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<document id="C1C58324CF3E07A088C74EEC84172B80" ID-DOI="10.11646/zootaxa.4039.3.3" ID-GBIF-Dataset="0ad500ca-d193-4331-90af-70a6018a5cfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="238528" ID-ZooBank="87A108DC-6967-4CAD-8F6C-DAF7ADD622CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461331934834" checkinUser="plazi" docAuthor="Irmler, Ulrich" docDate="2015" docId="07009178FF88BB22FF4886633ABB2BA7" docLanguage="en" docName="zt04039p444.pdf" docOrigin="Zootaxa 4039 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thoracophorus exilis Erichson 1840" docType="treatment" docVersion="7" lastPageNumber="438" masterDocId="FB39E900FF8FBB25FFDF840938402C7E" masterDocTitle="Four new species of the genus Thoracophorus Motschulsky, 1857 for the Neotropical region (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="444" masterPageNumber="431" pageNumber="438" updateTime="1698618579127" updateUser="plazi">
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<mods:title id="F8DEA2B87CA861DF74C9B6C0479AAF48">Four new species of the genus Thoracophorus Motschulsky, 1857 for the Neotropical region (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
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<mods:namePart id="4F36F15D041F7CA41718AB6081B4ECF1">Irmler, Ulrich</mods:namePart>
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<mods:date id="823FF2120A707FEFCCAD2A84144BB7C3">2015</mods:date>
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<treatment id="07009178FF88BB22FF4886633ABB2BA7" ID-DOI="http://doi.org/10.5281/zenodo.6107936" ID-GBIF-Taxon="119644042" ID-Zenodo-Dep="6107936" LSID="urn:lsid:plazi:treatment:07009178FF88BB22FF4886633ABB2BA7" httpUri="http://treatment.plazi.org/id/07009178FF88BB22FF4886633ABB2BA7" lastPageNumber="438" pageId="7" pageNumber="438">
<subSubSection id="C7B373E5FF88BB22FF48866339372ED8" pageId="7" pageNumber="438" type="nomenclature">
<paragraph id="8F16206EFF88BB22FF4886633A2D2EFA" blockId="7.[151,621,618,678]" box="[151,621,618,644]" pageId="7" pageNumber="438">
<heading id="D45E9702FF88BB22FF4886633A2D2EFA" bold="true" box="[151,621,618,644]" fontSize="11" level="1" pageId="7" pageNumber="438" reason="1">
<taxonomicName id="48A95BEDFF88BB22FF4886633A2D2EFA" ID-CoL="56KC6" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[151,621,618,644]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="BDDDFC7CFF88BB22FF4886633A2D2EFA" bold="true" box="[151,621,618,644]" pageId="7" pageNumber="438">
<emphasis id="BDDDFC7CFF88BB22FF48866339D82EFA" bold="true" box="[151,408,618,644]" italics="true" pageId="7" pageNumber="438">Thoracophorus exilis</emphasis>
(
<bibRefCitation id="EB385D9FFF88BB22FE7786633A252EFA" author="Erichson" box="[424,613,618,644]" pageId="7" pageNumber="438" refString="Erichson, W. F. (1840) Genera et species Staphylinorum Insectorum Coleopterorum Familiae. Morin, Berlin, 954 pp. http: // dx. doi. org / 10.5962 / bhl. title. 59644" type="book" year="1840">Erichson, 1840</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8F16206EFF88BB22FF48868439372ED8" blockId="7.[151,621,618,678]" box="[151,375,653,678]" pageId="7" pageNumber="438">
<figureCitation id="17923CEBFF88BB22FF48868438A32ED8" box="[151,227,653,678]" captionStart="FIGURES 4 8" captionStartId="4.[151,264,1958,1981]" captionTargetBox="[171,1410,614,1931]" captionTargetId="figure@4.[162,1422,603,1945]" captionTargetPageId="4" captionText="FIGURES 4 8. Thoracophorus struyvei (4); T. perplexus (5); T. proximus (6); T. exilis (7); T. aequalis (8); aedeagus in lateral aspect (A), antenna (B), shape of temples (C); scale bar: 0.1 mm." httpUri="https://zenodo.org/record/238530/files/figure.png" pageId="7" pageNumber="438">Figs. 7</figureCitation>
A, B, C, 10A
</paragraph>
</subSubSection>
