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<document id="8812C4F81F3814FA9F9B9D017CBFAFE3" ID-CLB-Dataset="7950" ID-DOI="10.11646/zootaxa.4878.3.3" ID-GBIF-Dataset="b9c711ba-146d-447b-ac2d-e1a0fcce0053" ID-ISSN="1175-5326" ID-Zenodo-Dep="4425178" ID-ZooBank="B660B3B2-7D4A-44C4-92C9-4BE24A1EA950" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610040207257" checkinUser="plazi" docAuthor="Sirenko, Boris I." docDate="2020" docId="039487CEF006FFB8FF5DFABCFDAAF866" docLanguage="en" docName="zootaxa.4878.3.3.pdf" docOrigin="Zootaxa 4878 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptochiton kantori Sirenko 2016" docType="treatment" docVersion="8" lastPageNumber="472" masterDocId="FFADFFB6F003FFBDFFCAFF86FFD6FF91" masterDocTitle="New shelf and bathyal Philippine leptochitons (Mollusca: Polyplacophora Lepidopleurida)" masterLastPageNumber="500" masterPageNumber="467" pageNumber="472" updateTime="1698923802988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6DA48B2CC25DB0C22AAEB07BBEDEAEEE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D8F4C06AC73DB3A486CD2EA71874E600">
<mods:title id="B03B28831DAAF61B1E051124A56D4952">New shelf and bathyal Philippine leptochitons (Mollusca: Polyplacophora Lepidopleurida)</mods:title>
</mods:titleInfo>
<mods:name id="1ED60A8A80FDF4E41C261BD5CA8ABB38" type="personal">
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<mods:namePart id="72459D07B39566AEDF599094E919447A">Sirenko, Boris I.</mods:namePart>
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<mods:title id="26C74817B0CEF899246ADA208EE0DA8B">Zootaxa</mods:title>
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<mods:part id="84C550AC38E5BADEBFE5250AC10DE089">
<mods:date id="69E463D6CBD546A5E4CEE95092070B3A">2020</mods:date>
<mods:detail id="50BDB4B5FE4EA89A9FCE75A76AE01E87" type="pubDate">
<mods:number id="71647D63B6CD09A791513B29F9F01205">2020-11-16</mods:number>
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<mods:classification id="A41188CB01A5E7D180008F21F53F7BB7">journal article</mods:classification>
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<mods:identifier id="CBB418A1935A03DB6DB9F0C295A11614" type="DOI">10.11646/zootaxa.4878.3.3</mods:identifier>
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<treatment id="039487CEF006FFB8FF5DFABCFDAAF866" ID-DOI="http://doi.org/10.5281/zenodo.4426174" ID-GBIF-Taxon="180324288" ID-Zenodo-Dep="4426174" LSID="urn:lsid:plazi:treatment:039487CEF006FFB8FF5DFABCFDAAF866" httpUri="http://treatment.plazi.org/id/039487CEF006FFB8FF5DFABCFDAAF866" lastPageNumber="472" pageId="5" pageNumber="472">
<subSubSection id="C3276553F006FFB8FF5DFABCFDECFAC4" box="[151,570,1338,1365]" pageId="5" pageNumber="472" type="nomenclature">
<paragraph id="8B8236D8F006FFB8FF5DFABCFDECFAC4" blockId="5.[151,570,1338,1402]" box="[151,570,1338,1365]" pageId="5" pageNumber="472">
<heading id="D0CA81B4F006FFB8FF5DFABCFDECFAC4" bold="true" box="[151,570,1338,1365]" fontSize="11" level="1" pageId="5" pageNumber="472" reason="1">
<taxonomicName id="4C3D4D5BF006FFB8FF5DFABCFDECFAC4" ID-CoL="3TLXT" authority="Sirenko, 2016" authorityName="Sirenko" authorityYear="2016" box="[151,570,1338,1365]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="472" phylum="Mollusca" rank="species" species="kantori">
<emphasis id="B949EACAF006FFB8FF5DFABCFDECFAC4" bold="true" box="[151,570,1338,1365]" pageId="5" pageNumber="472">
<emphasis id="B949EACAF006FFB8FF5DFABCFE5EFAC4" bold="true" box="[151,392,1338,1365]" italics="true" pageId="5" pageNumber="472">Leptochiton kantori</emphasis>
Sirenko, 2016
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3276553F006FFB8FF5DFAD9FEC4FAEB" box="[151,274,1375,1402]" pageId="5" pageNumber="472" type="description">
<paragraph id="8B8236D8F006FFB8FF5DFAD9FEC4FAEB" blockId="5.[151,570,1338,1402]" box="[151,274,1375,1402]" pageId="5" pageNumber="472">
