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<mods:title id="98D3246ED746F3E388BF0B2613350712">Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species</mods:title>
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<mods:namePart id="AE68606C8A86E9C8C8F72AE75FA8D446">Tan, Ming Kai</mods:namePart>
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<mods:affiliation id="8C26819C5661BD21DBE9F6561A0E74D9">Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE 1410, Brunei Darussalam.</mods:affiliation>
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<mods:affiliation id="46910EB56FE4B9AE41B6601F3AB0FE51">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia.</mods:affiliation>
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<mods:namePart id="119DCAEE4C62DAD7F4143D4B154B3DC0">Chung, Authur Y. C.</mods:namePart>
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<mods:affiliation id="5897D0CBD794DA6D81AE89E71F23F329">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia.</mods:affiliation>
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<treatment id="039487A4FFD2FFC2FF1FF7A6FCA7F811" ID-DOI="http://doi.org/10.5281/zenodo.5170045" ID-GBIF-Taxon="184010544" ID-Zenodo-Dep="5170045" LSID="urn:lsid:zoobank.org:act:A3AE87F7-8196-4C9B-BDEF-8E9D21B92BE5" httpUri="http://treatment.plazi.org/id/039487A4FFD2FFC2FF1FF7A6FCA7F811" lastPageNumber="63" pageId="62" pageNumber="63">
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<taxonomicName id="4C3D4D31FFD2FFC2FF1FF7A6FD4FFB24" ID-CoL="8P8MK" authority="(Rehn, 1909)" baseAuthorityName="Rehn" baseAuthorityYear="1909" box="[255,666,1219,1245]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B949EAA0FFD2FFC2FF1FF7A6FE2DFB24" bold="true" box="[255,504,1219,1245]" italics="true" pageId="62" pageNumber="63">Nisitrus sumatrensis</emphasis>
(
<bibRefCitation id="EFAC4B43FFD2FFC2FDE7F7A6FD44FB24" author="Rehn" box="[519,657,1219,1245]" firstAuthor="Rehn" pageId="62" pageNumber="63" pagination="177 - 211" refId="ref44962" refString="Rehn J. A. G. 1909. A contribution to the knowledge of the Orthoptera of Sumatra. Bulletin of the American Museum of Natural History 26: 177 - 211." type="journal article" year="1909">Rehn, 1909</bibRefCitation>
)
</taxonomicName>
junior synonym of
<taxonomicName id="4C3D4D31FFD2FFC2FC61F7A6FB6AFB24" authority="Saussure, 1878" authorityName="Saussure" authorityYear="1878" box="[897,1215,1219,1245]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B949EAA0FFD2FFC2FC61F7A6FBD1FB24" bold="true" box="[897,1028,1219,1245]" italics="true" pageId="62" pageNumber="63">N. insignis</emphasis>
<bibRefCitation id="EFAC4B43FFD2FFC2FBEBF7A6FB6AFB24" author="Saussure" box="[1035,1215,1219,1245]" firstAuthor="Saussure" pageId="62" pageNumber="63" pagination="369 - 704" refId="ref45577" refString="Saussure H. 1878. Melanges orthopterologiques. VI. Fascicule Gryllides. Memoires de la Societe de Physique et d'Histoire naturelle de Geneve 25 (2): 369 - 704 [505 - 834]." type="journal article" year="1878">Saussure, 1878</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A27A57DBFFD2FFC2FB2BF7A1FAE0FB27" box="[1227,1333,1220,1246]" pageId="62" pageNumber="63">syn. nov.</taxonomicNameLabel>
<tableCitation id="C6BF0309FFD2FFC2FD0EF78DFC90FAFB" box="[750,837,1256,1282]" captionStart="Table 2" captionStartId="10.[189,258,265,291]" captionTargetPageId="10" captionText="Table 2. Status of original species epithets associated with Nisitrus Saussure, 1878." httpUri="http://table.plazi.org/id/DF42663AFFE6FFF6FF5DF26CFB5AFEDA" pageId="62" pageNumber="63" tableUuid="DF42663AFFE6FFF6FF5DF26CFB5AFEDA">Table 2</tableCitation>
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<treatmentCitation id="0A9C10A3FFD2FFC2FF5DF649FD33FABF" author="Rehn J. A. G." box="[189,742,1324,1350]" page="209" pageId="62" pageNumber="63" year="1909">
<taxonomicName id="4C3D4D31FFD2FFC2FF5DF649FD33FABF" ID-CoL="8P8MK" authority="Rehn, 1909: 209" authorityName="Rehn" authorityPageNumber="209" authorityYear="1909" box="[189,742,1324,1350]" class="Insecta" family="Gryllidae" genus="Crystallomorpha" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B949EAA0FFD2FFC2FF5DF649FDCDFABC" box="[189,536,1324,1350]" italics="true" pageId="62" pageNumber="63">Crystallomorpha sumatrensis</emphasis>
<bibRefCitation id="EFAC4B43FFD2FFC2FDFFF649FD33FABF" author="Rehn" box="[543,742,1324,1350]" firstAuthor="Rehn" page="209" pageId="62" pageNumber="63" pagination="177 - 211" refId="ref44962" refString="Rehn J. A. G. 1909. A contribution to the knowledge of the Orthoptera of Sumatra. Bulletin of the American Museum of Natural History 26: 177 - 211." type="journal article" year="1909">Rehn, 1909: 209</bibRefCitation>
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.
