treatments-xml/data/03/94/87/03948799FFFEFFE9FF24FF72FE04FE1E.xml

<document id="669D36639148E5B78BB11BA4696552F7" ID-CLB-Dataset="20492" ID-DOI="10.5169/seals-787049" ID-GBIF-Dataset="ce0756e2-5184-4b74-8472-ab69fed7951c" ID-ISSN="2624-9170" ID-Zenodo-Dep="6285683" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935326602" checkinUser="carolina" docAuthor="Huber, Charles &amp; Schmidt, Joachim" docDate="2017" docId="03948799FFFEFFE9FF24FF72FE04FE1E" docLanguage="en" docName="ContribNatHist.36.1-85.pdf.imf" docOrigin="Contributions to Natural History 36" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Nebria (Epinebriola) rupina Huber &amp; Schmidt, 2017, sp. nov." docType="treatment" docVersion="12" lastPageNumber="42" masterDocId="FFADFFE1FFDBFFC0FF9CFFE9FFB0FFA7" masterDocTitle="Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="38" updateTime="1699470525451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="00E558C809102FBBAAEF3B0D65F1F560">Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)</mods:title>
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<mods:namePart id="A1A4AF0178D0CFAC8C5DA75C54D0991D">Huber, Charles</mods:namePart>
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<mods:namePart id="4759968F76B2A11A3F11ECEC17E0DAA3">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="04F86A0572DF113AC78AB983A8DFEDFD">text</mods:typeOfResource>
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<mods:title id="B2F78EF231A0906F730521B3647342A4">Contributions to Natural History</mods:title>
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<mods:part id="AF1C18064604F552565E573E9449EE38">
<mods:date id="C1273DAD3EE7F4833960A01DE181037F">2017</mods:date>
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<mods:number id="6783C8FBB995E4B74D214960B2EEC1C2">2017-07-17</mods:number>
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<subSubSection id="C3276504FFFEFFE5FF24FF72FD1FFF13" box="[184,687,155,180]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8B82368FFFFEFFE5FF24FF72FD1FFF13" blockId="37.[184,687,155,180]" box="[184,687,155,180]" pageId="37" pageNumber="38">
<heading id="D0CA81E3FFFEFFE5FF24FF72FD1FFF13" box="[184,687,155,180]" centered="true" fontSize="12" level="2" pageId="37" pageNumber="38" reason="2">
<emphasis id="B949EA9DFFFEFFE5FF24FF72FDFDFF13" bold="true" box="[184,589,155,180]" pageId="37" pageNumber="38">
<taxonomicName id="4C3D4D0CFFFEFFE5FF24FF72FE5FFF13" ID-CoL="8MFFP" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[184,495,155,180]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="rupina" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFEFFE5FF24FF72FEB4FF13" bold="true" box="[184,260,155,180]" italics="true" pageId="37" pageNumber="38">Nebria</emphasis>
(
<emphasis id="B949EA9DFFFEFFE5FE8FFF72FE25FF13" bold="true" box="[275,405,155,180]" italics="true" pageId="37" pageNumber="38">Epinebriola</emphasis>
) 
<emphasis id="B949EA9DFFFEFFE5FE39FF72FE5FFF13" bold="true" box="[421,495,155,180]" italics="true" pageId="37" pageNumber="38">rupina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFEFFE5FE69FF72FDFDFF13" box="[501,589,155,180]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13062A0AFFFEFFE5FDC5FF75FD16FF13" box="[601,678,156,180]" captionStart="Fig" captionStartId="40.[221,250,160,178]" captionTargetBox="[463,1106,153,1367]" captionTargetId="figure-100@40.[460,1108,150,1367]" captionTargetPageId="40" captionText="Fig. 21: Habitus of Nebria (Epinebriola) rupina sp. nov. Male paratype" pageId="37" pageNumber="38">Fig. 21</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3276504FFFEFFE5FF24FF0FFC04FE1E" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8B82368FFFFEFFE5FF24FF0FFDEAFE83" blockId="37.[184,1069,227,1636]" pageId="37" pageNumber="38">
<materialsCitation id="3B553CD2FFFEFFE5FF24FF0FFDEAFE83" ID-GBIF-Occurrence="3497686341" collectingDate="2001-06-24" collectionCode="cSCHM" collectorName="S. Tamang" country="Nepal" county="Rupina La" location="Manaslu Himal" municipality="Rumche Tal" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="5486882DFFFEFFE5FF24FF0FFE90FF59" box="[184,288,230,254]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3BFD06FFFEFFE5FEBAFF0AFE8AFF59" box="[294,314,227,254]" pageId="37" pageNumber="38" type="male">♂</specimenCount>
: 
<collectingCountry id="F32A761FFFFEFFE5FED4FF0FFE3FFF59" box="[328,399,230,254]" name="Nepal" pageId="37" pageNumber="38">Nepal</collectingCountry>
, 
<location id="8EE26054FFFEFFE5FE04FF0FFDF5FF59" LSID="urn:lsid:plazi:treatment:03948799FFFEFFE9FF24FF72FE04FE1E:8EE26054FFFEFFE5FE04FF0FFDF5FF59" box="[408,581,230,254]" country="Nepal" county="Rupina La" municipality="Rumche Tal" name="Manaslu Himal" pageId="37" pageNumber="38">Manaslu Himal</location>
, 
<location id="8EE26054FFFEFFE5FDD3FF0FFC13FF59" LSID="urn:lsid:plazi:treatment:03948799FFFEFFE9FF24FF72FE04FE1E:8EE26054FFFEFFE5FDD3FF0FFC13FF59" box="[591,931,230,254]" country="Nepal" county="Rupina La" municipality="Rumche Tal" name="Rumche Tal to Rupina La pass" pageId="37" pageNumber="38">Rumche Tal to Rupina La pass</location>
, 4200 mNN, 
<date id="FF83104FFFFEFFE5FF24FEE5FE87FE83" box="[184,311,268,292]" pageId="37" pageNumber="38" value="2001-06-24">
<collectingDate id="EFC7E9A7FFFEFFE5FF24FEE5FE87FE83" box="[184,311,268,292]" pageId="37" pageNumber="38" value="2001-06-24">24. 6. 2001</collectingDate>
</date>
, leg. 
