C3C802D42B22FFA0FF2ED64F539690B6" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12" type="description">
<paragraph id="8B6D515F2B22FFA1FF2ED64F57F5963C" blockId="11.[151,1436,1595,2014]" lastBlockId="14.[151,1437,1496,2025]" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<emphasis id="B9A68D4D2B22FFA4FF2ED64F5640960D" box="[199,367,1846,1871]" italics="true" pageId="11" pageNumber="12">Male genitalia</emphasis>
. Pygofer in ventral view ovoid in shape, with upper pygofer lobes widest points, the shape generally tapered from upper pygofer lobes to base; pygofer with distal shoulders developed; upper lobes moderately well developed, narrow and gently curved in ventral view, set well removed from dorsal beak; gently rounded in lateral view; basal lobes undivided, well developed, partially rounded in lateral view, abutted against pygofer margin; dorsal beak elongated as a pointed apex, forms part of chitinised pygofer. Uncus relatively short, somewhat flattened, median lobe gently curved in ventral view, roughly tongue-shaped. Claspers well developed, restraining aedeagus; prominent in lateral and ventral views, terminal claw with minimal cavity developed ventrally, terminations widely separated; gently rounded inward-facing swelling present on inner margins, apices diverging, in ventral view the widest dimension of claspers. Aedeagus with basal plate in lateral view undulated, depressed on dorsal midline, in dorsal view short and broad, apically broadened with 'ears', much broader than long; basal part of basal plate directed forwards away from thecal shaft, ventral rib fused with basal plate, small functional 'hinge' compressed between theca and basal plate; thecal shaft almost straight and parallel-sided, relatively thickly developed; pseudoparameres present, developed sub-apically, lateral of theca, to which they have fused for at least part of their length, in dorsal view their apices widely separated, diverging throughout their length, in lateral view directed upwards beyond thecal shaft; endotheca not visible; ventral support absent, thecal apex chitinised, thecal subapical cerci, flabellum and conjunctival claws each absent; vesical opening apical on theca.
</paragraph>
<caption id="DFAD01D72B25FFA3FF7ED7F355DC96B0" httpUri="https://zenodo.org/record/1221906/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1674,1696]" targetBox="[291,1294,185,1662]" targetPageId="12">
<paragraph id="8B6D515F2B25FFA3FF7ED7F355DC96B0" blockId="12.[151,1436,1674,2034]" pageId="12" pageNumber="13">
<emphasis id="B9A68D4D2B25FFA3FF7ED7F3563C97E2" bold="true" box="[151,275,1674,1696]" pageId="12" pageNumber="13">FIGURE 9.</emphasis>
<taxonomicName id="4CD22ADC2B25FFA3FEF2D7F356C497E2" baseAuthorityName="Distant" baseAuthorityYear="1911" box="[283,491,1674,1696]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B9A68D4D2B25FFA3FEF2D7F356C497E2" box="[283,491,1674,1696]" italics="true" pageId="12" pageNumber="13">Mugadina marshalli</emphasis>
</taxonomicName>
song. Waveform (A, B, D) and complimentary envelope (C, E) plots, of the ticking song of 
<taxonomicName id="4CD22ADC2B25FFA3FF7ED7D3563097FD" baseAuthorityName="Distant" baseAuthorityYear="1911" box="[151,287,1705,1727]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B9A68D4D2B25FFA3FF7ED7D3563097FD" box="[151,287,1705,1727]" italics="true" pageId="12" pageNumber="13">M. marshalli</emphasis>
</taxonomicName>
, recorded in open net, in grassland near Kaimkillenbun, southeast Queensland (A to C) and Yelarbon, S. Queensland, 
<emphasis id="B9A68D4D2B25FFA3FEF2D7B05675979C" box="[283,346,1737,1758]" italics="true" pageId="12" pageNumber="13">in situ</emphasis>
with parabola in grassland (D, E). Fig. 9A shows an overview plot of the repeated pulses ('ticks'), the pulse repetition rates (PRR) varying between 4.9–5.4 Hz. Figs. 9B, C, are time expanded illustrating details of an individual pulse, showing that when viewed in detail, each actually comprises two dominant (primary) pulses, each followed by a weaker, but clearly separate pulse, the time gaps being 1.9 and 2.8 ms. These are interpreted as secondary pulses, produced by timbal 'OUT' (relaxation) movement. The two initial strong pulses exhibit weaker and almost coalesced pulses, which is suggested to result from the 'firing' of long ribs 2 to 5 (implying that the dominant pulses are produced by the 'IN' movements of timbal 1). The presence of two complete sets of pulses that comprise each 'tick' are inferred to result from a very short but significant delay in the contraction (firing) between the two timbals. Figs. 9D, E, represent time expanded plots from another 
<taxonomicName id="4CD22ADC2B25FFA3FB27D6D9527D96F6" baseAuthorityName="Distant" baseAuthorityYear="1911" box="[1230,1362,1950,1973]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B9A68D4D2B25FFA3FB27D6D9527D96F6" box="[1230,1362,1950,1973]" italics="true" pageId="12" pageNumber="13">M. marshalli</emphasis>
</taxonomicName>
from a separate location, showing the similar arrangement and structures of the pulses, although the gaps between the primary and secondary pulses are significantly longer. Filtered to 1 kHz.