(
<figureCitation id="13062A5DF006FFB8FF55FAD9FF39FAE8" box="[159,239,1375,1401]" captionStart="FIGURE 4" captionStartId="4.[151,250,1933,1957]" captionTargetBox="[228,1360,636,1905]" captionTargetId="figure-196@4.[225,1363,633,1909]" captionTargetPageId="4" captionText="FIGURE 4. Leptochiton kantori, Philippines (MNHN-IM-2007-35505), BL 5.1 mm: A. Valve I, dorsal view; B. Valve III, dorsal view; C. Valve IV, dorsal view; D. Valve VIII, dorsal view; E. Valve III, detail of tegmentum in central area; F. Valve III, rostral view; G. Valve VIII, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4425186" httpUri="https://zenodo.org/record/4425186/files/figure.png" pageId="5" pageNumber="472">Figs. 4</figureCitation>
,
<figureCitation id="13062A5DF006FFB8FF31FAD9FEDFFAEB" box="[251,265,1375,1402]" captionStart="FIGURE 5" captionStartId="5.[151,250,1207,1231]" captionTargetBox="[217,1371,186,1179]" captionTargetId="figure-16@5.[213,1375,181,1183]" captionTargetPageId="5" captionText="FIGURE 5. Leptochiton kantori, Philippines (MNHN-IM-2007-35505), BL 5.1 mm:A, B. Dorsal, marginal and ventral scales; C. Sutural scales; D. Radula." figureDoi="http://doi.org/10.5281/zenodo.4425190" httpUri="https://zenodo.org/record/4425190/files/figure.png" pageId="5" pageNumber="472">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3276553F006FFB8FF5DFA18FD7BFA27" box="[151,685,1438,1462]" pageId="5" pageNumber="472" type="reference_group">
<paragraph id="8B8236D8F006FFB8FF5DFA18FD7BFA27" blockId="5.[151,685,1438,1462]" box="[151,685,1438,1462]" pageId="5" pageNumber="472">
<taxonomicName id="4C3D4D5BF006FFB8FF5DFA18FDC0FA27" ID-CoL="3TLXT" authority="Sirenko 2016: 53" authorityName="Sirenko" authorityPageNumber="53" authorityYear="2016" box="[151,534,1438,1462]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="472" phylum="Mollusca" rank="species" species="kantori">
<emphasis id="B949EACAF006FFB8FF5DFA18FEB6FA27" box="[151,352,1438,1462]" italics="true" pageId="5" pageNumber="472">Leptochiton kantori</emphasis>
Sirenko 2016: 53
</taxonomicName>
, figs 172–204.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F006FFB8FF5DFA62FB62F91E" pageId="5" pageNumber="472" type="materials_examined">
<paragraph id="8B8236D8F006FFB8FF5DFA62FEF9F9B3" blockId="5.[151,1436,1508,2039]" pageId="5" pageNumber="472">
<emphasis id="B949EACAF006FFB8FF5DFA62FE93FA6F" bold="true" box="[151,325,1508,1534]" pageId="5" pageNumber="472">Type material.</emphasis>
<materialsCitation id="3B553C85F006FFB8FE86FA62FEF9F9B3" ID-GBIF-Occurrence="3044017306" collectionCode="MNHN" pageId="5" pageNumber="472" specimenCode="IM-2000-31608, IM-2000-31609, 31610" specimenCount="3" typeStatus="holotype">
<typeStatus id="5486887AF006FFB8FE86FA62FE62FA6F" box="[332,436,1508,1534]" pageId="5" pageNumber="472" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED2CAE1DF006FFB8FE09FA62FDCCFA6F" box="[451,538,1508,1534]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="472">MNHN</collectionCode>
<specimenCode id="DB9B9EA3F006FFB8FDEBFA62FD03FA6E" box="[545,725,1508,1535]" collectionCode="IM-" pageId="5" pageNumber="472">IM-2000-31608</specimenCode>
) and
<specimenCount id="9D3BFD51F006FFB8FCDEFA62FC42FA6E" box="[788,916,1508,1535]" pageId="5" pageNumber="472" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
(
<collectionCode id="ED2CAE1DF006FFB8FC69FA62FC2CFA6F" box="[931,1018,1508,1534]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="472">MNHN</collectionCode>
<specimenCode id="DB9B9EA3F006FFB8FBC8FA62FB61FA6F" box="[1026,1207,1508,1534]" collectionCode="IM-" pageId="5" pageNumber="472">IM-2000-31609</specimenCode>
-
<specimenCode id="DB9B9EA3F006FFB8FB74FA62FAD0FA6F" box="[1214,1286,1508,1534]" collectionCode="IM-" pageId="5" pageNumber="472">31610</specimenCode>
, MNHN-IM- 2007-17439).