<subSubSection id="C7B373E5FF88BB22FF4886DC3C872E90" box="[151,1223,725,750]" pageId="7" pageNumber="438" type="materials_examined">
<paragraph id="8F16206EFF88BB22FF4886DC3C872E90" blockId="7.[151,1437,725,2010]" box="[151,1223,725,750]" pageId="7" pageNumber="438">
<emphasis id="BDDDFC7CFF88BB22FF4886DC39FF2E90" bold="true" box="[151,447,725,750]" pageId="7" pageNumber="438">
<typeStatus id="50129ECCFF88BB22FF4886DC38922E90" box="[151,210,725,750]" pageId="7" pageNumber="438">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="50129ECCFF88BB22FE1A86DF3A1D2E90" box="[453,605,726,750]" pageId="7" pageNumber="438" type="holotype">HOLOTYPE</typeStatus>
. male,
<emphasis id="BDDDFC7CFF88BB22FD7386DC3B712E90" bold="true" box="[684,817,725,750]" pageId="7" pageNumber="438">St. Thomas</emphasis>
without more information (
<collectionCode id="E9B8B8ABFF88BB22FBB686DF3CFF2E90" box="[1129,1215,726,750]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="7" pageNumber="438">ZMUH</collectionCode>
)
</paragraph>
</subSubSection>
<subSubSection id="C7B373E5FF88BB22FF1886F03A842804" pageId="7" pageNumber="438" type="diagnosis">
<paragraph id="8F16206EFF88BB22FF1886F03A842804" blockId="7.[151,1437,725,2010]" pageId="7" pageNumber="438">
<emphasis id="BDDDFC7CFF88BB22FF1886F039012F6C" bold="true" box="[199,321,761,786]" pageId="7" pageNumber="438">Diagnosis.</emphasis>
The species extremely resembles
<taxonomicName id="48A95BEDFF88BB22FD1686F33BA62F6C" authority="Irmler, 1985" authorityName="Irmler" authorityYear="1985" box="[713,998,761,786]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="BDDDFC7CFF88BB22FD1686F33B0D2F6F" box="[713,845,762,785]" italics="true" pageId="7" pageNumber="438">T. proximus</emphasis>
<bibRefCitation id="EB385D9FFF88BB22FC8986F03BA62F6C" author="Irmler" box="[854,998,761,786]" pageId="7" pageNumber="438" refString="Irmler, U. (1985) Neue Arten der Gattungen Aneucamptus und Thoracophorus (Col., Staphylinidae) aus der Neotropis. Entomologische Blatter, 81, 41 - 58." type="journal article" year="1985">Irmler, 1985</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="48A95BEDFF88BB22FC2D86F33C282F6F" box="[1010,1128,762,785]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="struyvei" status="sp. nov.">
<emphasis id="BDDDFC7CFF88BB22FC2D86F33C282F6F" box="[1010,1128,762,785]" italics="true" pageId="7" pageNumber="438">T. struyvei</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF88BB22FBAE86F03CF12F6C" bold="true" box="[1137,1201,761,786]" pageId="7" pageNumber="438">
<taxonomicNameLabel id="A6EE4107FF88BB22FBAE86F03CF12F6C" box="[1137,1201,761,786]" pageId="7" pageNumber="438" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="48A95BEDFF88BB22FB2C86F33D3D2F6C" box="[1267,1405,761,786]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="perplexus" status="sp. nov.">
<emphasis id="BDDDFC7CFF88BB22FB2C86F33D3D2F6C" box="[1267,1405,761,786]" italics="true" pageId="7" pageNumber="438">T. perplexus</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF88BB22FA5A86F038F82F48" bold="true" pageId="7" pageNumber="438">
<taxonomicNameLabel id="A6EE4107FF88BB22FA5A86F038F82F48" pageId="7" pageNumber="438" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. It can be mainly identified by the differences of the aedeagus. In contrast to these three species, the paramera do not or only slightly exceed the apex of the central lobe. In lateral aspect, the inner edge of the apical lobe is slightly sinuate such as in
<taxonomicName id="48A95BEDFF88BB22FE63876F3A042F00" box="[444,580,869,894]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="perplexus" status="sp. nov.">
<emphasis id="BDDDFC7CFF88BB22FE63876F3A042F00" box="[444,580,869,894]" italics="true" pageId="7" pageNumber="438">T. perplexus</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF88BB22FD95876C3AC82F00" bold="true" box="[586,648,869,894]" pageId="7" pageNumber="438">
