313 lines
38 KiB
XML
313 lines
38 KiB
XML
<document id="99A49556F3B8E24F6FCCD05AEDF7E6E0" ID-CLB-Dataset="2516" ID-DOI="10.21248/contrib.entomol.65.2.287-295" ID-GBIF-Dataset="d90577e5-63ce-465c-a7de-f23e166ad4fe" ID-ISSN="0005-805X" ID-Zenodo-Dep="5370974" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630269881874" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2015" docId="039487C55952744BFF69BF33FBB8FA63" docLanguage="en" docName="ContribEntomol.2015.65.2.287-295.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 65 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1884" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Sunius marratus Assing 2015, spec. nov." docType="treatment" docVersion="8" lastPageNumber="292" masterDocId="FFADFFBD5957744EFFD4BB00FFB3FF82" masterDocTitle="A revision of Sunius XV. Six new species from Kyrgyzstan and the Himalaya, and additional records (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="295" masterPageNumber="287" pageNumber="292" updateTime="1699233740109" updateUser="ExternalLinkService">
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<mods:title id="2B35AF4EF7728B7E7CE821EB7356B0FB">A revision of Sunius XV. Six new species from Kyrgyzstan and the Himalaya, and additional records (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:name id="922B2279E128C35DA93F3DAE6DDB98CC" type="personal">
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<mods:namePart id="AEF15F459D87603AEDFF723A6CBE95FF">Assing, Volker</mods:namePart>
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<mods:title id="D206AD147E5EA1BA883E316E690F4DAA">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="64691D64F5542695385CC576FAFA2FFB">
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<mods:date id="888C006481C16D3A161596196C5481E4">2015</mods:date>
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<mods:number id="05B5971A72B1CE1728CEE167DF2CB724">2015-12-21</mods:number>
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<mods:number id="B4515DBC06FCA01398D30D47BF521825">65</mods:number>
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<mods:identifier id="7414803DF7CB38FC448AD240DE756439" type="DOI">10.21248/contrib.entomol.65.2.287-295</mods:identifier>
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<treatment id="039487C55952744BFF69BF33FBB8FA63" ID-DOI="http://doi.org/10.5281/zenodo.5878366" ID-GBIF-Taxon="192428291" ID-Zenodo-Dep="5878366" LSID="urn:lsid:plazi:treatment:039487C55952744BFF69BF33FBB8FA63" httpUri="http://treatment.plazi.org/id/039487C55952744BFF69BF33FBB8FA63" lastPageNumber="292" pageId="5" pageNumber="292">
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<subSubSection id="C32765585952744BFF69BF33FE66FBCC" box="[189,469,1075,1102]" pageId="5" pageNumber="292" type="nomenclature">
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<paragraph id="8B8236D35952744BFF69BF33FE66FBCC" blockId="5.[189,469,1075,1135]" box="[189,469,1075,1102]" pageId="5" pageNumber="292">
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<heading id="D0CA81BF5952744BFF69BF33FE66FBCC" box="[189,469,1075,1102]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="292" reason="3">
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<taxonomicName id="4C3D4D505952744BFF69BF33FED6FBCC" ID-CoL="53GPY" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[189,357,1075,1102]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="marratus" status="sp. nov.">
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<emphasis id="B949EAC15952744BFF69BF33FED6FBCC" box="[189,357,1075,1102]" italics="true" pageId="5" pageNumber="292">Sunius marratus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57BA5952744BFEB8BF33FE66FBCC" box="[364,469,1075,1102]" pageId="5" pageNumber="292" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32765585952744BFF69BF59FEA8FBED" box="[189,283,1113,1135]" pageId="5" pageNumber="292" type="description">
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<paragraph id="8B8236D35952744BFF69BF59FEA8FBED" blockId="5.[189,469,1075,1135]" box="[189,283,1113,1135]" pageId="5" pageNumber="292">
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(
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<figureCitation id="13062A565952744BFF15BF59FEA5FBED" box="[193,278,1113,1135]" captionStart="Figs 1–14" captionStartId="4.[189,229,1969,1991]" captionTargetBox="[195,1426,266,1947]" captionTargetId="figure-136@4.[189,1436,260,1963]" captionTargetPageId="4" captionText="Figs 1–14: Sunius alaianus spec. nov. (1–6), S. marratus spec. nov. (7–10), and S. virilis spec. nov. (11–14): habitus (1); forebody (2, 7, 11); male sternite VIII (3, 8, 12); aedeagus in lateral and in ventral view (4, 9–10, 13–14); ventral process of aedeagus in ventral view (5); internal structures of aedeagus in ventral view (6). Scale bars: 1: 1.0 mm; 2, 7, 11: 0.5 mm; 3, 8, 12: 0.2 mm; 4–6, 9–10, 13–14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5370976" httpUri="https://zenodo.org/record/5370976/files/figure.png" pageId="5" pageNumber="292">Figs 7–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32765585952744BFF43BF89FEEFFA43" pageId="5" pageNumber="292" type="materials_examined">