</paragraph>
</caption>
<caption id="DFAD01D72B24FFA2FF7ED66F53CB96A6" httpUri="https://zenodo.org/record/1221908/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1814,1836]" targetBox="[248,1339,193,1793]" targetPageId="13">
<paragraph id="8B6D515F2B24FFA2FF7ED66F53CB96A6" blockId="13.[151,1437,1814,2020]" pageId="13" pageNumber="14">
<emphasis id="B9A68D4D2B24FFA2FF7ED66F560D966E" bold="true" box="[151,290,1814,1836]" pageId="13" pageNumber="14">FIGURE 10.</emphasis>
Waveform (A, B, D) and complementary envelope curves (C, E) of the ticking song of 
<taxonomicName id="4CD22ADC2B24FFA2FB2FD66F525C966F" box="[1222,1395,1814,1837]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="emma">
<emphasis id="B9A68D4D2B24FFA2FB2FD66F525C966F" box="[1222,1395,1814,1837]" italics="true" pageId="13" pageNumber="14">Mugadina emma</emphasis>
</taxonomicName>
, all recordings made near the Fairbairn Dam Rd, ~7 km S.W. of Emerald, eastern central Queensland, recorded 'in situ' in an open net in grassland. Fig. 10A presents an overview segment showing the multiple repeated 'ticks', the PRR varying between 2.5 to 4.6 Hz. As in the 
<taxonomicName id="4CD22ADC2B24FFA2FEA6D60A56FB96CA" baseAuthorityName="Distant" baseAuthorityYear="1911" box="[335,468,1906,1928]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B9A68D4D2B24FFA2FEA6D60A56FB96CA" box="[335,468,1906,1928]" italics="true" pageId="13" pageNumber="14">M. marshalli</emphasis>
</taxonomicName>
song, each 'tick' comprises a set of two pulses, with each initial 'primary' pulse followed by a weaker secondary pulse, the intervals between the two pulses being 1.4–3.4 ms, slightly variable, although similar to the 
<taxonomicName id="4CD22ADC2B24FFA2FA69D6EB57D79684" baseAuthorityName="Distant" baseAuthorityYear="1911" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B9A68D4D2B24FFA2FA69D6EB57D79684" italics="true" pageId="13" pageNumber="14">M. marshalli</emphasis>
</taxonomicName>
data. The detailed pulse structures within the primary and secondary pulses are also similar to those of 
<taxonomicName id="4CD22ADC2B24FFA2FAFAD6C852BA9684" baseAuthorityName="Distant" baseAuthorityYear="1911" box="[1299,1429,1968,1990]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B9A68D4D2B24FFA2FAFAD6C852BA9684" box="[1299,1429,1968,1990]" italics="true" pageId="13" pageNumber="14">M. marshalli</emphasis>
</taxonomicName>
, and the two sets of pulses are similarly interpreted as time offsets in the firing of the two timbals. Unfiltered.