<taxonomicNameLabel id="A6EE4107FF88BB22FD95876C3AC82F00" box="[586,648,869,894]" pageId="7" pageNumber="438" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, whereas the inner edges of the apical lobes are straight in
<taxonomicName id="48A95BEDFF88BB22FAC5876F39612FDC" authority="Irmler, 1985" authorityName="Irmler" authorityYear="1985" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="BDDDFC7CFF88BB22FAC5876F3DDC2F03" box="[1306,1436,870,893]" italics="true" pageId="7" pageNumber="438">T. proximus</emphasis>
<bibRefCitation id="EB385D9FFF88BB22FF48878039612FDC" author="Irmler" box="[151,289,905,930]" pageId="7" pageNumber="438" refString="Irmler, U. (1985) Neue Arten der Gattungen Aneucamptus und Thoracophorus (Col., Staphylinidae) aus der Neotropis. Entomologische Blatter, 81, 41 - 58." type="journal article" year="1985">Irmler, 1985</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="48A95BEDFF88BB22FE89878339892FDF" box="[342,457,906,929]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="struyvei" status="sp. nov.">
<emphasis id="BDDDFC7CFF88BB22FE89878339892FDF" box="[342,457,906,929]" italics="true" pageId="7" pageNumber="438">T. struyvei</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF88BB22FE0F87803A4E2FDC" bold="true" box="[464,526,905,930]" pageId="7" pageNumber="438">
<taxonomicNameLabel id="A6EE4107FF88BB22FE0F87803A4E2FDC" box="[464,526,905,930]" pageId="7" pageNumber="438" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Moreover, the shape of the apical transparent plate of the paramera differs among the three species: small plate in
<taxonomicName id="48A95BEDFF88BB22FDC187A73B0B2FB8" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[542,843,941,966]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="BDDDFC7CFF88BB22FDC187A73A3A2FB8" box="[542,634,941,966]" italics="true" pageId="7" pageNumber="438">T. exilis</emphasis>
(
<bibRefCitation id="EB385D9FFF88BB22FD5187A43B032FB8" author="Erichson" box="[654,835,941,966]" pageId="7" pageNumber="438" refString="Erichson, W. F. (1840) Genera et species Staphylinorum Insectorum Coleopterorum Familiae. Morin, Berlin, 954 pp. http: // dx. doi. org / 10.5962 / bhl. title. 59644" type="book" year="1840">Erichson, 1840</bibRefCitation>
)
</taxonomicName>
, large asymmetric plate in
<taxonomicName id="48A95BEDFF88BB22FB4A87A73D4F2FBB" box="[1173,1295,942,965]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="struyvei" status="sp. nov.">
<emphasis id="BDDDFC7CFF88BB22FB4A87A73D4F2FBB" box="[1173,1295,942,965]" italics="true" pageId="7" pageNumber="438">T. struyvei</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF88BB22FAC487A43D1F2FB8" bold="true" box="[1307,1375,941,966]" pageId="7" pageNumber="438">
<taxonomicNameLabel id="A6EE4107FF88BB22FAC487A43D1F2FB8" box="[1307,1375,941,966]" pageId="7" pageNumber="438" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and intermediate large plate in
<taxonomicName id="48A95BEDFF88BB22FE1687DB3AA52F94" authority="Irmler, 1985" authorityName="Irmler" authorityYear="1985" box="[457,741,977,1002]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="BDDDFC7CFF88BB22FE1687DB3A0D2F97" box="[457,589,978,1001]" italics="true" pageId="7" pageNumber="438">T. proximus</emphasis>
<bibRefCitation id="EB385D9FFF88BB22FD8A87D83AA52F94" author="Irmler" box="[597,741,977,1002]" pageId="7" pageNumber="438" refString="Irmler, U. (1985) Neue Arten der Gattungen Aneucamptus und Thoracophorus (Col., Staphylinidae) aus der Neotropis. Entomologische Blatter, 81, 41 - 58." type="journal article" year="1985">Irmler, 1985</bibRefCitation>
</taxonomicName>
. Regarding the
<typeStatus id="50129ECCFF88BB22FC4587D83BBC2F94" box="[922,1020,977,1002]" pageId="7" pageNumber="438" type="holotype">holotype</typeStatus>
specimen,