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<paragraph id="8B8236D35952744BFF43BF89FEEFFA43" blockId="5.[151,757,1161,1473]" pageId="5" pageNumber="292">
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<emphasis id="B949EAC15952744BFF43BF89FEF1FB23" bold="true" box="[151,322,1161,1185]" pageId="5" pageNumber="292">Type material</emphasis>
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||
:
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||
<materialsCitation id="3B553C8E5952744BFEB5BF89FE5FFA83" ID-GBIF-Occurrence="3446988356" collectingDate="1978-10-14" collectionCode="MHNG" collectorName="W. Bengal & Besuchet- Lobl" country="India" determinerName="V. Assing" elevation="2000" location="Ghoom - Lopchu" municipality="Darjeeling distr." pageId="5" pageNumber="292" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
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||
<typeStatus id="548688715952744BFEB5BF89FE75FB23" box="[353,454,1161,1185]" pageId="5" pageNumber="292" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D3BFD5A5952744BFE0BBF8AFE47FB23" box="[479,500,1162,1185]" pageId="5" pageNumber="292" type="male">♂</specimenCount>
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: “
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<collectingCountry id="F32A76435952744BFDCFBF89FDDBFB23" box="[539,616,1161,1185]" name="India" pageId="5" pageNumber="292">INDIA</collectingCountry>
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<collectorName id="26C853055952744BFD54BF8AFD5CFB23" box="[640,751,1161,1185]" pageId="5" pageNumber="292">W. Bengal</collectorName>
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,
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<collectingMunicipality id="6BE6ACA95952744BFF43BFA9FEE2FB43" box="[151,337,1193,1217]" country="India" municipality="Ghoom" name="Darjeeling distr." pageId="5" pageNumber="292">Darjeeling distr.</collectingMunicipality>
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,
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<location id="8EE260085952744BFEB6BFA9FDA5FB43" LSID="urn:lsid:plazi:treatment:039487C55952744BFF69BF33FBB8FA63:8EE260085952744BFEB6BFA9FDA5FB43" box="[354,534,1193,1217]" country="India" municipality="Darjeeling distr." name="Ghoom - Lopchu" pageId="5" pageNumber="292">Ghoom–Lopchu</location>
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</quantity>
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,
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<collectorName id="26C853055952744BFD5CBFA9FF7BFB63" pageId="5" pageNumber="292">BesuchetLöbl</collectorName>
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<date id="FF8310135952744BFF04BFC9FE91FB63" box="[208,290,1225,1249]" pageId="5" pageNumber="292" value="1978-10-14">
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<collectingDate id="EFC7E9FB5952744BFF04BFC9FE91FB63" box="[208,290,1225,1249]" pageId="5" pageNumber="292" value="1978-10-14">14.X.78</collectingDate>
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</date>
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/
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<typeStatus id="548688715952744BFEEFBFC9FE1FFB63" box="[315,428,1225,1249]" pageId="5" pageNumber="292" type="holotype">Holotypus</typeStatus>
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<specimenCount id="9D3BFD5A5952744BFE62BFCAFE78FB63" box="[438,459,1226,1249]" pageId="5" pageNumber="292" type="male">♂</specimenCount>
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||
<taxonomicName id="4C3D4D505952744BFE05BFC9FD31FB63" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[465,642,1225,1249]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="marratus" status="sp. nov.">
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<emphasis id="B949EAC15952744BFE05BFC9FD31FB63" box="[465,642,1225,1249]" italics="true" pageId="5" pageNumber="292">
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<location id="8EE260085952744BFE05BFC9FDAAFB62" LSID="urn:lsid:plazi:treatment:039487C55952744BFF69BF33FBB8FA63:8EE260085952744BFE05BFC9FDAAFB62" box="[465,537,1225,1248]" country="India" municipality="Darjeeling distr." name="Sunius" pageId="5" pageNumber="292">Sunius</location>
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marratus
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||
</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A27A57BA5952744BFD5FBFCAFD47FB63" box="[651,756,1226,1249]" pageId="5" pageNumber="292" rank="species">spec. nov.</taxonomicNameLabel>
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det.