</paragraph>
</caption>
<caption id="DFAD01D72B27FFA1FF7ED47B521094F3" httpUri="https://zenodo.org/record/1221910/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1282,1304]" targetBox="[151,1436,193,1261]" targetPageId="14">
<paragraph id="8B6D515F2B27FFA1FF7ED47B521094F3" blockId="14.[151,1436,1282,1457]" pageId="14" pageNumber="15">
<emphasis id="B9A68D4D2B27FFA1FF7ED47B5630945A" bold="true" box="[151,287,1282,1304]" pageId="14" pageNumber="15">FIGURE 11.</emphasis>
Pulse repetition rates (Hz) versus durations (ms) between the doublet primary pulses in the songs of 
<taxonomicName id="4CD22ADC2B27FFA1FADDD47B57D79475" baseAuthorityName="Distant" baseAuthorityYear="1911" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B9A68D4D2B27FFA1FADDD47B57D79475" italics="true" pageId="14" pageNumber="15">Mugadina marshalli</emphasis>
</taxonomicName>
and 
<taxonomicName id="4CD22ADC2B27FFA1FEC1D45B56A99475" box="[296,390,1314,1335]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="emma">
<emphasis id="B9A68D4D2B27FFA1FEC1D45B56A99475" box="[296,390,1314,1335]" italics="true" pageId="14" pageNumber="15">M. emma</emphasis>
</taxonomicName>
. These data are based on multiple recordings, from multiple locations, at multiple dates, including field and container recordings (recordings by AE and LWP). The PRR of the 
<taxonomicName id="4CD22ADC2B27FFA1FC61D43953229417" baseAuthorityName="Distant" baseAuthorityYear="1911" box="[904,1037,1343,1365]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B9A68D4D2B27FFA1FC61D43953229417" box="[904,1037,1343,1365]" italics="true" pageId="14" pageNumber="15">M. marshalli</emphasis>
</taxonomicName>
songs are consistently higher than the 
<taxonomicName id="4CD22ADC2B27FFA1FF7ED42657C19436" box="[151,238,1375,1396]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="emma">
<emphasis id="B9A68D4D2B27FFA1FF7ED42657C19436" box="[151,238,1375,1396]" italics="true" pageId="14" pageNumber="15">M.emma</emphasis>
</taxonomicName>
songs, with minimal overlap of data. The inset shows 
<emphasis id="B9A68D4D2B27FFA1FCF5D42454679431" bold="true" box="[796,840,1373,1395]" italics="true" pageId="14" pageNumber="15">only</emphasis>
PRR data from recordings made 2 km west of Macalister, southeastern Queensland, where both species were found to occur together. These recordings are container recordings made under identical conditions. They exhibit a marked separation of PRR between the two species at this overlap location.
</paragraph>
</caption>
<paragraph id="8B6D515F2B27FFA0FF2ED6F1539690B6" blockId="14.[151,1437,1496,2025]" lastBlockId="15.[151,1436,151,501]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B9A68D4D2B27FFA1FF2ED6F1533596E3" bold="true" box="[199,1050,1928,1953]" pageId="14" pageNumber="15">
Distinguishing characters between 
<taxonomicName id="4CD22ADC2B27FFA1FD9DD6F155C396E3" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[628,748,1928,1953]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B27FFA1FD9DD6F155C396E3" bold="true" box="[628,748,1928,1953]" italics="true" pageId="14" pageNumber="15">Mugadina</emphasis>
</taxonomicName>
and 
<taxonomicName id="4CD22ADC2B27FFA1FCC7D6F1549296E3" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[814,957,1928,1953]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B27FFA1FCC7D6F1549296E3" bold="true" box="[814,957,1928,1953]" italics="true" pageId="14" pageNumber="15">Heremusina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B27FFA1FC21D6F1533596E3" box="[968,1050,1928,1953]" pageId="14" pageNumber="15" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
The above listing of the generic characters of 
<taxonomicName id="4CD22ADC2B27FFA1FEDFD6D756919687" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[310,446,1966,1989]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B27FFA1FEDFD6D756919687" box="[310,446,1966,1989]" italics="true" pageId="14" pageNumber="15">Heremusina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B27FFA1FE23D6D455339684" box="[458,540,1965,1990]" pageId="14" pageNumber="15" rank="genus">