</materialsCitation>
</paragraph>
<paragraph id="8B8236D8F006FFB8FF0DF9AAFCE9F9D6" blockId="5.[151,1436,1508,2039]" box="[199,831,1580,1607]" pageId="5" pageNumber="472">
<materialsCitation id="3B553C85F006FFB8FF0DF9AAFCE9F9D6" ID-GBIF-Occurrence="3044017310" box="[199,831,1580,1607]" country="Vanuatu" elevation="664" latitude="-14.833333" location="Vanuatu" longLatPrecision="1286" longitude="167.05" pageId="5" pageNumber="472" specimenCount="1" typeStatus="holotype">
<emphasis id="B949EACAF006FFB8FF0DF9AAFEB3F9D7" bold="true" box="[199,357,1580,1606]" pageId="5" pageNumber="472">
<typeStatus id="5486887AF006FFB8FF0DF9AAFED4F9D7" box="[199,258,1580,1606]" pageId="5" pageNumber="472">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32A7648F006FFB8FEA6F9ABFE1DF9D6" box="[364,459,1581,1607]" name="Vanuatu" pageId="5" pageNumber="472">Vanuatu</collectingCountry>
,
<geoCoordinate id="EE09501FF006FFB8FE1CF9AAFDE2F9D6" box="[470,564,1580,1607]" degrees="14" direction="south" minutes="50" orientation="latitude" pageId="5" pageNumber="472" precision="925" value="-14.833333">14°50’S</geoCoordinate>
,
<geoCoordinate id="EE09501FF006FFB8FD8AF9AAFD78F9D7" box="[576,686,1580,1606]" degrees="167" direction="east" minutes="03" orientation="longitude" pageId="5" pageNumber="472" precision="925" value="167.05">167°03’E</geoCoordinate>
,
<quantity id="4CC59B3DF006FFB8FD73F9AAFCEDF9D6" box="[697,827,1580,1607]" metricMagnitude="2" metricUnit="m" metricValue="6.64" metricValueMax="7.2299999999999995" metricValueMin="6.05" pageId="5" pageNumber="472" unit="m" value="664.0" valueMax="723.0" valueMin="605.0">
<elevation id="0010D1EBF006FFB8FD73F9AAFCEDF9D6" box="[697,827,1580,1607]" metricMagnitude="2" metricUnit="m" metricValue="6.64" metricValueMax="7.2299999999999995" metricValueMin="6.05" pageId="5" pageNumber="472" unit="m" value="664.0" valueMax="723.0" valueMin="605.0">605–723 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8236D8F006FFB8FF0DF9D6FB62F91E" blockId="5.[151,1436,1508,2039]" pageId="5" pageNumber="472">
<emphasis id="B949EACAF006FFB8FF0DF9D6FE66F9FB" bold="true" box="[199,432,1616,1642]" pageId="5" pageNumber="472">Material examined.</emphasis>
<materialsCitation id="3B553C85F006FFB8FE7CF9D6FB79F91E" ID-GBIF-Occurrence="3044017322" collectingDate="2005-05-26" country="Philippines" elevation="530" latitude="8.885" location="Mindanao Sea" longLatPrecision="129" longitude="123.558334" pageId="5" pageNumber="472" specimenCount="2" stateProvince="Bohol" typeStatus="holotype">
<typeStatus id="5486887AF006FFB8FE7CF9D6FDC8F9FB" box="[438,542,1616,1642]" pageId="5" pageNumber="472" type="holotype">Holotype</typeStatus>
and
<typeStatus id="5486887AF006FFB8FD99F9D7FD61F9FA" box="[595,695,1617,1643]" pageId="5" pageNumber="472" type="paratype">paratype</typeStatus>
;
<collectingCountry id="F32A7648F006FFB8FD0BF9D6FC95F9FB" box="[705,835,1616,1642]" name="Philippines" pageId="5" pageNumber="472">Philippines</collectingCountry>
. PANGLAO 2005,
<collectingRegion id="49F9F83AF006FFB8FBD1F9D6FB88F9FB" box="[1051,1118,1616,1642]" country="Philippines" name="Bohol" pageId="5" pageNumber="472">Bohol</collectingRegion>
/
<location id="8EE26003F006FFB8FBADF9D6FADCF9FB" LSID="urn:lsid:plazi:treatment:039487CEF006FFB8FF5DFABCFDAAF866:8EE26003F006FFB8FBADF9D6FADCF9FB" box="[1127,1290,1616,1642]" country="Philippines" latitude="8.885" longLatPrecision="129" longitude="123.558334" name="Mindanao Sea" pageId="5" pageNumber="472" stateProvince="Bohol">Mindanao Sea</location>
, stn CP2361,