<taxonomicName id="48A95BEDFF88BB22FBA287DB3DDC2F94" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[1149,1436,977,1002]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="BDDDFC7CFF88BB22FBA287DB3C962F94" box="[1149,1238,977,1002]" italics="true" pageId="7" pageNumber="438">T. exilis</emphasis>
(
<bibRefCitation id="EB385D9FFF88BB22FB3987D83DD42F94" author="Erichson" box="[1254,1428,977,1002]" pageId="7" pageNumber="438" refString="Erichson, W. F. (1840) Genera et species Staphylinorum Insectorum Coleopterorum Familiae. Morin, Berlin, 954 pp. http: // dx. doi. org / 10.5962 / bhl. title. 59644" type="book" year="1840">Erichson, 1840</bibRefCitation>
)
</taxonomicName>
seems to be lighter than the other three species that are darker brown or partly orange brown. The elytral carinae are more distinct than in
<taxonomicName id="48A95BEDFF88BB22FE1D80133A0E284C" box="[450,590,1049,1074]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="perplexus" status="sp. nov.">
<emphasis id="BDDDFC7CFF88BB22FE1D80133A0E284C" box="[450,590,1049,1074]" italics="true" pageId="7" pageNumber="438">T. perplexus</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF88BB22FD8580103ADC284C" bold="true" box="[602,668,1049,1074]" pageId="7" pageNumber="438">
<taxonomicNameLabel id="A6EE4107FF88BB22FD8580103ADC284C" box="[602,668,1049,1074]" pageId="7" pageNumber="438" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but similar to
<taxonomicName id="48A95BEDFF88BB22FC8880133C38284C" authority="Irmler, 1985" authorityName="Irmler" authorityYear="1985" box="[855,1144,1049,1074]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="BDDDFC7CFF88BB22FC8880133B9D284F" box="[855,989,1050,1073]" italics="true" pageId="7" pageNumber="438">T. proximus</emphasis>
<bibRefCitation id="EB385D9FFF88BB22FC3680103C38284C" author="Irmler" box="[1001,1144,1049,1074]" pageId="7" pageNumber="438" refString="Irmler, U. (1985) Neue Arten der Gattungen Aneucamptus und Thoracophorus (Col., Staphylinidae) aus der Neotropis. Entomologische Blatter, 81, 41 - 58." type="journal article" year="1985">Irmler, 1985</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="48A95BEDFF88BB22FB6980133D6F284F" box="[1206,1327,1050,1073]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="struyvei" status="sp. nov.">
<emphasis id="BDDDFC7CFF88BB22FB6980133D6F284F" box="[1206,1327,1050,1073]" italics="true" pageId="7" pageNumber="438">T. struyvei</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF88BB22FAE680103D3C284C" bold="true" box="[1337,1404,1049,1074]" pageId="7" pageNumber="438">
<taxonomicNameLabel id="A6EE4107FF88BB22FAE680103D3C284C" box="[1337,1404,1049,1074]" pageId="7" pageNumber="438" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="48A95BEDFF88BB22FA58801339CC2828" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BDDDFC7CFF88BB22FA58801338B62828" italics="true" pageId="7" pageNumber="438">T. aequalis</emphasis>
<bibRefCitation id="EB385D9FFF88BB22FEDE803439CC2828" author="Sharp" box="[257,396,1085,1110]" pageId="7" pageNumber="438" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
also resembles
<taxonomicName id="48A95BEDFF88BB22FD9580373B302828" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[586,880,1085,1110]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="BDDDFC7CFF88BB22FD9580373AE52828" box="[586,677,1085,1110]" italics="true" pageId="7" pageNumber="438">T. exilis</emphasis>
(
<bibRefCitation id="EB385D9FFF88BB22FD6780343B282828" author="Erichson" box="[696,872,1085,1110]" pageId="7" pageNumber="438" refString="Erichson, W. F. (1840) Genera et species Staphylinorum Insectorum Coleopterorum Familiae. Morin, Berlin, 954 pp. http: // dx. doi. org / 10.5962 / bhl. title. 59644" type="book" year="1840">Erichson, 1840</bibRefCitation>
)
</taxonomicName>
in the shape of the head, the antennae and the aedeagus (Figs (8A, B, C), but is distinctly larger.