<emphasis id="B9A68D4D2B27FFA1FE23D6D455339684" bold="true" box="[458,540,1965,1990]" pageId="14" pageNumber="15">n. gen.</emphasis>
</taxonomicNameLabel>
extensively follows those defined by 
<bibRefCitation id="EF432CAE2B27FFA1FC09D6D453A49687" box="[992,1163,1965,1990]" pageId="14" pageNumber="15" refId="ref21391" refString="Moulds, M. S. (2012) A review of the genera of Australian cicadas (Hemiptera: Cicadoidea). Zootaxa, 3287, 1 - 262." type="journal article">Moulds (2012)</bibRefCitation>
for 
<taxonomicName id="4CD22ADC2B27FFA1FB2BD6D4521A9684" authorityName="Ewart" authorityYear="2018" box="[1218,1333,1965,1990]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B9A68D4D2B27FFA1FB2BD6D4521A9684" box="[1218,1333,1965,1990]" italics="true" pageId="14" pageNumber="15">Mugadina</emphasis>
</taxonomicName>
, thereby emphasising the closeness of many of these characters to those of 
<taxonomicName id="4CD22ADC2B27FFA1FC72D6A9532196AB" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[923,1038,2000,2025]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B9A68D4D2B27FFA1FC72D6A9532196AB" box="[923,1038,2000,2025]" italics="true" pageId="14" pageNumber="15">Mugadina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B27FFA1FBF1D6A8531396AB" box="[1048,1084,2001,2025]" pageId="14" pageNumber="15" sensu="stricto">s.s.</taxonomicNameLabel>
Although the lateral abdomen shapes of 
<taxonomicName id="4CD22ADC2B26FFA0FEEFD1EE565691F2" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[262,377,151,176]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B26FFA0FEEFD1EE565691F2" box="[262,377,151,176]" italics="true" pageId="15" pageNumber="16">Mugadina</emphasis>
</taxonomicName>
and 
<taxonomicName id="4CD22ADC2B26FFA0FE46D1E0551891F2" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[431,567,153,176]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B26FFA0FE46D1E0551891F2" box="[431,567,153,176]" italics="true" pageId="15" pageNumber="16">Heremusina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B26FFA0FDD7D1EE55A291F2" box="[574,653,151,176]" pageId="15" pageNumber="16" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FDD7D1EE55A291F2" bold="true" box="[574,653,151,176]" pageId="15" pageNumber="16">n. gen.</emphasis>
</taxonomicNameLabel>
are somewhat similar (
<figureCitation id="13E94DDA2B26FFA0FC7BD1EE533B91ED" box="[914,1044,151,176]" captionStart="FIGURE 1" captionStartId="4.[151,250,1864,1886]" captionTargetBox="[283,1306,215,1807]" captionTargetId="figure@4.[272,1315,215,1815]" captionTargetPageId="4" captionText="FIGURE 1. Mugadina emma and M. marshalli, comparison of selected morphological characters of both species. (A), (B), lateral views of abdomen. (C), (D), lateral views of male genitalia, both lengths 1.9 mm. (E), (F), ventral views of same genitalia; (G, H), timbals; dorsal to right, and (I), (J) left opercula. Figs. C, D based on specimens from 4 km north of Amby, southern Queensland and Figs. D, F from Lake Broadwater, southeast Queensland. Figs. A, G, I, based on specimens from Boonah, southeastern Queensland and Figs. B,H, J from Drillham, southern Queensland.The scale bars represent 1 mm." httpUri="https://zenodo.org/record/1221890/files/figure.png" pageId="15" pageNumber="16">Figs. 1A, B</figureCitation>
and 
<figureCitation id="13E94DDA2B26FFA0FBA0D1E153A191F2" box="[1097,1166,152,176]" captionStart="FIGURE2" captionStartId="5.[151,250,1814,1836]" captionTargetBox="[211,1376,193,1783]" captionTargetId="figure@5.[211,1376,193,1793]" captionTargetPageId="5" captionText="FIGURE2. Lateralabdomen views of the five new species described in thispaper;(A), (B), Heremusina udeoecetes n. sp.and H. pipatio n. sp.; (C) to (E), Xeropsalta thomsoni n. sp., X. aridula n. sp., X.rattrayi n. sp., respectively." httpUri="https://zenodo.org/record/1221892/files/figure.png" pageId="15" pageNumber="16">2A, B</figureCitation>
), significant differences are seen in the general shapes of the genitalia (cf. 