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||
<determinerName id="7E5EDA175952744BFF1EBFEAFE9CFA83" box="[202,303,1257,1281]" pageId="5" pageNumber="292">V. Assing</determinerName>
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2015” (
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)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B553C8E5952744BFE2DBFEAFDA7FAA3" ID-GBIF-Occurrence="3446988323" collectionCode="MHNG" collectorName="W. Bengal" country="India" determinerName="V. Assing" elevation="2000" municipality="Ghoom" pageId="5" pageNumber="292" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
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||
<typeStatus id="548688715952744BFE2DBFEAFDD6FA80" box="[505,613,1258,1282]" pageId="5" pageNumber="292" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="9D3BFD5A5952744BFDA6BFE9FD1EFA83" box="[626,685,1257,1281]" pageId="5" pageNumber="292" type="female">3 ♀♀</specimenCount>
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||
: same data as holotype (
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||
<collectionCode id="ED2CAE165952744BFEB8BE09FE77FAA3" box="[364,452,1289,1313]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="292" type="Museum">MHNG</collectionCode>
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, cAss)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B553C8E5952744BFDF2BE09FD07FAC3" ID-GBIF-Occurrence="3446988364" collectingDate="1978-10-12" collectionCode="MHNG" collectorName="W. Bengal" country="India" determinerName="V. Assing" elevation="2000" municipality="Ghoom" pageId="5" pageNumber="292" specimenCount="2" specimenCount-female="1" typeStatus="paratype">
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||
<specimenCount id="9D3BFD5A5952744BFDF2BE09FDF8FAA3" box="[550,587,1289,1313]" pageId="5" pageNumber="292" type="female">1 ♀</specimenCount>
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,
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<specimenCount id="9D3BFD5A5952744BFD8BBE09FD21FAA3" box="[607,658,1289,1313]" pageId="5" pageNumber="292" type="generic">1 ex.</specimenCount>
|
||
without abdomen: same data, but “
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||
<date id="FF8310135952744BFE09BE29FD83FAC3" box="[477,560,1321,1345]" pageId="5" pageNumber="292" value="1978-10-12">
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<collectingDate id="EFC7E9FB5952744BFE09BE29FD83FAC3" box="[477,560,1321,1345]" pageId="5" pageNumber="292" value="1978-10-12">12.X.78</collectingDate>
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||
</date>
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||
” (
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||
<collectionCode id="ED2CAE165952744BFD87BE29FD1EFAC3" box="[595,685,1321,1345]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="292" type="Museum">MHNG</collectionCode>
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||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B553C8E5952744BFD1CBE29FD92FA03" ID-GBIF-Occurrence="3446988309" collectingDate="1978-10-15" collectionCode="MHNG" collectorName="Besuchet-Lobl" country="India" county="Besuchet-Lobl" elevation="1500" location="Darjeeling distr." municipality="Ghoom" pageId="5" pageNumber="292" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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||
<specimenCount id="9D3BFD5A5952744BFD1CBE29FD5DFAC3" box="[712,750,1321,1345]" pageId="5" pageNumber="292" type="female">1 ♀</specimenCount>
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||
: “
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||
<collectingCountry id="F32A76435952744BFF75BE49FF5DFAE3" box="[161,238,1353,1377]" name="India" pageId="5" pageNumber="292">INDIA</collectingCountry>
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||
W. Bengal,
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||
<location id="8EE260085952744BFEAFBE49FD80FAE3" LSID="urn:lsid:plazi:treatment:039487C55952744BFF69BF33FBB8FA63:8EE260085952744BFEAFBE49FD80FAE3" box="[379,563,1353,1377]" country="India" county="Besuchet-Lobl" municipality="Ghoom" name="Darjeeling distr." pageId="5" pageNumber="292">Darjeeling distr.</location>
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,
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||
<location id="8EE260085952744BFD90BE49FD24FAE3" LSID="urn:lsid:plazi:treatment:039487C55952744BFF69BF33FBB8FA63:8EE260085952744BFD90BE49FD24FAE3" box="[580,663,1353,1377]" country="India" county="Besuchet-Lobl" municipality="Ghoom" name="Ghoom" pageId="5" pageNumber="292">Ghoom</location>
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<quantity id="4CC59B365952744BFD70BE49FD47FAE3" box="[676,756,1353,1377]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="292" unit="m" value="1500.0">
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<elevation id="0010D1E05952744BFD70BE49FD47FAE3" box="[676,756,1353,1377]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="292" unit="m" value="1500.0">1500 m</elevation>
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||
</quantity>
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||
<date id="FF8310135952744BFF43BE69FF58FA03" box="[151,235,1385,1409]" pageId="5" pageNumber="292" value="1978-10-15">