<collectorName id="26C85359FFFEFFE5FEEFFEE5FE57FE83" box="[371,487,268,292]" pageId="37" pageNumber="38">S. Tamang</collectorName>
(
<collectionCode id="ED2CAE4AFFFEFFE5FE6FFEE5FDE0FE83" box="[499,592,268,292]" pageId="37" pageNumber="38">cSCHM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B82368FFFFEFFE5FF41FED8FC04FE1E" blockId="37.[184,1069,227,1636]" pageId="37" pageNumber="38">
<materialsCitation id="3B553CD2FFFEFFE5FF41FED8FC15FEEE" ID-GBIF-Occurrence="3497686311" box="[221,933,302,329]" collectingDate="2001-06-24" collectionCode="NMBE, cSCHM" collectorName="S. Tamang" country="Nepal" county="Rupina La" location="Manaslu Himal" municipality="Rumche Tal" pageId="37" pageNumber="38" specimenCount="21" specimenCount-female="7" specimenCount-male="14" typeStatus="paratype">
<typeStatus id="5486882DFFFEFFE5FF41FED8FEE6FEEE" box="[221,342,305,329]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D3BFD06FFFEFFE5FEFDFED8FE29FEEE" box="[353,409,302,329]" pageId="37" pageNumber="38" type="male">14 ♂</specimenCount>
<specimenCount id="9D3BFD06FFFEFFE5FE3DFED8FE7BFEEE" box="[417,459,302,329]" pageId="37" pageNumber="38" type="female">7 ♀</specimenCount>
, same data as holotype (
<collectionCode id="ED2CAE4AFFFEFFE5FD71FED8FC8AFEEE" box="[749,826,305,329]" collectionName="Switzerland, Bern, Naturhistorische Museums" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="37" pageNumber="38">NMBE</collectionCode>
, 
<collectionCode id="ED2CAE4AFFFEFFE5FCD9FED8FC2FFEEE" box="[837,927,305,329]" pageId="37" pageNumber="38">cSCHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B553CD2FFFEFFE5FC2CFED8FDBBFE33" ID-GBIF-Occurrence="3497686384" collectingDate="2001-06-30" collectionCode="cSCHM" collectorName="S. Tamang &amp; H. S. Galel &amp; S. Gurung" country="Nepal" elevation="4350" location="Manaslu Himal" municipality="Dudh Pokhari" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3BFD06FFFEFFE5FC2CFED8FC6AFEEE" box="[944,986,302,329]" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
<collectingCountry id="F32A761FFFFEFFE5FC7EFED8FB99FEEE" box="[994,1065,305,329]" name="Nepal" pageId="37" pageNumber="38">Nepal</collectingCountry>
, 
<location id="8EE26054FFFEFFE5FF24FEBFFED6FEC9" LSID="urn:lsid:plazi:treatment:03948799FFFEFFE9FF24FF72FE04FE1E:8EE26054FFFEFFE5FF24FEBFFED6FEC9" box="[184,358,342,366]" country="Nepal" municipality="Dudh Pokhari" name="Manaslu Himal" pageId="37" pageNumber="38">Manaslu Himal</location>
, 
<location id="8EE26054FFFEFFE5FEEDFEBFFDA0FEC9" LSID="urn:lsid:plazi:treatment:03948799FFFEFFE9FF24FF72FE04FE1E:8EE26054FFFEFFE5FEEDFEBFFDA0FEC9" box="[369,528,342,366]" country="Nepal" municipality="Dudh Pokhari" name="Dudh Pokhari" pageId="37" pageNumber="38">Dudh Pokhari</location>
, 
<quantity id="4CC59B6AFFFEFFE5FD87FEBFFD71FEC9" box="[539,705,342,366]" metricMagnitude="3" metricUnit="m" metricValue="4.35" metricValueMax="4.4" metricValueMin="4.3" pageId="37" pageNumber="38" unit="m" value="4350.0" valueMax="4400.0" valueMin="4300.0">
<elevation id="0010D1BCFFFEFFE5FD87FEBFFD71FEC9" box="[539,705,342,366]" metricMagnitude="3" metricUnit="m" metricValue="4.35" metricValueMax="4.4" metricValueMin="4.3" pageId="37" pageNumber="38" unit="m" value="4350.0" valueMax="4400.0" valueMin="4300.0">4300–4400 m</elevation>
</quantity>
, 
<date id="FF83104FFFFEFFE5FD50FEBFFCFFFEC9" box="[716,847,342,366]" pageId="37" pageNumber="38" value="2001-06-30">
<collectingDate id="EFC7E9A7FFFEFFE5FD50FEBFFCFFFEC9" box="[716,847,342,366]" pageId="37" pageNumber="38" value="2001-06-30">30. 6. 2001</collectingDate>
</date>
, leg. 