<figureCitation id="13E94DDA2B26FFA0FD37D1C554739196" box="[734,860,188,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1864,1886]" captionTargetBox="[283,1306,215,1807]" captionTargetId="figure@4.[272,1315,215,1815]" captionTargetPageId="4" captionText="FIGURE 1. Mugadina emma and M. marshalli, comparison of selected morphological characters of both species. (A), (B), lateral views of abdomen. (C), (D), lateral views of male genitalia, both lengths 1.9 mm. (E), (F), ventral views of same genitalia; (G, H), timbals; dorsal to right, and (I), (J) left opercula. Figs. C, D based on specimens from 4 km north of Amby, southern Queensland and Figs. D, F from Lake Broadwater, southeast Queensland. Figs. A, G, I, based on specimens from Boonah, southeastern Queensland and Figs. B,H, J from Drillham, southern Queensland.The scale bars represent 1 mm." httpUri="https://zenodo.org/record/1221890/files/figure.png" pageId="15" pageNumber="16">Figs. 1C–F</figureCitation>
, and 
<figureCitation id="13E94DDA2B26FFA0FC75D1C553309197" box="[924,1055,188,213]" captionStart="FIGURE 6" captionStartId="9.[151,250,1814,1836]" captionTargetBox="[219,1367,193,1790]" captionTargetId="figure@9.[219,1367,193,1793]" captionTargetPageId="9" captionText="FIGURE 6. Pygofer and male genitalia, lateral views, respectively, of the five new species described in this paper; (A), (B), Heremusina udeoecetes n. sp. and H. pipatio n. sp.; (C) to (E), Xeropsalta thomsoni n. sp. (see also Pl.4), X.aridula n. sp., X. rattrayi n. sp. Pygofer lengths are, respectively, 1.60, 1.70, 1.86, 1.91, and 1.9 mm." httpUri="https://zenodo.org/record/1221900/files/figure.png" pageId="15" pageNumber="16">Figs. 6A–B</figureCitation>
and 
<figureCitation id="13E94DDA2B26FFA0FBB2D1C5538E9196" box="[1115,1185,188,212]" captionStart="FIGURE7" captionStartId="10.[151,250,1814,1836]" captionTargetBox="[226,1361,193,1778]" captionTargetId="figure@10.[226,1361,193,1793]" captionTargetPageId="10" captionText="FIGURE7. Pygofer and male genitalia,ventral views, respectively,of the five new species described in thispaper; (A), (B),H. udeoecetes n. sp. and H.pipatio n. sp.; (C) to (E), X. thomsoni n. sp. (see also Pl. 4), X. aridula n. sp., X. rattrayi n. sp. Sizes as in Fig. 6." httpUri="https://zenodo.org/record/1221902/files/figure.png" pageId="15" pageNumber="16">7A–B</figureCitation>
), the pygofer outlines being more elongated in lateral and ventral views in 
<taxonomicName id="4CD22ADC2B26FFA0FD01D198545F91BA" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[744,880,225,248]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B26FFA0FD01D198545F91BA" box="[744,880,225,248]" italics="true" pageId="15" pageNumber="16">Heremusina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B26FFA0FC9ED1A654E991BA" box="[887,966,223,248]" pageId="15" pageNumber="16" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FC9ED1A654E991BA" bold="true" box="[887,966,223,248]" pageId="15" pageNumber="16">n. gen.</emphasis>
</taxonomicNameLabel>
The dorsal beaks are more elongated and gently curved in 
<taxonomicName id="4CD22ADC2B26FFA0FE8BD07C56C3905E" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[354,492,261,284]" class="Insecta" family="Cicadidae" genus="Herumusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B26FFA0FE8BD07C56C3905E" box="[354,492,261,284]" italics="true" pageId="15" pageNumber="16">Herumusina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B26FFA0FE11D07D5564905F" box="[504,587,260,285]" pageId="15" pageNumber="16" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FE11D07D5564905F" bold="true" box="[504,587,260,285]" pageId="15" pageNumber="16">n. gen.</emphasis>
</taxonomicNameLabel>
; the claspers exhibit sharply diverging distal terminations in 
<taxonomicName id="4CD22ADC2B26FFA0FACBD07D52BA905F" authorityName="Ewart" authorityYear="2018" box="[1314,1429,260,285]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FACBD07D52BA905F" box="[1314,1429,260,285]" italics="true" pageId="15" pageNumber="16">Mugadina</emphasis>