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<collectingDate id="EFC7E9FB5952744BFF43BE69FF58FA03" box="[151,235,1385,1409]" pageId="5" pageNumber="292" value="1978-10-15">15.X.78</collectingDate>
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</date>
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,
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||
<collectorName id="26C853055952744BFED4BE69FE2DFA03" box="[256,414,1385,1409]" pageId="5" pageNumber="292">Besuchet-Löbl</collectorName>
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||
” (
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||
<collectionCode id="ED2CAE165952744BFE6BBE69FDAAFA03" box="[447,537,1385,1409]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="292" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B553C8E5952744BFDE1BE69FEEBFA43" ID-GBIF-Occurrence="3446988327" collectingDate="1978-10-16" collectorName="W. Bengal" country="India" county="Besuchet-Lobl" elevation="3100" location="Darjeeling distr." municipality="Tonglu" pageId="5" pageNumber="292" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D3BFD5A5952744BFDE1BE69FDE8FA03" box="[565,603,1385,1409]" pageId="5" pageNumber="292" type="male">1 ♂</specimenCount>
|
||
: “
|
||
<collectingCountry id="F32A76435952744BFDAEBE69FD74FA03" box="[634,711,1385,1409]" name="India" pageId="5" pageNumber="292">INDIA</collectingCountry>
|
||
W. Bengal,
|
||
<location id="8EE260085952744BFF3BBE89FE12FA23" LSID="urn:lsid:plazi:treatment:039487C55952744BFF69BF33FBB8FA63:8EE260085952744BFF3BBE89FE12FA23" box="[239,417,1417,1441]" country="India" county="Besuchet-Lobl" municipality="Tonglu" name="Darjeeling distr." pageId="5" pageNumber="292">Darjeeling distr.</location>
|
||
,
|
||
<collectingMunicipality id="6BE6ACA95952744BFE78BE89FE4BFA23" box="[428,504,1417,1441]" pageId="5" pageNumber="292">Tonglu</collectingMunicipality>
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||
<quantity id="4CC59B365952744BFE2BBE89FDFCFA23" box="[511,591,1417,1441]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="292" unit="m" value="3100.0">
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<elevation id="0010D1E05952744BFE2BBE89FDFCFA23" box="[511,591,1417,1441]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="292" unit="m" value="3100.0">3100 m</elevation>
|
||
</quantity>
|
||
<date id="FF8310135952744BFD82BE89FD18FA23" box="[598,683,1417,1441]" pageId="5" pageNumber="292" value="1978-10-16">
|
||
<collectingDate id="EFC7E9FB5952744BFD82BE89FD18FA23" box="[598,683,1417,1441]" pageId="5" pageNumber="292" value="1978-10-16">16.X.78</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingCounty id="62E34E5F5952744BFD62BE8AFEB3FA43" pageId="5" pageNumber="292">Besuchet-Löbl</collectingCounty>
|
||
” (cAss)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32765585952744BFF43BEE9FDFDF9C3" pageId="5" pageNumber="292" type="etymology">
|
||
<paragraph id="8B8236D35952744BFF43BEE9FDFDF9C3" blockId="5.[150,756,1513,1601]" pageId="5" pageNumber="292">
|
||
<emphasis id="B949EAC15952744BFF43BEE9FEBDF983" bold="true" box="[151,270,1513,1537]" pageId="5" pageNumber="292">Etymology</emphasis>
|
||
: The specific epithet is an adjective derived from the Latin noun marra (hoe) and alludes to the shape of the internal structures of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32765585952744BFF43BD69FAAFFAC3" pageId="5" pageNumber="292" type="description">
|
||
<paragraph id="8B8236D35952744BFF43BD69FD04F943" blockId="5.[150,756,1641,2049]" pageId="5" pageNumber="292">
|
||
<emphasis id="B949EAC15952744BFF43BD69FEAAF903" bold="true" box="[151,281,1641,1665]" pageId="5" pageNumber="292">Description</emphasis>
|
||
: Body length 3.0–4.0 mm; length of forebody
|
||
<quantity id="4CC59B365952744BFF42BD89FEABF923" box="[150,280,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="5" pageNumber="292" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.6–1.7 mm</quantity>
|
||
. Coloration: body black; legs pale-brown with slightly darker femora; antennae pale-reddish.
|
||
</paragraph>
|
||
<paragraph id="8B8236D35952744BFF43BDC9FECFF823" blockId="5.[150,756,1641,2049]" pageId="5" pageNumber="292">
|
||
Head (
|
||
<figureCitation id="13062A565952744BFF38BDC9FE9AF963" box="[236,297,1737,1761]" captionStart="Figs 1–14" captionStartId="4.[189,229,1969,1991]" captionTargetBox="[195,1426,266,1947]" captionTargetId="figure-136@4.[189,1436,260,1963]" captionTargetPageId="4" captionText="Figs 1–14: Sunius alaianus spec. nov. (1–6), S. marratus spec. nov. (7–10), and S. virilis spec. nov. (11–14): habitus (1); forebody (2, 7, 11); male sternite VIII (3, 8, 12); aedeagus in lateral and in ventral view (4, 9–10, 13–14); ventral process of aedeagus in ventral view (5); internal structures of aedeagus in ventral view (6). Scale bars: 1: 1.0 mm; 2, 7, 11: 0.5 mm; 3, 8, 12: 0.2 mm; 4–6, 9–10, 13–14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5370976" httpUri="https://zenodo.org/record/5370976/files/figure.png" pageId="5" pageNumber="292">Fig. 7</figureCitation>
|
||
) as long as broad or weakly transverse, tapering posteriad in posterior half; punctation conspicuously coarse and dense in anterior dorsal portion, very sparse and less coarse in posterior dorsal portion; interstices without microsculpture. Eyes enormous and bulging, more than twice as long as postocular region in dorsal view.