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</paragraph>
<paragraph id="8B8236B2FFD2FFC2FF5DF615FDB7FA73" blockId="62.[189,1296,1391,1455]" box="[189,610,1391,1418]" pageId="62" pageNumber="63">
<taxonomicName id="4C3D4D31FFD2FFC2FF5DF615FEE0FA70" ID-CoL="73ZB5" baseAuthorityName="Haan" baseAuthorityYear="1944" box="[189,309,1392,1417]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="vittatus">
<emphasis id="B949EAA0FFD2FFC2FF5DF615FEE0FA70" box="[189,309,1392,1417]" italics="true" pageId="62" pageNumber="63">N. vittatus</emphasis>
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<bibRefCitation id="EFAC4B43FFD2FFC2FEB3F60AFE2FFA70" author="Chopard" box="[339,506,1391,1417]" firstAuthor="Chopard" pageId="62" pageNumber="63" pagination="213 - 500" refId="ref43495" refString="Chopard L. 1968. Fam. Gryllidae: Subfam. Mogoplistinae, Myrmecophilinae, Scleropterinae, Cachoplistinae, Pteroplistinae, Pentacentrinae, Phalangopsinae, Trigonidiinae, Eneopterinae; Fam. Oecanthidae, Gryllotalpidae. In: Beier M. (ed.) Orthopterorum Catalogus 12: 213 - 500. Junk, ' s-Gravenhage [The Hague]." type="journal article" year="1968">Chopard 1968</bibRefCitation>
12: 352.
</paragraph>
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<treatmentCitationGroup id="AB2D119CFFD2FFC2FF5DF6F0FAC5FA56" box="[189,1296,1428,1455]" pageId="62" pageNumber="63">
<taxonomicName id="4C3D4D31FFD2FFC2FF5DF6F0FE79FA57" ID-CoL="8P8MK" baseAuthorityName="Rehn" baseAuthorityYear="1909" box="[189,428,1429,1454]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B949EAA0FFD2FFC2FF5DF6F0FE79FA57" box="[189,428,1429,1454]" italics="true" pageId="62" pageNumber="63">Nisitrus sumatrensis</emphasis>
</taxonomicName>
<treatmentCitation id="0A9C10A3FFD2FFC2FE2AF6F1FDBAFA57" author="Otte D." box="[458,623,1428,1454]" page="67" pageId="62" pageNumber="63" year="1994">
<bibRefCitation id="EFAC4B43FFD2FFC2FE2AF6F1FDBAFA57" author="Otte" box="[458,623,1428,1454]" firstAuthor="Otte" page="67" pageId="62" pageNumber="63" refId="ref44815" refString="Otte D. 1994. Orthoptera Species File. Vol. 1. Crickets (Grylloidea). Orthopterists' Society and the ANSP, Philadelphia." type="book" year="1994">Otte 1994: 67</bibRefCitation>
</treatmentCitation>
. —
<bibRefCitation id="EFAC4B43FFD2FFC2FD7EF6F1FC5DFA57" author="Cigliano" box="[670,904,1428,1454]" etAl="et al." firstAuthor="Cigliano" pageId="62" pageNumber="63" refId="ref43370" refString="Cigliano M. M., Braun H., Eades D. C. &amp; Otte D. 2020. Orthoptera Species File Online. Version 5 (5.0). Available from http: // orthoptera. speciesfile. org / HomePage / Orthoptera / HomePage. aspx [accessed 8 Mar. 2020]." type="url" year="2020">
Cigliano
<emphasis id="B949EAA0FFD2FFC2FCEBF6F0FC90FA57" box="[779,837,1428,1454]" italics="true" pageId="62" pageNumber="63">et al.</emphasis>
2020
</bibRefCitation>
(
<taxonomicName id="4C3D4D31FFD2FFC2FC76F6F1FBC2FA57" ID-CoL="8NKFC" authorityName="Latreille" authorityYear="1793" box="[918,1047,1428,1454]" class="Insecta" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
Species File online).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFD2FFC2FF5DF6B2FE7DF942" pageId="62" pageNumber="63" type="materials_examined">