</taxonomicName>
, contrasting with 
<taxonomicName id="4CD22ADC2B26FFA0FEBED05056F09002" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[343,479,297,320]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="47" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B26FFA0FEBED05056F09002" box="[343,479,297,320]" italics="true" pageId="15" pageNumber="16">Heremusina</emphasis>
</taxonomicName>
; marked differences in opercula outline shapes are evident (
<figureCitation id="13E94DDA2B26FFA0FB7AD05E5229907D" box="[1171,1286,295,320]" captionStart="FIGURE 1" captionStartId="4.[151,250,1864,1886]" captionTargetBox="[283,1306,215,1807]" captionTargetId="figure@4.[272,1315,215,1815]" captionTargetPageId="4" captionText="FIGURE 1. Mugadina emma and M. marshalli, comparison of selected morphological characters of both species. (A), (B), lateral views of abdomen. (C), (D), lateral views of male genitalia, both lengths 1.9 mm. (E), (F), ventral views of same genitalia; (G, H), timbals; dorsal to right, and (I), (J) left opercula. Figs. C, D based on specimens from 4 km north of Amby, southern Queensland and Figs. D, F from Lake Broadwater, southeast Queensland. Figs. A, G, I, based on specimens from Boonah, southeastern Queensland and Figs. B,H, J from Drillham, southern Queensland.The scale bars represent 1 mm." httpUri="https://zenodo.org/record/1221890/files/figure.png" pageId="15" pageNumber="16">
Figs. 
<date id="FF6C779F2B26FFA0FB3DD05153C29002" box="[1236,1261,296,320]" pageId="15" pageNumber="16">1I</date>
, J
</figureCitation>
compared to 
<figureCitation id="13E94DDA2B26FFA0FF7ED035560B9026" box="[151,292,332,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1814,1836]" captionTargetBox="[217,1369,193,1775]" captionTargetId="figure@6.[217,1369,193,1793]" captionTargetPageId="6" captionText="FIGURE 3. Male opercula, all left opercula, of the five new species described in this paper; (A), (B), Heremusina udeoecetes n. sp. and H. pipatio n. sp.; (C) to (E), Xeropsalta thomsoni n. sp., X. aridula n. sp., X. rattrayi n. sp., respectively. Opercula nomenclature shown in (E)." httpUri="https://zenodo.org/record/1221894/files/figure.png" pageId="15" pageNumber="16">Figs. 3A, B</figureCitation>
), noting the broader and more blunted and rounded terminations (between the medial and distal margins) of the 
<taxonomicName id="4CD22ADC2B26FFA0FEA0D00856FE90CA" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[329,465,369,392]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B26FFA0FEA0D00856FE90CA" box="[329,465,369,392]" italics="true" pageId="15" pageNumber="16">Heremusina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B26FFA0FE30D016550790CA" box="[473,552,367,392]" pageId="15" pageNumber="16" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FE30D016550790CA" bold="true" box="[473,552,367,392]" pageId="15" pageNumber="16">n. gen.</emphasis>
</taxonomicNameLabel>
opercula; the intercalary timbal ribs of 
<taxonomicName id="4CD22ADC2B26FFA0FC0FD008534190CA" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[998,1134,369,392]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B26FFA0FC0FD008534190CA" box="[998,1134,369,392]" italics="true" pageId="15" pageNumber="16">Heremusina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B26FFA0FB9CD01653EB90CA" box="[1141,1220,367,392]" pageId="15" pageNumber="16" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FB9CD01653EB90CA" bold="true" box="[1141,1220,367,392]" pageId="15" pageNumber="16">n. gen.</emphasis>
</taxonomicNameLabel>
are unusually thin, and the ventral terminations of long ribs 1 to 4 are clearly separated in 