<collectorName id="26C85359FFFEFFE5FC13FEBFFBB9FEC9" box="[911,1033,342,366]" pageId="37" pageNumber="38">S. Tamang</collectorName>
, 
<collectorName id="26C85359FFFEFFE5FB88FEBFFEBDFE33" pageId="37" pageNumber="38">H. S. Galel</collectorName>
&amp; 
<collectorName id="26C85359FFFEFFE5FEB0FE95FE2BFE33" box="[300,411,380,404]" pageId="37" pageNumber="38">S. Gurung</collectorName>
(
<collectionCode id="ED2CAE4AFFFEFFE5FE34FE95FDB5FE33" box="[424,517,380,404]" pageId="37" pageNumber="38">cSCHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B553CD2FFFEFFE5FD88FE95FC1FFE1E" ID-GBIF-Occurrence="3497686398" collectingDate="1983-08-09" collectionCode="SMNS" collectorName="Martens &amp; Schawaller" country="Nepal" county="Rupina La" elevation="4450" location="Gorkha Dist." municipality="Tabruk" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3BFD06FFFEFFE5FD88FE95FD8DFE33" box="[532,573,377,404]" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
<collectingCountry id="F32A761FFFFEFFE5FDDFFE95FD3AFE33" box="[579,650,380,404]" name="Nepal" pageId="37" pageNumber="38">Nepal</collectingCountry>
, 
<location id="8EE26054FFFEFFE5FD0FFE95FC94FE33" LSID="urn:lsid:plazi:treatment:03948799FFFEFFE9FF24FF72FE04FE1E:8EE26054FFFEFFE5FD0FFE95FC94FE33" box="[659,804,380,404]" country="Nepal" county="Rupina La" municipality="Tabruk" name="Gorkha Dist." pageId="37" pageNumber="38">Gorkha Dist.</location>
, 
<location id="8EE26054FFFEFFE5FCB1FE95FF67FE1E" LSID="urn:lsid:plazi:treatment:03948799FFFEFFE9FF24FF72FE04FE1E:8EE26054FFFEFFE5FCB1FE95FF67FE1E" country="Nepal" county="Rupina La" municipality="Tabruk" name="zw. Tabruk und Rupina La" pageId="37" pageNumber="38">zw. Tabruk und Rupina La</location>
, 
<quantity id="4CC59B6AFFFEFFE5FF7CFE48FE33FE1E" box="[224,387,417,441]" metricMagnitude="3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="37" pageNumber="38" unit="m" value="4450.0" valueMax="4500.0" valueMin="4400.0">
<elevation id="0010D1BCFFFEFFE5FF7CFE48FE33FE1E" box="[224,387,417,441]" metricMagnitude="3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="37" pageNumber="38" unit="m" value="4450.0" valueMax="4500.0" valueMin="4400.0">4400–4500 m</elevation>
</quantity>
, 
<date id="FF83104FFFFEFFE5FE11FE48FDA9FE1E" box="[397,537,417,441]" pageId="37" pageNumber="38" value="1983-08-09">
<collectingDate id="EFC7E9A7FFFEFFE5FE11FE48FDA9FE1E" box="[397,537,417,441]" pageId="37" pageNumber="38" value="1983-08-09">9. Aug. 1983</collectingDate>
</date>
, leg. 
<collectorName id="26C85359FFFEFFE5FDCAFE48FD03FE1E" box="[598,691,417,441]" pageId="37" pageNumber="38">Martens</collectorName>
&amp; 
<collectorName id="26C85359FFFEFFE5FD4EFE48FCFEFE1E" box="[722,846,417,441]" pageId="37" pageNumber="38">Schawaller</collectorName>
(
<collectionCode id="ED2CAE4AFFFEFFE5FCC6FE48FC1AFE1E" box="[858,938,417,441]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="37" pageNumber="38">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276504FFFEFFE7FF41FE2FFC80FB93" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="8B82368FFFFEFFE5FF41FE2FFC39FDA3" blockId="37.[184,1069,227,1636]" pageId="37" pageNumber="38">
Body length 
<quantity id="4CC59B6AFFFEFFE5FEF2FE2FFE6AFE79" box="[366,474,454,478]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="37" pageNumber="38" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
. Colour dark brown, appendages of head and antennae brown. Pronotum brown, lighter than the head. Legs brown.
</paragraph>
<paragraph id="8B82368FFFFEFFE5FF41FDF8FCB6FC63" blockId="37.[184,1069,227,1636]" pageId="37" pageNumber="38">
Head with a shallow transverse impression behind the eyes. Labrum with anterior margin faintly bisinuate, bearing six setae. The angles of the labrum rounded but distinct. Apical margin of clypeus straight. Frons transversally wrinkled, vertex impunctate. Eyes prominent, frons above the eyes longitudinally wrinkled. Antennae long and slender extending to one third of the elytra. Antennal scape elongate, as long as the eye's diameter, basally narrowed, subcylindrical apically, with one dorsal seta. 2 
<superScript id="7C489BC7FFFEFFE5FD4CFD07FD57FCA6" attach="left" box="[720,743,750,769]" fontSize="8" pageId="37" pageNumber="38">nd</superScript>
antennomere with one seta ventroapically. Apical third of the 4 
<superScript id="7C489BC7FFFEFFE5FDC3FCFAFDC2FC81" attach="left" box="[607,626,787,806]" fontSize="8" pageId="37" pageNumber="38">th</superScript>
antennomere with a few short setae in addition to the apical collar of long setae; the additional ones sometimes hardly visible. Penultimate labial palpomere trisetose. Ligula long, spiny, with two lateral setae. Mentum with a bifid medial tooth. Submentum with a row of 12 setae. Microreticulation of the head isodiametric.
</paragraph>
<paragraph id="8B82368FFFFEFFE5FF41FC38FE56F9BE" blockId="37.[184,1069,227,1636]" pageId="37" pageNumber="38">
Pronotum faintly transverse, ratio width/length of the pronotum = 1.30. Lateral margin convex, rounded to the anterior angles, rectilinearly narrowed to the posterior angles (
<figureCitation id="13062A0AFFFEFFE5FE4CFBF5FD84FB93" box="[464,564,1052,1076]" captionStart="Fig" captionStartId="61.[184,213,1317,1335]" captionTargetBox="[185,1068,153,1276]" captionTargetId="figure-106@61.[551,733,151,716]" captionTargetPageId="61" captionText="Fig. 31: Left part of the pronotum of Epinebriola species. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) retingensis sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov.; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285816" httpUri="https://zenodo.org/record/6285816/files/figure.png" pageId="37" pageNumber="38">Fig. 31B</figureCitation>
). The posterior angles just a little narrower than the anterior angles. Lateral explanation narrow with a conspicuous groove, broadened apically and basally. Lateral margin basally obliquely upturned but not blade-like. Posterior angles acute, protruding posteriorly. Anterior angles faintly rounded, weakly protruding. Basal margin bisinuate. Pronotal disc weakly convex. Basal fovea deep, posterior transverse impression distinct, anterior transverse impression and median line shallow. Basal fovea, anterior and posterior transverse sulci and lateral groove impunctate. Apical margination of the pronotum restricted to lateral one-fourths, merges with the anterior angle. Base of the pronotum not margined. Basolateral seta present. One midlateral seta in the apical half of the pronotum, inserted in a distinct groove at the inside of the slightly upturned margin. Microreticulation of the pronotum isodiametric, impunctate on disc. Proepisternum smooth. Prosternal process flattened and unmargined.
</paragraph>
<paragraph id="8B82368FFFFEFFE7FF41F9CFFE27FF59" blockId="37.[184,1069,227,1636]" lastBlockId="39.[183,1069,156,1636]" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">Elytral silhouette slender, ellipsoid, widest behind the middle. Lateral margin subrectilinear at basal third. Shoulder distinct. Elytral apex acutely rounded. Apical carina present. Basal margination straight. Humeral carina present. Striae distinct on disc, weakly punctate. Striae obliterate towards the apex; striae 7 and 8 scarcely visible. Intervals scarcely convex, more convex on disc; interval 3 without setae. Scutellar seta absent. Microreticulation isodiametric or faintly oblong. Mesepisterna smooth. Metepisterna smooth, 1.8 times as long as wide. Metacoxa with two basal and one apical setae. Hindwings present as a strap-like vestige.</paragraph>
<caption id="DF426607FFFDFFE6FF41FACCFE34FA05" ID-DOI="http://doi.org/10.5281/zenodo.6285764" ID-Zenodo-Dep="6285764" httpUri="https://zenodo.org/record/6285764/files/figure.png" pageId="38" pageNumber="39" startId="38.[221,250,1317,1335]" targetBox="[221,1106,153,1296]" targetPageId="38">
<paragraph id="8B82368FFFFDFFE6FF41FACCFE34FA05" blockId="38.[221,1107,1317,1442]" pageId="38" pageNumber="39">
Fig. 20: Male edeagi of new 
<taxonomicName id="4C3D4D0CFFFDFFE6FE41FACCFDEFFA90" box="[477,607,1317,1335]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B949EA9DFFFDFFE6FE41FACCFDA7FA90" bold="true" box="[477,535,1317,1335]" italics="true" pageId="38" pageNumber="39">Nebria</emphasis>
species
</taxonomicName>
of the subgenus 
<taxonomicName id="4C3D4D0CFFFDFFE6FD60FACCFC42FA90" authority="K. DANIEL, 1904" authorityName="K. DANIEL" authorityYear="1904" box="[764,1010,1317,1335]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FD60FACCFCD0FA90" bold="true" box="[764,864,1317,1335]" italics="true" pageId="38" pageNumber="39">Epinebriola</emphasis>
K. 
<bibRefCitation id="EFAC4B7EFFFDFFE6FCE3FACCFC42FA90" author="Daniel, K." box="[895,1010,1317,1335]" pageId="38" pageNumber="39" pagination="1904 - 1906" refId="ref32922" refString="Daniel, K. 1904: [new taxa]. In: Daniel, K. &amp; Daniel, J.: Neue palaarktische Koleopteren. - Munchener Koleopterologische Zeitschrift 2 [1904 - 1906]: 76 - 93." type="journal article" year="1904">DANIEL, 1904</bibRefCitation>
</taxonomicName>
. A: 
<taxonomicName id="4C3D4D0CFFFDFFE6FB84FACCFE16FAF5" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="triseriata" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FB84FACCFBE2FA90" bold="true" box="[1048,1106,1317,1335]" italics="true" pageId="38" pageNumber="39">Nebria</emphasis>
(
<emphasis id="B949EA9DFFFDFFE6FF79FAA9FEF9FAF5" bold="true" box="[229,329,1344,1362]" italics="true" pageId="38" pageNumber="39">Epinebriola</emphasis>
) 
<emphasis id="B949EA9DFFFDFFE6FECAFAA9FE16FAF5" bold="true" box="[342,422,1344,1362]" italics="true" pageId="38" pageNumber="39">triseriata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFDFFE6FE37FAA9FE43FAF5" box="[427,499,1344,1362]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
; B: 
<taxonomicName id="4C3D4D0CFFFDFFE6FD88FAA9FD38FAF5" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[532,648,1344,1362]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="2" phylum="Arthropoda" rank="species" species="rupina" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FD88FAA9FD91FAF5" bold="true" box="[532,545,1344,1362]" italics="true" pageId="38" pageNumber="39">N</emphasis>
. (
<emphasis id="B949EA9DFFFDFFE6FDAFFAA9FD8EFAF5" bold="true" box="[563,574,1344,1362]" italics="true" pageId="38" pageNumber="39">E</emphasis>
.) 
<emphasis id="B949EA9DFFFDFFE6FDCCFAA9FD38FAF5" bold="true" box="[592,648,1344,1362]" italics="true" pageId="38" pageNumber="39">rupina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFDFFE6FD12FAA9FD66FAF5" box="[654,726,1344,1362]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
; C: 
<taxonomicName id="4C3D4D0CFFFDFFE6FD68FAA9FC1DFAF5" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[756,941,1344,1362]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="2" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FD68FAA9FCB1FAF5" bold="true" box="[756,769,1344,1362]" italics="true" pageId="38" pageNumber="39">N</emphasis>
. (
<emphasis id="B949EA9DFFFDFFE6FC88FAA9FCAFFAF5" bold="true" box="[788,799,1344,1362]" italics="true" pageId="38" pageNumber="39">E</emphasis>
.) 
<emphasis id="B949EA9DFFFDFFE6FCADFAA9FC1DFAF5" bold="true" box="[817,941,1344,1362]" italics="true" pageId="38" pageNumber="39">montisanimae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFDFFE6FC2EFAA9FC4AFAF5" box="[946,1018,1344,1362]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
; D: 
<taxonomicName id="4C3D4D0CFFFDFFE6FB87FAA9FE8FFACB" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="2" phylum="Arthropoda" rank="species" species="impunctata" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FB87FAA9FB98FAF5" bold="true" box="[1051,1064,1344,1362]" italics="true" pageId="38" pageNumber="39">N</emphasis>
. (
<emphasis id="B949EA9DFFFDFFE6FBA6FAA9FBF5FAF5" bold="true" box="[1082,1093,1344,1362]" italics="true" pageId="38" pageNumber="39">E</emphasis>
.) 
<emphasis id="B949EA9DFFFDFFE6FF41FAB3FE8FFACB" bold="true" box="[221,319,1370,1388]" italics="true" pageId="38" pageNumber="39">impunctata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFDFFE6FED9FAB3FE3EFACB" box="[325,398,1370,1388]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
; E: 
<taxonomicName id="4C3D4D0CFFFDFFE6FE32FAB3FDF7FACB" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[430,583,1370,1388]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="2" phylum="Arthropoda" rank="species" species="numburica" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FE32FAB3FE0BFACB" bold="true" box="[430,443,1370,1388]" italics="true" pageId="38" pageNumber="39">N</emphasis>
. (
<emphasis id="B949EA9DFFFDFFE6FE51FAB3FE68FACB" bold="true" box="[461,472,1370,1388]" italics="true" pageId="38" pageNumber="39">E</emphasis>
.) 
<emphasis id="B949EA9DFFFDFFE6FE77FAB3FDF7FACB" bold="true" box="[491,583,1370,1388]" italics="true" pageId="38" pageNumber="39">numburica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFDFFE6FDD1FAB3FD24FACB" box="[589,660,1370,1388]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, with parameres; F: 
<taxonomicName id="4C3D4D0CFFFDFFE6FCD4FAB3FC57FACB" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[840,999,1370,1388]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="2" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FCD4FAB3FCE5FACB" bold="true" box="[840,853,1370,1388]" italics="true" pageId="38" pageNumber="39">N</emphasis>
. (
<emphasis id="B949EA9DFFFDFFE6FCFBFAB3FCC2FACB" bold="true" box="[871,882,1370,1388]" italics="true" pageId="38" pageNumber="39">E</emphasis>
.) 
<emphasis id="B949EA9DFFFDFFE6FC19FAB3FC57FACB" bold="true" box="[901,999,1370,1388]" italics="true" pageId="38" pageNumber="39">retingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFDFFE6FC71FAB3FB86FACB" box="[1005,1078,1370,1388]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
; G: 
<taxonomicName id="4C3D4D0CFFFDFFE6FF41FA9CFEC0FA20" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[221,368,1397,1415]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="2" phylum="Arthropoda" rank="species" species="incarinata" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FF41FA9CFF5AFA20" bold="true" box="[221,234,1397,1415]" italics="true" pageId="38" pageNumber="39">N</emphasis>
. (
<emphasis id="B949EA9DFFFDFFE6FF60FA9CFEB7FA20" bold="true" box="[252,263,1397,1415]" italics="true" pageId="38" pageNumber="39">E</emphasis>
.) 
<emphasis id="B949EA9DFFFDFFE6FE84FA9CFEC0FA20" bold="true" box="[280,368,1397,1415]" italics="true" pageId="38" pageNumber="39">incarinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFDFFE6FEE8FA9CFE0CFA20" box="[372,444,1397,1415]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
; H: 
<taxonomicName id="4C3D4D0CFFFDFFE6FE47FA9CFDECFA20" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[475,604,1397,1415]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="2" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FE47FA9CFE58FA20" bold="true" box="[475,488,1397,1415]" italics="true" pageId="38" pageNumber="39">N</emphasis>
. (
<emphasis id="B949EA9DFFFDFFE6FE66FA9CFDB5FA20" bold="true" box="[506,517,1397,1415]" italics="true" pageId="38" pageNumber="39">E</emphasis>
.) 
<emphasis id="B949EA9DFFFDFFE6FD8AFA9CFDECFA20" bold="true" box="[534,604,1397,1415]" italics="true" pageId="38" pageNumber="39">delicata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFDFFE6FDFDFA9CFD18FA20" box="[609,680,1397,1415]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
; I: 
<taxonomicName id="4C3D4D0CFFFDFFE6FD5DFA9CFCE4FA20" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[705,852,1397,1415]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="2" phylum="Arthropoda" rank="species" species="incarinata" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFFDFFE6FD5DFA9CFD7EFA20" bold="true" box="[705,718,1397,1415]" italics="true" pageId="38" pageNumber="39">N</emphasis>
. (
<emphasis id="B949EA9DFFFDFFE6FD43FA9CFD5AFA20" bold="true" box="[735,746,1397,1415]" italics="true" pageId="38" pageNumber="39">E</emphasis>
.) 
<emphasis id="B949EA9DFFFDFFE6FD60FA9CFCE4FA20" bold="true" box="[764,852,1397,1415]" italics="true" pageId="38" pageNumber="39">incarinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFDFFE6FCC4FA9CFC2EFA20" box="[856,926,1397,1415]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, apex in dorsal view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8B82368FFFFCFFE7FF41FEE5FE17FEC9" blockId="39.[183,1069,156,1636]" pageId="39" pageNumber="40">Sternum 3 medially asetose. Sterna 4–6 each with 4–5 posterior paramedial setae. Anal sternum with two paramedial setae in the male. All sterna with faint impressions laterally.</paragraph>
<paragraph id="8B82368FFFFCFFE7FF41FE95FD11FDA3" blockId="39.[183,1069,156,1636]" pageId="39" pageNumber="40">Legs long and slender. All tarsi dorsally glabrous. Metatarsomere 4 with a ventroapical tooth. Meso- and metatarsomeres 2–4 ventroapically with long setae. All tarsomeres 5 ventrally with two rows of 4–5 short setae. Metatarsomere 5 as long as the metatarsomeres 3+4.</paragraph>
<paragraph id="8B82368FFFFCFFE7FF41FDF8FE6DFD19" blockId="39.[183,1069,156,1636]" pageId="39" pageNumber="40">
Male genitalia: Edeagus (
<figureCitation id="13062A0AFFFCFFE7FE66FDF8FDEDFD8E" box="[506,605,529,553]" captionStart="Fig" captionStartId="38.[221,250,1317,1335]" captionTargetBox="[221,1106,153,1296]" captionTargetId="figure-70@38.[221,704,627,867]" captionTargetPageId="38" captionText="Fig. 20: Male edeagi of new Nebria species of the subgenus Epinebriola K. DANIEL, 1904. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) numburica sp. nov., with parameres; F: N. (E.) retingensis sp. nov.; G: N. (E.) incarinata sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov., apex in dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285764" httpUri="https://zenodo.org/record/6285764/files/figure.png" pageId="39" pageNumber="40">Fig. 20B</figureCitation>
): Basal part of the median lobe wide, the basolateral lobes small. Mid-shaft slender, strongly curved at base, moderately curved and progressively narrowed to the apex. Apex straight, in front of the tip on the ventral side faintly convex. Tip of the apex long and slender. Apex deflected to the left.
</paragraph>
<paragraph id="8B82368FFFFCFFE7FF41FD25FD2AFC39" blockId="39.[183,1069,156,1636]" pageId="39" pageNumber="40">
Female genitalia: Gonocoxa (
<figureCitation id="13062A0AFFFCFFE7FDB0FD25FD21FD43" box="[556,657,716,740]" captionStart="Fig" captionStartId="20.[221,250,1430,1448]" captionTargetBox="[228,1106,153,1365]" captionTargetId="figure-16@20.[574,757,751,1260]" captionTargetPageId="20" captionText="Fig. 10: Right gonocoxae of Epinebriola species, in ventral view. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) numburica sp. nov. in lateral view with the distale gonocoxite 2 dorsally grooved (arrow); H: N. (E.) christinae HUBER &amp; SCHMIDT, 2007; I: N. (E.) pseudorestias sp. nov.; J: N. (E.) retingensis sp. nov.; Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285726" httpUri="https://zenodo.org/record/6285726/files/figure.png" pageId="39" pageNumber="40">Fig. 10B</figureCitation>
): Gonocoxite 1 slight and relatively short, gonocoxite 2 elongate-triangular, two-thirds the length of gonocoxite 1, slightly narrowed near base. Apex acuminate, dorsally arcuate, dorsally faintly grooved. Ventral preapical insertion furrow elongate-oval, with two nematiforme setae. Gonocoxites ventrally unjointed and continuously sclerotized, dorsally separated by a membranous area.
</paragraph>
<paragraph id="8B82368FFFFCFFE7FF41FC45FC80FB93" blockId="39.[183,1069,156,1636]" pageId="39" pageNumber="40">Body ratios: hea.w/fro.w = 1.51±0.024 (1.48–1.55); prm.w/hea.w = 1.16± 0.018 (1.15–1.19); prm.w/prp.w = 1.53±0.050 (1.45–1.59); pra.w/prp.w = 1.13± 0.042 (1.06–1.19); ely.w/prm.w = 1.61±0.046 (1.56–1.70); prm.w/prm.l = 1.30± 0.032 (1.25–1.33); ely.l/ely.w = 1.63± 0.030 (1.59–1.68).</paragraph>
</subSubSection>
<subSubSection id="C3276504FFFCFFE7FF41FBA8FE33FBD9" pageId="39" pageNumber="40" type="etymology">
<paragraph id="8B82368FFFFCFFE7FF41FBA8FE33FBD9" blockId="39.[183,1069,156,1636]" pageId="39" pageNumber="40">
Etymology: The specific epithet (noun in apposition) refers to the 
<typeStatus id="5486882DFFFCFFE7FC64FBA8FB9CFBFE" box="[1016,1068,1089,1113]" pageId="39" pageNumber="40">type</typeStatus>
locality Rupina La.
</paragraph>
</subSubSection>
<subSubSection id="C3276504FFFCFFE7FF41FB65FBB0F9C3" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8B82368FFFFCFFE7FF41FB65FBB0F9C3" blockId="39.[183,1069,156,1636]" pageId="39" pageNumber="40">
Diagnosis: 
<taxonomicName id="4C3D4D0CFFFCFFE7FEC4FB65FE70FB03" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[344,448,1164,1188]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="rupina" status="sp. nov.">
<emphasis id="B949EA9DFFFCFFE7FEC4FB65FE70FB03" box="[344,448,1164,1188]" italics="true" pageId="39" pageNumber="40">N. rupina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFCFFE7FE5BFB65FDAEFB03" box="[455,542,1164,1188]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
stands out within the subgenus 
<taxonomicName id="4C3D4D0CFFFCFFE7FC10FB65FBBAFB03" authorityName="K. DANIEL" authorityYear="1904" box="[908,1034,1164,1188]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
<emphasis id="B949EA9DFFFCFFE7FC10FB65FBBAFB03" box="[908,1034,1164,1188]" italics="true" pageId="39" pageNumber="40">Epinebriola</emphasis>
</taxonomicName>
by the additional apical pubescence of the 4 
<superScript id="7C489BC7FFFCFFE7FDE6FB47FD3CFB66" attach="left" box="[634,652,1198,1217]" fontSize="8" pageId="39" pageNumber="40">th</superScript>
antennomere and the presence of the basolateral seta of the pronotum. An apical pubescence of the 4 
<superScript id="7C489BC7FFFCFFE7FC1EFB3AFC24FB41" attach="left" box="[898,916,1235,1254]" fontSize="8" pageId="39" pageNumber="40">th</superScript>
antennomere is rarely present in the sympatric species 
<taxonomicName id="4C3D4D0CFFFCFFE7FD1FFB15FCF0FAB3" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[643,832,1276,1300]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
<emphasis id="B949EA9DFFFCFFE7FD1FFB15FCF0FAB3" box="[643,832,1276,1300]" italics="true" pageId="39" pageNumber="40">N. montisanimae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFCFFE7FCDBFB15FC2DFAB3" box="[839,925,1276,1300]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
(see below), but the basolateral seta of the pronotum is absent. Furthermore, the lateral margin of the pronotum is rounded to the anterior angles in a lesser extent than in than 
<taxonomicName id="4C3D4D0CFFFCFFE7FF6DFA85FE1EFA23" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[241,430,1388,1412]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
<emphasis id="B949EA9DFFFCFFE7FF6DFA85FE1EFA23" box="[241,430,1388,1412]" italics="true" pageId="39" pageNumber="40">N. montisanimae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFCFFE7FE29FA85FDBFFA23" box="[437,527,1388,1412]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, and the apex of the anterior angle of the pronotum is without ridge along the lateral groove (in 
<taxonomicName id="4C3D4D0CFFFCFFE7FD7CFA78FC10FA0E" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[736,928,1425,1449]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
<emphasis id="B949EA9DFFFCFFE7FD7CFA78FC10FA0E" box="[736,928,1425,1449]" italics="true" pageId="39" pageNumber="40">N. montisanimae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFCFFE7FC34FA78FBB0FA0E" box="[936,1024,1425,1449]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
the apical margination of the pronotum merges with a distinct rigde which continues to the lateral seta). In 
<taxonomicName id="4C3D4D0CFFFCFFE7FE31FA35FDA4FA53" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[429,532,1500,1524]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="rupina" status="sp. nov.">