|
||
</paragraph>
|
||
<paragraph id="8B8236D35952744BFF43BCAAFE15F783" blockId="5.[150,756,1641,2049]" pageId="5" pageNumber="292">
|
||
Pronotum (
|
||
<figureCitation id="13062A565952744BFEC1BCA9FEE0F843" box="[277,339,1961,1985]" captionStart="Figs 1–14" captionStartId="4.[189,229,1969,1991]" captionTargetBox="[195,1426,266,1947]" captionTargetId="figure-136@4.[189,1436,260,1963]" captionTargetPageId="4" captionText="Figs 1–14: Sunius alaianus spec. nov. (1–6), S. marratus spec. nov. (7–10), and S. virilis spec. nov. (11–14): habitus (1); forebody (2, 7, 11); male sternite VIII (3, 8, 12); aedeagus in lateral and in ventral view (4, 9–10, 13–14); ventral process of aedeagus in ventral view (5); internal structures of aedeagus in ventral view (6). Scale bars: 1: 1.0 mm; 2, 7, 11: 0.5 mm; 3, 8, 12: 0.2 mm; 4–6, 9–10, 13–14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5370976" httpUri="https://zenodo.org/record/5370976/files/figure.png" pageId="5" pageNumber="292">Fig. 7</figureCitation>
|
||
) weakly oblong and slightly narrower than head; punctation coarse and dense; midline moderately broadly impunctate.
|
||
</paragraph>
|
||
<paragraph id="8B8236D35952744BFCCEBBE9FBC1FEC3" blockId="5.[793,1401,233,705]" pageId="5" pageNumber="292">
|
||
Elytra (
|
||
<figureCitation id="13062A565952744BFCBDBBE9FC15FE83" box="[873,934,233,257]" captionStart="Figs 1–14" captionStartId="4.[189,229,1969,1991]" captionTargetBox="[195,1426,266,1947]" captionTargetId="figure-136@4.[189,1436,260,1963]" captionTargetPageId="4" captionText="Figs 1–14: Sunius alaianus spec. nov. (1–6), S. marratus spec. nov. (7–10), and S. virilis spec. nov. (11–14): habitus (1); forebody (2, 7, 11); male sternite VIII (3, 8, 12); aedeagus in lateral and in ventral view (4, 9–10, 13–14); ventral process of aedeagus in ventral view (5); internal structures of aedeagus in ventral view (6). Scale bars: 1: 1.0 mm; 2, 7, 11: 0.5 mm; 3, 8, 12: 0.2 mm; 4–6, 9–10, 13–14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5370976" httpUri="https://zenodo.org/record/5370976/files/figure.png" pageId="5" pageNumber="292">Fig. 7</figureCitation>
|
||
) short, approximately 0.65 times as long as pronotum; punctation dense, irregular, and ill-defined. Hind wings completely reduced.
|
||
</paragraph>
|
||
<paragraph id="8B8236D35952744BFCCEBA49FC0EFE63" blockId="5.[793,1401,233,705]" pageId="5" pageNumber="292">Abdomen slightly broader than elytra; punctation fine, moderately dense anteriorly, gradually becoming less dense towards posterior tergites; interstices with shallow microsculpture; posterior margin of tergite VII without palisade fringe.</paragraph>
|
||
<paragraph id="8B8236D35952744BFCCEBAEAFAC4FD43" blockId="5.[793,1401,233,705]" pageId="5" pageNumber="292">
|
||
<specimenCount id="9D3BFD5A5952744BFCCEBAEAFC9CFD83" box="[794,815,490,513]" pageId="5" pageNumber="292" type="male">♂</specimenCount>
|
||
: sternite VII without distinct modifications; sternite VIII (
|
||
<figureCitation id="13062A565952744BFC67B909FC43FDA3" box="[947,1008,521,545]" captionStart="Figs 1–14" captionStartId="4.[189,229,1969,1991]" captionTargetBox="[195,1426,266,1947]" captionTargetId="figure-136@4.[189,1436,260,1963]" captionTargetPageId="4" captionText="Figs 1–14: Sunius alaianus spec. nov. (1–6), S. marratus spec. nov. (7–10), and S. virilis spec. nov. (11–14): habitus (1); forebody (2, 7, 11); male sternite VIII (3, 8, 12); aedeagus in lateral and in ventral view (4, 9–10, 13–14); ventral process of aedeagus in ventral view (5); internal structures of aedeagus in ventral view (6). Scale bars: 1: 1.0 mm; 2, 7, 11: 0.5 mm; 3, 8, 12: 0.2 mm; 4–6, 9–10, 13–14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5370976" httpUri="https://zenodo.org/record/5370976/files/figure.png" pageId="5" pageNumber="292">Fig. 8</figureCitation>
|
||
) approximately as broad as long, pubescence unmodified, posterior excision of subtriangular shape, its depth approximately one-tenth the length of sternite; aedeagus (
|
||
<figureCitation id="13062A565952744BFBDEB969FBC7FD03" box="[1034,1140,617,641]" captionStart="Figs 1–14" captionStartId="4.[189,229,1969,1991]" captionTargetBox="[195,1426,266,1947]" captionTargetId="figure-136@4.[189,1436,260,1963]" captionTargetPageId="4" captionText="Figs 1–14: Sunius alaianus spec. nov. (1–6), S. marratus spec. nov. (7–10), and S. virilis spec. nov. (11–14): habitus (1); forebody (2, 7, 11); male sternite VIII (3, 8, 12); aedeagus in lateral and in ventral view (4, 9–10, 13–14); ventral process of aedeagus in ventral view (5); internal structures of aedeagus in ventral view (6). Scale bars: 1: 1.0 mm; 2, 7, 11: 0.5 mm; 3, 8, 12: 0.2 mm; 4–6, 9–10, 13–14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5370976" httpUri="https://zenodo.org/record/5370976/files/figure.png" pageId="5" pageNumber="292">Figs 9–10</figureCitation>
|
||
)
|
||
<quantity id="4CC59B365952744BFB51B969FB57FD03" box="[1157,1252,617,641]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="292" unit="mm" value="0.45">0.45 mm</quantity>
|
||
long; ventral process stout and of distinctive shape; internal sac with a pair of long and similarly shaped sclerotized structures.