<taxonomicName id="4CD22ADC2B26FFA0FC1CD0EC535290EE" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1013,1149,405,428]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B26FFA0FC1CD0EC535290EE" box="[1013,1149,405,428]" italics="true" pageId="15" pageNumber="16">Heremusina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B26FFA0FB60D0ED53F390EF" box="[1161,1244,404,429]" pageId="15" pageNumber="16" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FB60D0ED53F390EF" bold="true" box="[1161,1244,404,429]" pageId="15" pageNumber="16">n. gen.</emphasis>
</taxonomicNameLabel>
as compared to 
<taxonomicName id="4CD22ADC2B26FFA0FF7ED0CE56259092" authorityName="Ewart" authorityYear="2018" box="[151,266,439,464]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FF7ED0CE56259092" box="[151,266,439,464]" italics="true" pageId="15" pageNumber="16">Mugadina</emphasis>
</taxonomicName>
(
<figureCitation id="13E94DDA2B26FFA0FEF3D0CE56B4908D" box="[282,411,439,464]" captionStart="FIGURE 1" captionStartId="4.[151,250,1864,1886]" captionTargetBox="[283,1306,215,1807]" captionTargetId="figure@4.[272,1315,215,1815]" captionTargetPageId="4" captionText="FIGURE 1. Mugadina emma and M. marshalli, comparison of selected morphological characters of both species. (A), (B), lateral views of abdomen. (C), (D), lateral views of male genitalia, both lengths 1.9 mm. (E), (F), ventral views of same genitalia; (G, H), timbals; dorsal to right, and (I), (J) left opercula. Figs. C, D based on specimens from 4 km north of Amby, southern Queensland and Figs. D, F from Lake Broadwater, southeast Queensland. Figs. A, G, I, based on specimens from Boonah, southeastern Queensland and Figs. B,H, J from Drillham, southern Queensland.The scale bars represent 1 mm." httpUri="https://zenodo.org/record/1221890/files/figure.png" pageId="15" pageNumber="16">Figs. 1G, H</figureCitation>
compared to 
<figureCitation id="13E94DDA2B26FFA0FDDFD0CE55959092" box="[566,698,439,464]" captionStart="FIGURE 4" captionStartId="7.[151,250,1919,1941]" captionTargetBox="[163,1417,193,1911]" captionTargetId="figure@7.[163,1427,193,1911]" captionTargetPageId="7" captionText="FIGURE 4. Left timbal, posterior margin to right, dorsal margin at top, of the five new species described in this paper; (A), (B), Heremusina udeoecetes n. sp. and H.pipatio n. sp.; (C) to (E), Xeropsalta thomsoni n. sp., X.aridula n. sp., X. rattrayi n. sp., respectively. For clarity, only selected long ribs and one short (intercalary) rib are labelled, as are the timbal plate and apodeme.5 long ribs are present, only numbers 1 and 5 labelled." httpUri="https://zenodo.org/record/1221896/files/figure.png" pageId="15" pageNumber="16">Figs. 4A, B</figureCitation>
); the 4 hind wing apical cells in 
<taxonomicName id="4CD22ADC2B26FFA0FBC3D0C0539D9092" authority="Ewart, 2018" authorityName="Ewart" authorityYear="2018" box="[1066,1202,441,464]" class="Insecta" family="Cicadidae" genus="Heremusina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9A68D4D2B26FFA0FBC3D0C0539D9092" box="[1066,1202,441,464]" italics="true" pageId="15" pageNumber="16">Heremusina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A29530362B26FFA0FB53D0CE52269092" box="[1210,1289,439,464]" pageId="15" pageNumber="16" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FB53D0CE52269092" bold="true" box="[1210,1289,439,464]" pageId="15" pageNumber="16">n. gen.</emphasis>
</taxonomicNameLabel>
compared to the 
<quantity id="4C2AFCBA2B26FFA0FF29D0A5563290B7" box="[192,285,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="15" pageNumber="16" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4 to 5 in</quantity>
<taxonomicName id="4CD22ADC2B26FFA0FECDD0A556B890B7" authorityName="Ewart" authorityYear="2018" box="[292,407,476,501]" class="Insecta" family="Cicadidae" genus="Mugadina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B9A68D4D2B26FFA0FECDD0A556B890B7" box="[292,407,476,501]" italics="true" pageId="15" pageNumber="16">Mugadina</emphasis>
</taxonomicName>
. As shown below, the calling songs have markedly differing structures.
</paragraph>
</subSubSection>
</treatment>
</document>