<geoCoordinate id="EE09501FF006FFB8FF5DF9F2FED4F91E" box="[151,258,1652,1679]" degrees="8" direction="north" minutes="53.1" orientation="latitude" pageId="5" pageNumber="472" precision="92" value="8.885">8°53.1’N</geoCoordinate>
,
<geoCoordinate id="EE09501FF006FFB8FEC7F9F2FE46F91E" box="[269,400,1652,1679]" degrees="123" direction="east" minutes="33.5" orientation="longitude" pageId="5" pageNumber="472" precision="92" value="123.558334">123°33.5’E</geoCoordinate>
,
<quantity id="4CC59B3DF006FFB8FE56F9F2FDCEF91E" box="[412,536,1652,1679]" metricMagnitude="2" metricUnit="m" metricValue="5.295" metricValueMax="5.43" metricValueMin="5.16" pageId="5" pageNumber="472" unit="m" value="529.5" valueMax="543.0" valueMin="516.0">
<elevation id="0010D1EBF006FFB8FE56F9F2FDCEF91E" box="[412,536,1652,1679]" metricMagnitude="2" metricUnit="m" metricValue="5.295" metricValueMax="5.43" metricValueMin="5.16" pageId="5" pageNumber="472" unit="m" value="529.5" valueMax="543.0" valueMin="516.0">516-543 m</elevation>
</quantity>
, 1 spm (MNHN-IM-2000-35505), BL
<quantity id="4CC59B3DF006FFB8FC00F9F2FBF4F91E" box="[970,1058,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="472" unit="mm" value="5.1">5.1 mm</quantity>
,
<date id="FF831018F006FFB8FBE4F9F2FB79F91E" box="[1070,1199,1652,1679]" pageId="5" pageNumber="472" value="2005-05-26">
<collectingDate id="EFC7E9F0F006FFB8FBE4F9F2FB79F91E" box="[1070,1199,1652,1679]" pageId="5" pageNumber="472" value="2005-05-26">26.05.2005</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F006FFB8FF0DF91EFBD6F947" pageId="5" pageNumber="472" type="distribution">
<paragraph id="8B8236D8F006FFB8FF0DF91EFBD6F947" blockId="5.[151,1436,1508,2039]" pageId="5" pageNumber="472">
<emphasis id="B949EACAF006FFB8FF0DF91EFE8FF923" bold="true" box="[199,345,1688,1714]" pageId="5" pageNumber="472">Distribution</emphasis>
. Off
<collectingCountry id="F32A7648F006FFB8FE56F91FFE2EF922" box="[412,504,1689,1715]" name="Vanuatu" pageId="5" pageNumber="472">Vanuatu</collectingCountry>
(
<quantity id="4CC59B3DF006FFB8FDC0F91EFD47F922" box="[522,657,1688,1715]" metricMagnitude="2" metricUnit="m" metricValue="6.64" metricValueMax="7.2299999999999995" metricValueMin="6.05" pageId="5" pageNumber="472" unit="m" value="664.0" valueMax="723.0" valueMin="605.0">605–723 m</quantity>
), the
<collectingCountry id="F32A7648F006FFB8FD1CF91EFC42F923" box="[726,916,1688,1714]" name="Solomon Islands" pageId="5" pageNumber="472">Solomon Islands</collectingCountry>
(
<quantity id="4CC59B3DF006FFB8FC6DF91EFBFAF922" box="[935,1068,1688,1715]" metricMagnitude="2" metricUnit="m" metricValue="2.325" metricValueMax="2.36" metricValueMin="2.29" pageId="5" pageNumber="472" unit="m" value="232.5" valueMax="236.0" valueMin="229.0">229–236 m</quantity>
) and the
<collectingCountry id="F32A7648F006FFB8FB55F91EFACBF923" box="[1183,1309,1688,1714]" name="Philippines" pageId="5" pageNumber="472">Philippines</collectingCountry>
(516–543) (herein). The new records expand the distribution for this species to the north.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F006FFB8FF0DF966FDAAF866" pageId="5" pageNumber="472" type="discussion">
<paragraph id="8B8236D8F006FFB8FF0DF966FDAAF866" blockId="5.[151,1436,1508,2039]" pageId="5" pageNumber="472">
<emphasis id="B949EACAF006FFB8FF0DF966FEEDF96B" bold="true" box="[199,315,1760,1786]" pageId="5" pageNumber="472">Remarks.</emphasis>
The studied specimen (MNHN-IM-2000-35505) is similar to the
<typeStatus id="5486887AF006FFB8FBD9F967FB95F96A" box="[1043,1091,1761,1787]" pageId="5" pageNumber="472">type</typeStatus>