|
||
</paragraph>
|
||
<paragraph id="8B8236D35952744BFCCEB9E9FAAFFAC3" blockId="5.[793,1400,745,1346]" pageId="5" pageNumber="292">
|
||
<emphasis id="B949EAC15952744BFCCEB9E9FC59FC83" bold="true" box="[794,1002,745,769]" pageId="5" pageNumber="292">Comparative notes</emphasis>
|
||
: Based on the external (coarse punctation, reduced elytra and hind wings, enormous eyes) and the male sexual characters (aedeagus with a pair of conspicuously long sclerotized structures in internal sac),
|
||
<taxonomicName id="4C3D4D505952744BFC81B869FC7DFC03" authorityName="Assing" authorityYear="2015" box="[853,974,873,897]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="marratus">
|
||
<emphasis id="B949EAC15952744BFC81B869FC7DFC03" box="[853,974,873,897]" italics="true" pageId="5" pageNumber="292">S. marratus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C3D4D505952744BFB57B869FA95FC03" authorityName="ASSING" authorityYear="2010" box="[1155,1318,873,897]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="manasluensis">
|
||
<emphasis id="B949EAC15952744BFB57B869FA95FC03" box="[1155,1318,873,897]" italics="true" pageId="5" pageNumber="292">S. manasluensis</emphasis>
|
||
</taxonomicName>
|
||
group, which is distributed in the Himalaya and
|
||
<collectingCountry id="F32A76435952744BFB2FB889FA8CFC23" box="[1275,1343,905,929]" name="China" pageId="5" pageNumber="292">China</collectingCountry>
|
||
, and which previously included eight species:
|
||
<taxonomicName id="4C3D4D505952744BFB36B8A9FC7DFC63" authority="ASSING, 2010" authorityName="ASSING" authorityYear="2010" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="manasluensis">
|
||
<emphasis id="B949EAC15952744BFB36B8A9FC80FC63" italics="true" pageId="5" pageNumber="292">S. manasluensis</emphasis>
|
||
<bibRefCitation id="EFAC4B225952744BFCE9B8C9FC7DFC63" author="ASSING, V." box="[829,974,969,993]" pageId="5" pageNumber="292" pagination="1045 - 1061" refId="ref6612" refString="ASSING, V. 2010: A revision of Palaearctic Sunius. XII. New species, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). - Linzer Biologische Beitrage 42 (2): 1045 - 1061." type="journal article" year="2010">ASSING, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F32A76435952744BFC0BB8C9FB90FC63" box="[991,1059,969,993]" name="Nepal" pageId="5" pageNumber="292">Nepal</collectingCountry>
|
||
: Manaslu),
|
||
<taxonomicName id="4C3D4D505952744BFB70B8C9FAA0FC63" baseAuthorityName="COIFFAIT" baseAuthorityYear="1987" box="[1188,1299,969,993]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="galiberti">
|
||
<emphasis id="B949EAC15952744BFB70B8C9FAA0FC63" box="[1188,1299,969,993]" italics="true" pageId="5" pageNumber="292">S. galiberti</emphasis>
|
||
</taxonomicName>
|
||
(Central
|
||
<collectingCountry id="F32A76435952744BFCCEB8E9FCEFFB83" box="[794,860,1001,1025]" name="Nepal" pageId="5" pageNumber="292">Nepal</collectingCountry>
|
||
: Phulchoki),
|
||
<taxonomicName id="4C3D4D505952744BFC21B8E9FAAAFB83" authority="ASSING, 2010" authorityName="ASSING" authorityYear="2010" box="[1013,1305,1001,1025]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="turgescens">
|
||
<emphasis id="B949EAC15952744BFC21B8E9FBC4FB83" box="[1013,1143,1001,1025]" italics="true" pageId="5" pageNumber="292">S. turgescens</emphasis>
|
||
<bibRefCitation id="EFAC4B225952744BFB50B8E9FAAAFB83" author="ASSING, V." box="[1156,1305,1001,1025]" pageId="5" pageNumber="292" pagination="1045 - 1061" refId="ref6612" refString="ASSING, V. 2010: A revision of Palaearctic Sunius. XII. New species, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). - Linzer Biologische Beitrage 42 (2): 1045 - 1061." type="journal article" year="2010">ASSING, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F32A76435952744BFAF9B8E9FAC0FB83" box="[1325,1395,1001,1025]" name="China" pageId="5" pageNumber="292">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F9F8315952744BFCCEBF09FCC1FBA3" box="[794,882,1033,1057]" country="China" name="Yunnan" pageId="5" pageNumber="292">Yunnan</collectingRegion>