</paragraph>
</subSubSection>
<subSubSection id="C7B373E5FF88BB22FF18808C3ABB2BA7" pageId="7" pageNumber="438" type="description">
<paragraph id="8F16206EFF88BB22FF18808C3C2828E0" blockId="7.[151,1437,725,2010]" box="[199,1128,1157,1182]" pageId="7" pageNumber="438">
<emphasis id="BDDDFC7CFF88BB22FF18808C391928E0" bold="true" box="[199,345,1157,1182]" pageId="7" pageNumber="438">Description.</emphasis>
Length: 2.0 mm. Colouration: light-brown; legs and antennae yellow.
</paragraph>
<paragraph id="8F16206EFF88BB22FF1880A03AD5292F" blockId="7.[151,1437,725,2010]" pageId="7" pageNumber="438">
Head:
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long,
<quantity id="48518D8BFF88BB22FE6380A33A6028BF" box="[444,544,1194,1218]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="438" unit="mm" value="0.31">0.31 mm</quantity>
wide; eyes prominent; approximately twice as long as temples; temples shortly rounded to neck; posterior angle of temples 114°; sides in front of eyes up to base of antennae approximately half as long as eyes and slightly narrowed; clypeus nearly triangular with short straight central part; deeper margined near base of antennae than at central clypeus; pair of swellings on posterior vertex distinct; without punctation; microsculpture deep and dense; surface matte
</paragraph>
<paragraph id="8F16206EFF88BB22FF1881573CC1299C" blockId="7.[151,1437,725,2010]" pageId="7" pageNumber="438">Antenna as long as head and half of pronotum combined; first and second antennomeres thicker than following four antennomeres; second global; 1.5 times wider than conical third; third antennomere nearly as long as second; antennomeres four to six much shorter, but as wide as third; width of following antennomeres increasing; antennomere four approximately quadrate; antennomere ten nearly twice as wide as long.</paragraph>
<paragraph id="8F16206EFF88BB22FF1881E73D6F2AEB" blockId="7.[151,1437,725,2010]" pageId="7" pageNumber="438">
Pronotum:
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long,
<quantity id="48518D8BFF88BB22FDDF81E43A292A7B" box="[512,617,1517,1542]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="438" unit="mm" value="0.35">0.35 mm</quantity>
wide; widest at middle; slightly narrowed to anterior angles; anterior angles nearly rectangular; anterior edge slightly prominent in convex curve; more strongly narrowed to posterior angles in concave curve; anterior edge approximately 1.5 times as wide as posterior edge; laterally with deep longitudinal impressions; slightly divided into two deeper impressions; central impression deep and with central swelling in posterior half; without punctures; isodiametric microsculpture deep and dense; surface matte.
</paragraph>
<paragraph id="8F16206EFF88BB22FF1882A839F02B7F" blockId="7.[151,1437,725,2010]" pageId="7" pageNumber="438">
Elytra:
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long,
<quantity id="48518D8BFF88BB22FE1582AB3A702AC7" box="[458,560,1698,1722]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="7" pageNumber="438" unit="mm" value="0.41">0.41 mm</quantity>
wide; shoulders and posterior angles shortly rounded; on each side of suture with three distinct longitudinal carinae; without punctures; isodiametric microsculpture deep, but less dense as on pronotum; surface matte.
</paragraph>
<paragraph id="8F16206EFF88BB22FF18830439752B13" blockId="7.[151,1437,725,2010]" pageId="7" pageNumber="438">Abdomen parallel; segments III to V with slight transverse impression at base; with extremely fine punctation; punctures with short white scales; isodiametric microsculpture dense and deep; slightly less deep than on elytra; surface matte.</paragraph>
<paragraph id="8F16206EFF88BB22FF1883703ABB2BA7" blockId="7.[151,1437,725,2010]" pageId="7" pageNumber="438">Aedeagus smoothly rounded; in lateral aspect, inner edge of apical lobe slightly sinuate; apical lobe not exceeding ventral sclerotised plate; paramera not exceeding apex of central lobe; short transparent plate at apex as wide as paramera; numerous sensillae along paramera.</paragraph>
</subSubSection>
</treatment>
</document>