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<document id="40E0377C685090E3F35131B44881E559" ID-DOI="10.35463/j.apr.2021.01.06" ID-ISSN="1842-371x" ID-Zenodo-Dep="10520821" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1705451288939" checkinUser="felipe" docAuthor="SCHLAGINTWEIT, FELIX, RASHIDI, KOOROSH &amp; MOSAVINIA, ABDOLMAJID" docDate="2021" docId="0394879D143CA31AFCAA3E1FF1EBF9B4" docLanguage="en" docName="ActaPalaeontolRom.17.1.51-64.pdf" docOrigin="Acta Palaeontologica Romaniae 17 (1)" docSource="http://dx.doi.org/10.35463/j.apr.2021.01.06" docStyle="DocumentStyle:CA474AAD92E97C533D63E0C5C0F159F1.1:ActaPalaeontolRom.2019-.journal_article" docStyleId="CA474AAD92E97C533D63E0C5C0F159F1" docStyleName="ActaPalaeontolRom.2019-.journal_article" docStyleVersion="1" docTitle="Cymopolia mayaense Johnson &amp; Kaska 1965" docType="treatment" docVersion="1" lastPageNumber="56" masterDocId="FFADFFE51439A31DFF9A385BF078FFD3" masterDocTitle="First Record Of Calcareous Green Algae (Dasycladales, Halimedaceae) From The Paleocene Chehel Kaman Formation Of North-Eastern Iran (Kopet-Dagh Basin)" masterLastPageNumber="64" masterPageNumber="51" pageNumber="56" updateTime="1708694136784" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="ABD8786518DD3675DFCB110AC9D725D2">First Record Of Calcareous Green Algae (Dasycladales, Halimedaceae) From The Paleocene Chehel Kaman Formation Of North-Eastern Iran (Kopet-Dagh Basin)</mods:title>
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<mods:namePart id="09A244661C258AD937638101AA5B2F9E">SCHLAGINTWEIT, FELIX</mods:namePart>
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<mods:namePart id="6A18F693BCBAFD5CDA390CADB26EF994">RASHIDI, KOOROSH</mods:namePart>
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<treatment id="0394879D143CA31AFCAA3E1FF1EBF9B4" LSID="urn:lsid:plazi:treatment:0394879D143CA31AFCAA3E1FF1EBF9B4" httpUri="http://treatment.plazi.org/id/0394879D143CA31AFCAA3E1FF1EBF9B4" lastPageId="7" lastPageNumber="56" pageId="5" pageNumber="56">
<subSubSection id="C3276500143CA318FCAA3E1FF562F98F" box="[816,1306,1604,1628]" pageId="5" pageNumber="56" type="nomenclature">
<paragraph id="8B82368B143CA318FCAA3E1FF562F98F" blockId="5.[816,1341,1571,1661]" box="[816,1306,1604,1628]" pageId="5" pageNumber="56">
<taxonomicName id="4C3D4D08143CA318FCAA3E1FF562F98F" authority="Johnson &amp; Kaska, 1965" authorityName="Johnson &amp; Kaska" authorityYear="1965" box="[816,1306,1604,1628]" class="Ulvophyceae" family="Dasycladaceae" genus="Cymopolia" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="56" phylum="Chlorophyta" rank="species" species="mayaense">
<emphasis id="B949EA99143CA318FCAA3E1FF46DF98F" box="[816,1045,1604,1628]" italics="true" pageId="5" pageNumber="56">Cymopolia mayaense</emphasis>
<bibRefCitation id="EFAC4B7A143CA318FB863E1FF562F98F" author="Johnson &amp; Kaska" box="[1052,1306,1604,1628]" firstAuthor="Johnson" pageId="5" pageNumber="56" pagination="1 - 152" refId="ref6929" refString="Johnson, J. H., Kaska, H. V., 1965. Fossil Algae from Guatemala. Professional Contributions of the Colorado School of Mines, 1: 1 - 152." type="journal article" year="1965">Johnson &amp; Kaska, 1965</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C3276500143CA318FCAA3E3EF47FF9AE" box="[816,1031,1637,1661]" pageId="5" pageNumber="56" type="description">