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<heading id="D0CA81DEFFD2FFC2FF5DF6B2FE7FFA08" bold="true" box="[189,426,1495,1521]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="B949EAA0FFD2FFC2FF5DF6B2FE7FFA08" bold="true" box="[189,426,1495,1521]" pageId="62" pageNumber="63">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFD2FFC2FF0AF56DFE89F9DB" blockId="62.[189,819,1544,1608]" box="[234,348,1544,1570]" pageId="62" pageNumber="63">
<emphasis id="B949EAA0FFD2FFC2FF0AF56DFE89F9DB" bold="true" box="[234,348,1544,1570]" pageId="62" pageNumber="63">
<typeStatus id="54868810FFD2FFC2FF0AF56DFE89F9DB" box="[234,348,1544,1570]" pageId="62" pageNumber="63" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8236B2FFD2FFC2FF5DF548FCE6F9BE" blockId="62.[189,819,1544,1608]" box="[189,819,1581,1608]" pageId="62" pageNumber="63">
<materialsCitation id="3B553CEFFFD2FFC2FF5DF548FCE6F9BE" ID-GBIF-Occurrence="3333037555" box="[189,819,1581,1608]" collectionCode="AMNH" collectorName="R. Weber" country="Indonesia" location="Sumatra" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="Sumatra" typeStatus="holotype">
<collectingCountry id="F32A7622FFD2FFC2FF5DF548FE8EF9BE" box="[189,347,1581,1607]" name="Indonesia" pageId="62" pageNumber="63">INDONESIA</collectingCountry>
<emphasis id="B949EAA0FFD2FFC2FE98F548FE30F9BE" bold="true" box="[376,485,1581,1607]" pageId="62" pageNumber="63">
<collectingRegion id="49F9F850FFD2FFC2FE98F548FE30F9BE" box="[376,485,1581,1607]" country="Indonesia" name="Sumatera Utara" pageId="62" pageNumber="63">Sumatra</collectingRegion>
</emphasis>
<specimenCount id="9D3BFD3BFFD2FFC2FE1EF54BFDC2F9B1" box="[510,535,1582,1608]" pageId="62" pageNumber="63" type="male"></specimenCount>
;
<collectorName id="26C85364FFD2FFC2FDC4F54BFD47F9BE" box="[548,658,1581,1608]" pageId="62" pageNumber="63">R. Weber</collectorName>
leg.;
<collectionCode id="ED2CAE77FFD2FFC2FD32F548FCE5F9BE" box="[722,816,1581,1607]" collectionName="USA, New York, New York, American Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="62" pageNumber="63">AMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8236B2FFD2FFC2FF5DF514FE88F973" blockId="62.[189,349,1648,1675]" box="[189,349,1648,1675]" pageId="62" pageNumber="63">
<heading id="D0CA81DEFFD2FFC2FF5DF514FE88F973" bold="true" box="[189,349,1648,1675]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="B949EAA0FFD2FFC2FF5DF514FE88F973" bold="true" box="[189,349,1648,1675]" pageId="62" pageNumber="63">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFD2FFC2FF5DF5C4FE7DF942" blockId="62.[189,424,1697,1723]" box="[189,424,1697,1723]" pageId="62" pageNumber="63">
<collectingCountry id="F32A7622FFD2FFC2FF5DF5C4FEE1F942" box="[189,308,1697,1723]" name="Indonesia" pageId="62" pageNumber="63">Indonesia</collectingCountry>
:
<collectingRegion id="49F9F850FFD2FFC2FEDFF5C4FE70F942" box="[319,421,1697,1723]" country="Indonesia" name="Sumatera Utara" pageId="62" pageNumber="63">Sumatra</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFD2FFC2FF5DF581FCA7F811" pageId="62" pageNumber="63" type="discussion">
<paragraph id="8B8236B2FFD2FFC2FF5DF581FEFAF907" blockId="62.[189,303,1764,1790]" box="[189,303,1764,1790]" pageId="62" pageNumber="63">
<heading id="D0CA81DEFFD2FFC2FF5DF581FEFAF907" bold="true" box="[189,303,1764,1790]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="B949EAA0FFD2FFC2FF5DF581FEFAF907" bold="true" box="[189,303,1764,1790]" pageId="62" pageNumber="63">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFD2FFC2FF5DF470FCA7F811" blockId="62.[187,1399,1813,2024]" pageId="62" pageNumber="63">
Based on the study of the
<typeStatus id="54868810FFD2FFC2FDE5F470FDECF8D6" box="[517,569,1813,1839]" pageId="62" pageNumber="63">type</typeStatus>
from
<collectingRegion id="49F9F850FFD2FFC2FD67F470FD3BF8D6" box="[647,750,1813,1839]" country="Indonesia" name="Sumatera Utara" pageId="62" pageNumber="63">Sumatra</collectingRegion>
, we consider
<taxonomicName id="4C3D4D31FFD2FFC2FC7AF470FB9AF8D7" baseAuthorityName="Rehn" baseAuthorityYear="1909" box="[922,1103,1813,1838]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B949EAA0FFD2FFC2FC7AF470FB9AF8D7" box="[922,1103,1813,1838]" italics="true" pageId="62" pageNumber="63">N. sumatrensis</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C3D4D31FFD2FFC2FF5DF45EFEE8F8AD" authorityName="Saussure" authorityYear="1878" box="[189,317,1851,1876]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B949EAA0FFD2FFC2FF5DF45EFEE8F8AD" box="[189,317,1851,1876]" italics="true" pageId="62" pageNumber="63">N. insignis</emphasis>
</taxonomicName>
, even though the colour pattern of the
<typeStatus id="54868810FFD2FFC2FCF5F45FFC86F8AD" box="[789,851,1850,1876]" pageId="62" pageNumber="63">types</typeStatus>
of both species is slightly different. After the examination of the material from
<collectingRegion id="49F9F850FFD2FFC2FDB0F43AFD67F880" box="[592,690,1887,1913]" country="Indonesia" name="Sumatera Utara" pageId="62" pageNumber="63">Sumatra</collectingRegion>
identified “
<taxonomicName id="4C3D4D31FFD2FFC2FCA0F405FC3BF880" baseAuthorityName="Rehn" baseAuthorityYear="1909" box="[832,1006,1888,1913]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B949EAA0FFD2FFC2FCA0F405FC3BF880" box="[832,1006,1888,1913]" italics="true" pageId="62" pageNumber="63">N. sumatrensis</emphasis>
</taxonomicName>
” and “
<taxonomicName id="4C3D4D31FFD2FFC2FBA5F405FB11F880" authorityName="Saussure" authorityYear="1878" box="[1093,1220,1888,1913]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B949EAA0FFD2FFC2FBA5F405FB11F880" box="[1093,1220,1888,1913]" italics="true" pageId="62" pageNumber="63">N. insignis</emphasis>
</taxonomicName>
”, we observed that the colour patterns form a continuum between a lighter version in
<taxonomicName id="4C3D4D31FFD2FFC2FC1EF4E0FBA9F867" authorityName="Saussure" authorityYear="1878" box="[1022,1148,1925,1950]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B949EAA0FFD2FFC2FC1EF4E0FBA9F867" box="[1022,1148,1925,1950]" italics="true" pageId="62" pageNumber="63">N. insignis</emphasis>
</taxonomicName>
and the darker, more contrasting specimens previously identified as in the
<typeStatus id="54868810FFD2FFC2FCF5F4CFFC92F83D" box="[789,839,1962,1988]" pageId="62" pageNumber="63">type</typeStatus>
of
<taxonomicName id="4C3D4D31FFD2FFC2FC8AF4CFFBC6F83A" baseAuthorityName="Rehn" baseAuthorityYear="1909" box="[874,1043,1962,1987]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B949EAA0FFD2FFC2FC8AF4CFFBC6F83A" box="[874,1043,1962,1987]" italics="true" pageId="62" pageNumber="63">N. sumatrensis</emphasis>
</taxonomicName>
. There are also little diagnostic differences in the male and female genitalia for both series.
</paragraph>
</subSubSection>
</treatment>
</document>