specimens except of the sculpture of the tegmentum of the head valve and the lateral areas of the intermediate valves which have granules (vs. lacking granules in
<taxonomicName id="4C3D4D5BF006FFB8FEB2F8AFFE3EF8D3" authorityName="Sirenko" authorityYear="2016" box="[376,488,1832,1858]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="472" phylum="Mollusca" rank="species" species="kantori">
<emphasis id="B949EACAF006FFB8FEB2F8AFFE3EF8D3" box="[376,488,1832,1858]" italics="true" pageId="5" pageNumber="472">L. kantori</emphasis>
</taxonomicName>
). The specimen from the
<collectingCountry id="F32A7648F006FFB8FCDAF8AEFC58F8D3" box="[784,910,1832,1858]" name="Philippines" pageId="5" pageNumber="472">Philippines</collectingCountry>
possesses four gills on each side and a radula that is
<quantity id="4CC59B3DF006FFB8FF2AF8CAFEE3F8F6" box="[224,309,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="472" unit="mm" value="1.5">1.5 mm</quantity>
long with 37 transverse rows of mature teeth. Each granule of the tegmentum in the studied specimen has five pores of aesthetes. The studied specimen is superficially similar to
<taxonomicName id="4C3D4D5BF006FFB8FC3FF8F7FB2AF81B" authority="Kaas, 1991" authorityName="Kaas" authorityYear="1991" box="[1013,1276,1904,1931]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="472" phylum="Mollusca" rank="species" species="perscitus">
<emphasis id="B949EACAF006FFB8FC3FF8F7FBA4F81B" box="[1013,1138,1905,1930]" italics="true" pageId="5" pageNumber="472">L perscitus</emphasis>
<bibRefCitation id="EFAC4B29F006FFB8FBB0F8F7FB2AF81B" author="Kaas, P." box="[1146,1276,1904,1931]" pageId="5" pageNumber="472" pagination="9 - 27" refId="ref11132" refString="Kaas, P. (1991) Mollusca, Polyplacophora: Deep-water chitons from New Caledonia. In: Resultats des Campagnes Musorstom, Volume 7. Memoires du Museum national d'Histoire Naturelle, 150, 9 - 27." type="journal article" year="1991">Kaas, 1991</bibRefCitation>
</taxonomicName>
, but differs in possessing four gills (vs. six gills in
<taxonomicName id="4C3D4D5BF006FFB8FDE6F813FD78F83F" authorityName="Kaas" authorityYear="1991" box="[556,686,1941,1966]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="472" phylum="Mollusca" rank="species" species="perscitus">
<emphasis id="B949EACAF006FFB8FDE6F813FD78F83F" box="[556,686,1941,1966]" italics="true" pageId="5" pageNumber="472">L. perscitus</emphasis>
</taxonomicName>
), longitudinal rows of granules on the central areas of intermediate valves (vs. granules arranged quincuncially in
<taxonomicName id="4C3D4D5BF006FFB8FD63F83FFCFAF843" authorityName="Kaas" authorityYear="1991" box="[681,812,1977,2002]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="472" phylum="Mollusca" rank="species" species="perscitus">
<emphasis id="B949EACAF006FFB8FD63F83FFCFAF843" box="[681,812,1977,2002]" italics="true" pageId="5" pageNumber="472">L. perscitus</emphasis>
</taxonomicName>
), and a tridentate head of the major lateral teeth of the radula (vs. unidentate head in
<taxonomicName id="4C3D4D5BF006FFB8FE23F85BFDBDF867" authorityName="Kaas" authorityYear="1991" box="[489,619,2013,2038]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="5" pageNumber="472" phylum="Mollusca" rank="species" species="perscitus">
<emphasis id="B949EACAF006FFB8FE23F85BFDBDF867" box="[489,619,2013,2038]" italics="true" pageId="5" pageNumber="472">L. perscitus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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