|
||
),
|
||
<taxonomicName id="4C3D4D505952744BFC58BF09FB10FBA3" authority="ASSING, 2010" authorityName="ASSING" authorityYear="2010" box="[908,1187,1033,1057]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="macrops">
|
||
<emphasis id="B949EAC15952744BFC58BF09FC4FFBA3" box="[908,1020,1033,1057]" italics="true" pageId="5" pageNumber="292">S. macrops</emphasis>
|
||
<bibRefCitation id="EFAC4B225952744BFBD8BF09FB10FBA3" author="ASSING, V." box="[1036,1187,1033,1057]" pageId="5" pageNumber="292" pagination="1045 - 1061" refId="ref6612" refString="ASSING, V. 2010: A revision of Palaearctic Sunius. XII. New species, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). - Linzer Biologische Beitrage 42 (2): 1045 - 1061." type="journal article" year="2010">ASSING, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F32A76435952744BFB6FBF09FAB2FBA3" box="[1211,1281,1033,1057]" name="China" pageId="5" pageNumber="292">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F9F8315952744BFAC0BF09FADFFBA3" box="[1300,1388,1033,1057]" country="China" name="Yunnan" pageId="5" pageNumber="292">Yunnan</collectingRegion>
|
||
),
|
||
<taxonomicName id="4C3D4D505952744BFCCEBF29FBECFBC3" authority="ASSING, 2011" authorityName="ASSING" authorityYear="2011" box="[794,1119,1065,1089]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="cursor">
|
||
<emphasis id="B949EAC15952744BFCCEBF29FCC7FBC3" box="[794,884,1065,1089]" italics="true" pageId="5" pageNumber="292">S. cursor</emphasis>
|
||
ASSING, 2011 (
|
||
<collectingCountry id="F32A76435952744BFBC6BF29FBE4FBC3" box="[1042,1111,1065,1089]" name="Nepal" pageId="5" pageNumber="292">Nepal</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D505952744BFBB2BF29FAC5FBC3" authority="ASSING, 2011" authorityName="ASSING" authorityYear="2011" box="[1126,1398,1065,1089]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="baculatus">
|
||
<emphasis id="B949EAC15952744BFBB2BF29FB50FBC3" box="[1126,1251,1065,1089]" italics="true" pageId="5" pageNumber="292">S. baculatus</emphasis>
|
||
ASSING, 2011
|
||
</taxonomicName>
|
||
(East
|
||
<collectingCountry id="F32A76435952744BFC8EBF49FC2EFBE3" box="[858,925,1097,1121]" name="Nepal" pageId="5" pageNumber="292">Nepal</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C3D4D505952744BFC61BF49FB0DFBE3" authority="(Darjeeling)" baseAuthorityName="Darjeeling" box="[949,1214,1097,1121]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="cameroni">
|
||
<emphasis id="B949EAC15952744BFC61BF49FB9CFBE3" box="[949,1071,1097,1121]" italics="true" pageId="5" pageNumber="292">S. cameroni</emphasis>
|
||
(Darjeeling)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3D4D505952744BFAD5BF49FB88FB03" authority="(COIFFAIT, 1978) (Bhutan)" baseAuthorityName="COIFFAIT" baseAuthorityYear="1978" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="bouddha">
|
||
<emphasis id="B949EAC15952744BFAD5BF49FAC4FBE3" box="[1281,1399,1097,1121]" italics="true" pageId="5" pageNumber="292">S. bouddha</emphasis>
|
||
(COIFFAIT, 1978) (
|
||
<collectingCountry id="F32A76435952744BFC34BF69FB80FB03" box="[992,1075,1129,1153]" name="Bhutan" pageId="5" pageNumber="292">Bhutan</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
. The aedeagus of
|
||
<taxonomicName id="4C3D4D505952744BFB2BBF69FACBFB03" authorityName="Assing" authorityYear="2015" box="[1279,1400,1129,1153]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="marratus">
|
||
<emphasis id="B949EAC15952744BFB2BBF69FACBFB03" box="[1279,1400,1129,1153]" italics="true" pageId="5" pageNumber="292">S. marratus</emphasis>
|
||
</taxonomicName>
|
||
is most similar to that of
|
||