<paragraph id="8B82368B143CA318FCAA3E3EF47FF9AE" blockId="5.[816,1341,1571,1661]" box="[816,1031,1637,1661]" pageId="5" pageNumber="56">
<figureCitation id="13062A0E143CA318FCAA3E3EF300F9AE" box="[816,888,1637,1661]" captionStart="Fig" captionStartId="3.[206,241,1021,1042]" captionTargetBox="[226,1362,157,982]" captionTargetId="figure-787@3.[226,1362,157,982]" captionTargetPageId="3" captionText="Fig. 3 Algal-bearing bioclastic packstone-wackestone from the Paleocene Chehel Kaman Formation of northeastern Iran with Halimeda? sp. (H in a), Jodotella veslensis Morellet &amp; Morellet (J in a-b), Cymopolia cf. mayaense Johnson &amp; Kaska (c-d), Acicularia sp. 1 (A in a), bryozoans (br in a), and common Rahaghia khorassanica (Rahaghi) (c). Locality see Figure 2. Thin-sections Nf 12-2 (a), Nf 14 (b), Nf 6-2 (c-d)." figureDoi="http://doi.org/10.5281/zenodo.10520827" httpUri="https://zenodo.org/record/10520827/files/figure.png" pageId="5" pageNumber="56">Figs. 3</figureCitation>
c-d, 8a-e, g-h
</paragraph>
</subSubSection>
<subSubSection id="C3276500143CA318FCAA3EFCF3C8F8B6" pageId="5" pageNumber="56" type="reference_group">
<paragraph id="8B82368B143CA318FCAA3EFCF439F933" blockId="5.[816,1437,1703,1893]" pageId="5" pageNumber="56">
*1965
<taxonomicName id="4C3D4D08143CA318FCE63EFCF41AF96C" authority="SCHLAGINTWEIT &amp; RASHIDI &amp; MOSAVINIA, 2021" authorityName="SCHLAGINTWEIT &amp; RASHIDI &amp; MOSAVINIA" authorityYear="2021" box="[892,1122,1703,1727]" class="Ulvophyceae" family="Dasycladaceae" genus="Cymopolia" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="56" phylum="Chlorophyta" rank="species" species="mayaense" status="sp. nov.">
<emphasis id="B949EA99143CA318FCE63EFCF41AF96C" box="[892,1122,1703,1727]" italics="true" pageId="5" pageNumber="56">Cymopolia mayaense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E2143CA318FBF13EF3F4DEF96C" box="[1131,1190,1704,1727]" pageId="5" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
Johnson and Kaska, p. 83-85, pl. 18, figs. 1-4.
</paragraph>
<paragraph id="8B82368B143CA318FCAA3EB1F3C8F8B6" blockId="5.[816,1437,1703,1893]" pageId="5" pageNumber="56">
2013
<taxonomicName id="4C3D4D08143CA318FCF43EB1F555F8D1" authority="Johnson and Kaska" authorityName="Johnson and Kaska" box="[878,1325,1770,1794]" class="Ulvophyceae" family="Dasycladaceae" genus="Cymopolia" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="56" phylum="Chlorophyta" rank="species" species="mayaense">
<emphasis id="B949EA99143CA318FCF43EB1F42CF8D1" box="[878,1108,1770,1794]" italics="true" pageId="5" pageNumber="56">Cymopolia mayaense</emphasis>
Johnson and Kaska
</taxonomicName>
Granier in Granier et al., p. 283,
<figureCitation id="13062A0E143CA318FBD83F50F4D7F8F0" box="[1090,1199,1803,1827]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[206,241,1021,1042]" captionStartId-1="5.[224,259,1415,1436]" captionTargetBox-0="[226,1362,157,982]" captionTargetBox-1="[224,1363,152,1376]" captionTargetId-0="figure-787@3.[226,1362,157,982]" captionTargetId-1="figure-389@5.[224,1363,152,1376]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Fig. 3 Algal-bearing bioclastic packstone-wackestone from the Paleocene Chehel Kaman Formation of northeastern Iran with Halimeda? sp. (H in a), Jodotella veslensis Morellet &amp; Morellet (J in a-b), Cymopolia cf. mayaense Johnson &amp; Kaska (c-d), Acicularia sp. 1 (A in a), bryozoans (br in a), and common Rahaghia khorassanica (Rahaghi) (c). Locality see Figure 2. Thin-sections Nf 12-2 (a), Nf 14 (b), Nf 6-2 (c-d)." captionText-1="Fig. 4 Acicularia sp. 1 (a-i, l-o), diverse oriented sections. Note the regular palisade-like arrangement of elongated cysts in h and the presence of a central hollow in some specimens (e.g., d, m). Acicularia sp. 2 (j-k) displaying spherical cysts. Thin-sections: Nf 12-1 (a, k), Nf 12-2 (b, d-e, g-j), Nf 12-3 (c, l), Nf 12 (f, m-o)." figureDoi-0="http://doi.org/10.5281/zenodo.10520827" figureDoi-1="http://doi.org/10.5281/zenodo.10520829" httpUri-0="https://zenodo.org/record/10520827/files/figure.png" httpUri-1="https://zenodo.org/record/10520829/files/figure.png" pageId="5" pageNumber="56">Fig. 3.1-4</figureCitation>
, pl. 4, figs. d-f, pl. 5, figs. a-i, pl. 6, figs. a-c, pl. 7, figs a-f, pl. 8, fig. e (with synonymy).
</paragraph>
</subSubSection>
<caption id="DF426603143CA318FF0D3F28F21AF832" ID-DOI="http://doi.org/10.5281/zenodo.10520831" ID-Zenodo-Dep="10520831" httpUri="https://zenodo.org/record/10520831/files/figure.png" pageId="5" pageNumber="56" startId="5.[151,186,1907,1928]" targetBox="[151,771,1531,1881]" targetPageId="5" targetType="figure">
<paragraph id="8B82368B143CA318FF0D3F28F21AF832" blockId="5.[151,772,1906,2017]" pageId="5" pageNumber="56">
<emphasis id="B949EA99143CA318FF0D3F28F0AAF85B" bold="true" box="[151,210,1907,1928]" pageId="5" pageNumber="56">Fig. 5</emphasis>
<emphasis id="B949EA99143CA318FF403F29F1A5F85B" box="[218,477,1906,1928]" italics="true" pageId="5" pageNumber="56">
<taxonomicName id="4C3D4D08143CA318FF403F29F127F85B" authorityName="Gumbel" authorityYear="1871" box="[218,351,1906,1928]" class="Ulvophyceae" family="Dasycladaceae" genus="Thyrsoporella" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="56" phylum="Chlorophyta" rank="genus">Thyrsoporella</taxonomicName>
/
<taxonomicName id="4C3D4D08143CA318FEF13F29F1A5F85B" authorityName="Raineri" authorityYear="1922" box="[363,477,1906,1928]" class="Ulvophyceae" family="Dasycladaceae" genus="Trinocladus" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="56" phylum="Chlorophyta" rank="genus">Trinocladus</taxonomicName>
</emphasis>
, tangential section (
<emphasis id="B949EA99143CA318FD3F3F28F2C8F85B" bold="true" box="[677,688,1907,1928]" pageId="5" pageNumber="56">c</emphasis>
),
<taxonomicName id="4C3D4D08143CA318FD5F3F28F10FF875" authority="(Elliott)" baseAuthorityName="Elliott" class="Ulvophyceae" family="Dasycladaceae" genus="Uteria" isUncertain="true" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="56" phylum="Chlorophyta" rank="species" species="merienda">
<emphasis id="B949EA99143CA318FD5F3F28F37BF85B" box="[709,771,1907,1928]" italics="true" pageId="5" pageNumber="56">Uteria</emphasis>
aff.
<emphasis id="B949EA99143CA318FF5E3FCBF167F875" box="[196,287,1936,1958]" italics="true" pageId="5" pageNumber="56">merienda</emphasis>
(Elliott)
</taxonomicName>
(
<emphasis id="B949EA99143CA318FE103FCBF1EFF876" bold="true" box="[394,407,1936,1957]" pageId="5" pageNumber="56">b</emphasis>
), oblique longitudinal section from Paleocene carbonates of the Chehel Kaman Formation, northeastern Iran. Thin-sections: Nf 13 (
<emphasis id="B949EA99143CA318FE723F97F18CF832" bold="true" box="[488,500,1996,2017]" pageId="5" pageNumber="56">a</emphasis>
), NF 4 (
<emphasis id="B949EA99143CA318FDDD3F97F22CF832" bold="true" box="[583,596,1996,2017]" pageId="5" pageNumber="56">b</emphasis>
).
</paragraph>
</caption>
<subSubSection id="C3276500143CA31AFCAA3FCBF1EBF9B4" lastPageId="7" lastPageNumber="58" pageId="5" pageNumber="56" type="discussion">
<paragraph id="8B82368B143CA31AFCAA3FCBF1EBF9B4" blockId="5.[816,1437,1935,2058]" lastBlockId="7.[151,772,1317,1639]" lastPageId="7" lastPageNumber="58" pageId="5" pageNumber="56">
<emphasis id="B949EA99143CA318FCAA3FCBF3D9F87B" bold="true" box="[816,929,1936,1960]" pageId="5" pageNumber="56">Remarks:</emphasis>
<taxonomicName id="4C3D4D08143CA318FC373FCBF441F874" box="[941,1081,1936,1959]" class="Ulvophyceae" family="Dasycladaceae" genus="Cymopolia" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="56" phylum="Chlorophyta" rank="species" species="mayaense">
<emphasis id="B949EA99143CA318FC373FCBF441F874" box="[941,1081,1936,1959]" italics="true" pageId="5" pageNumber="56">C. mayaense</emphasis>
</taxonomicName>
represents a species showing a rather wide variability of article morphology and range of biometric data. The type-material of the Johnson collection has been recently revised by Granier in
<bibRefCitation id="EFAC4B7A143CA31AFA923FA9F09AFAEE" author="Granier" etAl="et al." firstAuthor="Granier" lastPageId="7" lastPageNumber="58" pageId="5" pageNumber="56" pagination="2013" refId="ref6782" refString="Granier, B., Radoicic, R., Drobne, K., 2013. Revision of the Jesse Harlan Johnson Collection. Part 1. Some fossil Dasycladales from Guatemala. Carnets de Geologie [Notebooks on Geology]: Article 2013 / 07 (CG 2013 _ A 07)." type="journal article" year="2013">Granier et al. (2013)</bibRefCitation>
. Characteristics of the Iranian specimens are a smooth inner surface, comparably small ovoid sporangia surrounded by 4 to 5 secondary laterals (
<figureCitation id="13062A0E143EA31AFDF43D3CF2C9FAAC" box="[622,689,1383,1407]" captionStart="Fig" captionStartId="8.[217,252,1250,1271]" captionTargetBox="[217,1388,154,1208]" captionTargetId="figure-53@8.[217,1388,154,1208]" captionTargetPageId="8" captionText="Fig. 8 Cymopolia cf. mayaense Johnson &amp; Kaska (a-e, g-h) and Cymopolia sp. (f). a, Oblique transverse section. b, tangential section. c, tangential-oblique section. d, tangential section showing five secondaries (rectangle). e, longitudinal (above)-oblique (below) section. f, oblique transverse section. g, variously oriented sections together with Rahaghia khorassanica (Rahaghi) (left ablove). h, fragmentary tangential section showing five secondaries. Thin-sections: Nf 9 (a), Nf 6-2 (b, d, g), Nf 9 (c), Nf 8 (e), Nf 6-1 (f), Nf 4 (h)." figureDoi="http://doi.org/10.5281/zenodo.10520837" httpUri="https://zenodo.org/record/10520837/files/figure.png" pageId="7" pageNumber="58">Fig. 8</figureCitation>
c-e, h). Most specimens are represented by cylindrical segments (length up to 4.0 mm; diameter up to
<quantity id="4CC59B6E143EA31AFDA63DF2F2EEFA12" box="[572,662,1449,1473]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="58" unit="mm" value="0.9">0.9 mm</quantity>
). The primary laterals bearing numerous fertile ampullae at the distal end are shorther than the secondaries. In some samples,
<taxonomicName id="4C3D4D08143EA31AFF673E56F1D5F9F7" box="[253,429,1549,1573]" class="Ulvophyceae" family="Dasycladaceae" genus="Cymopolia" isUncertain="true" kingdom="Plantae" order="Dasycladales" pageId="7" pageNumber="58" phylum="Chlorophyta" rank="species" species="mayaense">
<emphasis id="B949EA99143EA31AFF673E56F177F9F7" box="[253,271,1549,1572]" italics="true" pageId="7" pageNumber="58">C</emphasis>
. cf.
<emphasis id="B949EA99143EA31AFEDE3E56F1D5F9F7" box="[324,429,1549,1572]" italics="true" pageId="7" pageNumber="58">mayaense</emphasis>
</taxonomicName>
shows an assemblage with the calcareous benthic foraminifera
<taxonomicName id="4C3D4D08143EA31AFD9B3E75F179F9B4" authority="(Rahaghi)" baseAuthorityName="Rahaghi" baseAuthorityYear="1976" class="Globothalamea" family="Rotaliidae" genus="Rahaghia" kingdom="Chromista" order="Rotaliida" pageId="7" pageNumber="58" phylum="Foraminifera" rank="species" species="khorassanica">
<emphasis id="B949EA99143EA31AFD9B3E75F37BF995" box="[513,771,1582,1606]" italics="true" pageId="7" pageNumber="58">Rahaghia khorassanica</emphasis>
(Rahaghi)
</taxonomicName>
(
<figureCitation id="13062A0E143EA31AFE8A3E14F126F9B4" box="[272,350,1615,1639]" captionStart="Fig" captionStartId="3.[206,241,1021,1042]" captionTargetBox="[226,1362,157,982]" captionTargetId="figure-787@3.[226,1362,157,982]" captionTargetPageId="3" captionText="Fig. 3 Algal-bearing bioclastic packstone-wackestone from the Paleocene Chehel Kaman Formation of northeastern Iran with Halimeda? sp. (H in a), Jodotella veslensis Morellet &amp; Morellet (J in a-b), Cymopolia cf. mayaense Johnson &amp; Kaska (c-d), Acicularia sp. 1 (A in a), bryozoans (br in a), and common Rahaghia khorassanica (Rahaghi) (c). Locality see Figure 2. Thin-sections Nf 12-2 (a), Nf 14 (b), Nf 6-2 (c-d)." figureDoi="http://doi.org/10.5281/zenodo.10520827" httpUri="https://zenodo.org/record/10520827/files/figure.png" pageId="7" pageNumber="58">Fig. 3c</figureCitation>
,
<figureCitation id="13062A0E143EA31AFEF33E14F1FDF9B4" box="[361,389,1615,1639]" captionStart="Fig" captionStartId="8.[217,252,1250,1271]" captionTargetBox="[217,1388,154,1208]" captionTargetId="figure-53@8.[217,1388,154,1208]" captionTargetPageId="8" captionText="Fig. 8 Cymopolia cf. mayaense Johnson &amp; Kaska (a-e, g-h) and Cymopolia sp. (f). a, Oblique transverse section. b, tangential section. c, tangential-oblique section. d, tangential section showing five secondaries (rectangle). e, longitudinal (above)-oblique (below) section. f, oblique transverse section. g, variously oriented sections together with Rahaghia khorassanica (Rahaghi) (left ablove). h, fragmentary tangential section showing five secondaries. Thin-sections: Nf 9 (a), Nf 6-2 (b, d, g), Nf 9 (c), Nf 8 (e), Nf 6-1 (f), Nf 4 (h)." figureDoi="http://doi.org/10.5281/zenodo.10520837" httpUri="https://zenodo.org/record/10520837/files/figure.png" pageId="7" pageNumber="58">
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).
</paragraph>
</subSubSection>
</treatment>
</document>