<emphasis id="B949EA9DFFFCFFE7FE31FA35FDA4FA53" box="[429,532,1500,1524]" italics="true" pageId="39" pageNumber="40">N. rupina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFCFFE7FD86FA35FDC1FA53" box="[538,625,1500,1524]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
the scutellar setae and and the setae of the 3 
<superScript id="7C489BC7FFFCFFE7FF69FA17FEB8F9B6" attach="left" box="[245,264,1534,1553]" fontSize="8" pageId="39" pageNumber="40">rd</superScript>
elytral interval are absent (present in 
<taxonomicName id="4C3D4D0CFFFCFFE7FD4CF9E8FC26F9BE" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[720,918,1537,1561]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
<emphasis id="B949EA9DFFFCFFE7FD4CF9E8FC26F9BE" box="[720,918,1537,1561]" italics="true" pageId="39" pageNumber="40">N. montisanimae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFCFFE7FC02F9E8FBB0F9BE" box="[926,1024,1537,1561]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
), 
<taxonomicName id="4C3D4D0CFFFCFFE7FB88F9E8FEB0F999" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="rupina" status="sp. nov.">
<emphasis id="B949EA9DFFFCFFE7FB88F9E8FEB0F999" italics="true" pageId="39" pageNumber="40">N. rupina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFCFFE7FE99F9CFFEEBF999" box="[261,347,1574,1598]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
is distinctly smaller than 
<taxonomicName id="4C3D4D0CFFFCFFE7FDECF9CFFC9CF999" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[624,812,1574,1598]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
<emphasis id="B949EA9DFFFCFFE7FDECF9CFFC9CF999" box="[624,812,1574,1598]" italics="true" pageId="39" pageNumber="40">N. montisanimae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFCFFE7FCADF9CFFC3BF999" box="[817,907,1574,1598]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, and the elytra are elongate-ovoid in contrast to the ovoid elytra in 
<taxonomicName id="4C3D4D0CFFFCFFE7FD77F9A5FC16F9C3" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[747,934,1612,1636]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
<emphasis id="B949EA9DFFFCFFE7FD77F9A5FC16F9C3" box="[747,934,1612,1636]" italics="true" pageId="39" pageNumber="40">N. montisanimae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFFCFFE7FC37F9A5FBB0F9C3" box="[939,1024,1612,1636]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<paragraph id="8B82368FFFF3FFE8FF41FF49FE83FEA5" blockId="40.[221,399,160,258]" pageId="40" pageNumber="41">
<emphasis id="B949EA9DFFF3FFE8FF41FF49FECBFF15" bold="true" box="[221,379,160,178]" pageId="40" pageNumber="41">Fig. 21: Habitus of</emphasis>
<taxonomicName id="4C3D4D0CFFF3FFE8FF41FF52FE3FFF6A" authority="" authorityName="K. DANIEL" authorityYear="1904" box="[221,399,187,205]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
<emphasis id="B949EA9DFFF3FFE8FF41FF52FEA7FF6A" bold="true" box="[221,279,187,205]" italics="true" pageId="40" pageNumber="41">Nebria</emphasis>
(
<emphasis id="B949EA9DFFF3FFE8FEBFFF52FE37FF6A" bold="true" box="[291,391,187,205]" italics="true" pageId="40" pageNumber="41">Epinebriola</emphasis>
)
</taxonomicName>
<emphasis id="B949EA9DFFF3FFE8FF41FF3CFE3DFF40" bold="true" box="[221,397,213,231]" pageId="40" pageNumber="41">
<taxonomicName id="4C3D4D0CFFF3FFE8FF41FF3CFEA5FF40" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[221,277,213,231]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="rupina" status="sp. nov.">
<emphasis id="B949EA9DFFF3FFE8FF41FF3CFEA5FF40" bold="true" box="[221,277,213,231]" italics="true" pageId="40" pageNumber="41">rupina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFF3FFE8FE86FF3CFEEEFF40" box="[282,350,213,231]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
Male
</emphasis>
<typeStatus id="5486882DFFF3FFE8FF41FF19FE9FFEA5" box="[221,303,240,258]" pageId="40" pageNumber="41" type="paratype">paratype</typeStatus>
<emphasis id="B949EA9DFFF3FFE8FEB3FF19FE83FEA5" bold="true" box="[303,307,240,258]" pageId="40" pageNumber="41">.</emphasis>
</paragraph>
<subSubSection id="C3276504FFF3FFE9FE9FFA78FC86FE83" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8B82368FFFF3FFE9FE9FFA78FC86FE83" blockId="40.[221,1106,1422,1636]" lastBlockId="41.[184,1069,156,441]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
Remarks: The apical pubescence of the 4 
<superScript id="7C489BC7FFF3FFE8FD57FA67FD6EFA06" attach="left" box="[715,734,1422,1441]" fontSize="8" pageId="40" pageNumber="41">th</superScript>
antennomere is rare in the genus 
<taxonomicName id="4C3D4D0CFFF3FFE8FF41FA5FFE97FA69" authorityName="LATREILLE" authorityYear="1802" box="[221,295,1462,1486]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA9DFFF3FFE8FF41FA5FFE97FA69" box="[221,295,1462,1486]" italics="true" pageId="40" pageNumber="41">Nebria</emphasis>
</taxonomicName>
. It is restricted to the subgenus 
<taxonomicName id="4C3D4D0CFFF3FFE8FD06FA5FFCA4FA69" authorityName="Jeannel" authorityYear="1937" box="[666,788,1462,1486]" class="Insecta" family="Carabidae" genus="Paranebria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA9DFFF3FFE8FD06FA5FFCA4FA69" box="[666,788,1462,1486]" italics="true" pageId="40" pageNumber="41">Paranebria</emphasis>
</taxonomicName>
(two species), to most species of the Himalayan subgenus 
<taxonomicName id="4C3D4D0CFFF3FFE8FDCDFA35FD43FA53" authorityName="Banninger" authorityYear="1923" box="[593,755,1500,1524]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Patrobonebria">
<emphasis id="B949EA9DFFF3FFE8FDCDFA35FD43FA53" box="[593,755,1500,1524]" italics="true" pageId="40" pageNumber="41">Patrobonebria</emphasis>
</taxonomicName>
, and to a few isolated species in different subgenera (
<bibRefCitation id="EFAC4B7EFFF3FFE8FE78F9E8FD63F9BE" author="Ledoux, G. &amp; Roux, P." box="[484,723,1537,1561]" pageId="40" pageNumber="41" refId="ref33536" refString="Ledoux, G. &amp; Roux, P. (2005): Nebria (Coleoptera, Nebriidae) Faune mondiale. - 976 pp., Saint-Just-la-Pendue." type="book" year="2005">Ledoux &amp; Roux 2005</bibRefCitation>
). It has not been reported for the subgenus 
<taxonomicName id="4C3D4D0CFFF3FFE8FECCF9CFFE7FF999" authorityName="K. DANIEL" authorityYear="1904" box="[336,463,1574,1598]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
<emphasis id="B949EA9DFFF3FFE8FECCF9CFFE7FF999" box="[336,463,1574,1598]" italics="true" pageId="40" pageNumber="41">Epinebriola</emphasis>
</taxonomicName>
so far. In 
<taxonomicName id="4C3D4D0CFFF3FFE8FDA0F9CFFD14F999" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[572,676,1574,1598]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="rupina" status="sp. nov.">
<emphasis id="B949EA9DFFF3FFE8FDA0F9CFFD14F999" box="[572,676,1574,1598]" italics="true" pageId="40" pageNumber="41">N. rupina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFF3FFE8FD36F9CFFCB0F999" box="[682,768,1574,1598]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
as well as in 
<taxonomicName id="4C3D4D0CFFF3FFE8FC08F9CFFBE2F999" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[916,1106,1574,1598]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
<emphasis id="B949EA9DFFF3FFE8FC08F9CFFBE2F999" box="[916,1106,1574,1598]" italics="true" pageId="40" pageNumber="41">N. montisanimae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFF3FFE8FF41F9A5FE83F9C3" box="[221,307,1612,1636]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
several characters speak for an assignment to the subgenus 
<taxonomicName id="4C3D4D0CFFF3FFE8FC4EF9A5FBE1F9C3" authorityName="K. DANIEL" authorityYear="1904" box="[978,1105,1612,1636]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
<emphasis id="B949EA9DFFF3FFE8FC4EF9A5FBE1F9C3" box="[978,1105,1612,1636]" italics="true" pageId="40" pageNumber="41">Epinebriola</emphasis>
</taxonomicName>
(and not to 
<taxonomicName id="4C3D4D0CFFF2FFE9FEA5FF75FE6AFF13" authorityName="Banninger" authorityYear="1923" box="[313,474,156,180]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Patrobonebria">
<emphasis id="B949EA9DFFF2FFE9FEA5FF75FE6AFF13" box="[313,474,156,180]" italics="true" pageId="41" pageNumber="42">Patrobonebria</emphasis>
</taxonomicName>
) such as the weak head constriction behind the eyes (distinct in 
<taxonomicName id="4C3D4D0CFFF2FFE9FEA5FF28FE6BFF7E" authorityName="Banninger" authorityYear="1923" box="[313,475,193,217]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Patrobonebria">
<emphasis id="B949EA9DFFF2FFE9FEA5FF28FE6BFF7E" box="[313,475,193,217]" italics="true" pageId="41" pageNumber="42">Patrobonebria</emphasis>
</taxonomicName>
), the impunctate head (punctate in 
<taxonomicName id="4C3D4D0CFFF2FFE9FCE6FF28FBACFF7E" authorityName="Banninger" authorityYear="1923" box="[890,1052,193,217]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Patrobonebria">
<emphasis id="B949EA9DFFF2FFE9FCE6FF28FBACFF7E" box="[890,1052,193,217]" italics="true" pageId="41" pageNumber="42">Patrobonebria</emphasis>
</taxonomicName>
), the penultimate trisetose palpomere (bisetose in 
<taxonomicName id="4C3D4D0CFFF2FFE9FD42FF0FFCCFFF59" authorityName="Banninger" authorityYear="1923" box="[734,895,230,254]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Patrobonebria">
<emphasis id="B949EA9DFFF2FFE9FD42FF0FFCCFFF59" box="[734,895,230,254]" italics="true" pageId="41" pageNumber="42">Patrobonebria</emphasis>
</taxonomicName>
), and the three dilated protarsomeres in the male (two in 
<taxonomicName id="4C3D4D0CFFF2FFE9FD18FEE5FC95FE83" authorityName="Banninger" authorityYear="1923" box="[644,805,268,292]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Patrobonebria">
<emphasis id="B949EA9DFFF2FFE9FD18FEE5FC95FE83" box="[644,805,268,292]" italics="true" pageId="41" pageNumber="42">Patrobonebria</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3276504FFF2FFE9FF41FED8FDCBFE33" pageId="41" pageNumber="42" type="distribution">
<paragraph id="8B82368FFFF2FFE9FF41FED8FDCBFE33" blockId="41.[184,1069,156,441]" pageId="41" pageNumber="42">
Distribution (
<figureCitation id="13062A0AFFF2FFE9FEF2FED8FE0AFEEE" box="[366,442,305,329]" captionStart="Fig" captionStartId="76.[221,250,160,178]" captionTargetId="figure-8@75.[184,1068,152,1645]" captionTargetPageId="75" captionText="Fig. 41: Locations with occurrences of Nebria (Epinebriola) species from the Central and Eastern Himalaya and from South Tibet." figureDoi="http://doi.org/10.5281/zenodo.6285892" httpUri="https://zenodo.org/record/6285892/files/figure.png" pageId="41" pageNumber="42">Fig. 41</figureCitation>
): Known only from the 
<typeStatus id="5486882DFFF2FFE9FD5DFED8FD43FEEE" box="[705,755,305,329]" pageId="41" pageNumber="42">type</typeStatus>
locality, the pass Rupina La on eastern slope of Himal Chuli (eastern part of the Manaslu Himal mountain range), western Central 
<collectingCountry id="F32A761FFFF2FFE9FE58FE95FDB5FE33" box="[452,517,380,404]" name="Nepal" pageId="41" pageNumber="42">Nepal</collectingCountry>
Himalaya.
</paragraph>
</subSubSection>
<subSubSection id="C3276504FFF2FFE9FF41FE48FE04FE1E" box="[221,436,417,441]" pageId="41" pageNumber="42" type="biology_ecology">
<paragraph id="8B82368FFFF2FFE9FF41FE48FE04FE1E" blockId="41.[184,1069,156,441]" box="[221,436,417,441]" pageId="41" pageNumber="42">Habitat: Not noted.</paragraph>
</subSubSection>
</treatment>
</document>