<taxonomicName id="4C3D4D505952744BFBE2BF89FB18FB23" baseAuthorityName="COIFFAIT" baseAuthorityYear="1978" box="[1078,1195,1161,1185]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="bouddha">
|
||
<emphasis id="B949EAC15952744BFBE2BF89FB18FB23" box="[1078,1195,1161,1185]" italics="true" pageId="5" pageNumber="292">S. bouddha</emphasis>
|
||
</taxonomicName>
|
||
, but distinguished by the slightly different shape of the ventral process and particularly by the distinctly shorter and differently shaped internal structures. For illustrations of
|
||
<taxonomicName id="4C3D4D505952744BFAD4BFE9FAC4FA83" baseAuthorityName="COIFFAIT" baseAuthorityYear="1978" box="[1280,1399,1257,1281]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="bouddha">
|
||
<emphasis id="B949EAC15952744BFAD4BFE9FAC4FA83" box="[1280,1399,1257,1281]" italics="true" pageId="5" pageNumber="292">S. bouddha</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="EFAC4B225952744BFC92BE09FC44FAA3" author="ASSING, V." box="[838,1015,1289,1313]" pageId="5" pageNumber="292" pagination="151 - 193" refId="ref6693" refString="ASSING, V. 2011 b: The Sunius species of the Palaearctic region (Coleoptera: Staphylinidae: Paederinae). - Linzer Biologische Beitrage 43 (1): 151 - 193." type="journal article" year="2011">ASSING (2011b)</bibRefCitation>
|
||
, for those of other species of the
|
||
<taxonomicName id="4C3D4D505952744BFCCEBE29FC0DFAC3" authorityName="ASSING" authorityYear="2010" box="[794,958,1321,1345]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="manasluensis">
|
||
<emphasis id="B949EAC15952744BFCCEBE29FC0DFAC3" box="[794,958,1321,1345]" italics="true" pageId="5" pageNumber="292">S. manasluensis</emphasis>
|
||
</taxonomicName>
|
||
group see
|
||
<bibRefCitation id="EFAC4B225952744BFBE4BE29FB70FAC3" author="ASSING, V." box="[1072,1219,1321,1345]" pageId="5" pageNumber="292" pagination="1045 - 1061" refId="ref6612" refString="ASSING, V. 2010: A revision of Palaearctic Sunius. XII. New species, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). - Linzer Biologische Beitrage 42 (2): 1045 - 1061." type="journal article" year="2010">ASSING (2010</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAC4B225952744BFB19BE29FAA2FAC3" author="ASSING, V." box="[1229,1297,1321,1345]" pageId="5" pageNumber="292" pagination="89 - 148" refId="ref6656" refString="ASSING, V. 2011 a: A revision of the genus Neosclerus CAMERON (Coleoptera: Staphylinidae: Paederinae). - Beitrage zur Entomologie, Keltern 61 (1): 89 - 148." type="journal article" year="2011">2011a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32765585952744BFCCEBE69FBB8FA63" pageId="5" pageNumber="292" type="distribution">
|
||
<paragraph id="8B8236D35952744BFCCEBE69FBB8FA63" blockId="5.[794,1399,1385,1505]" pageId="5" pageNumber="292">
|
||
<emphasis id="B949EAC15952744BFCCEBE69FB3AFA03" bold="true" box="[794,1161,1385,1409]" pageId="5" pageNumber="292">Distribution and natural history</emphasis>
|
||
: The
|
||
<typeStatus id="548688715952744BFB1BBE6AFB4EFA00" box="[1231,1277,1386,1410]" pageId="5" pageNumber="292">type</typeStatus>
|
||
specimens were collected in three geographically close localities in Darjeeling district,
|
||
<collectingRegion id="49F9F8315952744BFC26BEAAFBC9FA43" box="[1010,1146,1449,1473]" country="India" name="West Bengal" pageId="5" pageNumber="292">West Bengal</collectingRegion>
|
||
(North
|
||
<collectingCountry id="F32A76435952744BFB0EBEA9FAA5FA43" box="[1242,1302,1449,1473]" name="India" pageId="5" pageNumber="292">India</collectingCountry>
|
||
), at altitudes of
|
||
<quantity id="4CC59B365952744BFCA0BEC9FBB4FA63" box="[884,1031,1481,1505]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="3.1" metricValueMin="1.5" pageId="5" pageNumber="292" unit="m" value="2300.0" valueMax="3100.0" valueMin="1500.0">1500–3100 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |