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<document id="F8088B986D88EB37D2810F3437B488DA" ID-CLB-Dataset="26386" ID-DOI="10.11646/zootaxa.4624.1.3" ID-GBIF-Dataset="ef166e01-240e-4c35-a6b5-79da690f5d24" ID-ISSN="1175-5326" ID-Zenodo-Dep="3258458" ID-ZooBank="84AA439F-5A44-495B-A981-81ED53173690" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561648432921" checkinUser="plazi" docAuthor="Bochenski, Zbigniew M., Wertz, Krzysztof, Tomek, Teresa &amp; Gorobets, Leonid" docDate="2019" docId="842887D0EF30813FFF3EFE223F9E128F" docLanguage="en" docName="zootaxa.4624.1.3.pdf" docOrigin="Zootaxa 4624 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cherevychnavis umanskae Bochenski &amp; Wertz &amp; Tomek &amp; Gorobets 2019, gen. et sp. nov." docType="treatment" docVersion="12" lastPageNumber="53" masterDocId="7811FFA8EF3C8133FFA9FFFF3E5D1648" masterDocTitle="A new species of the late Miocene charadriiform bird (Aves: Charadriiformes), with a summary of all Paleogene and Miocene Charadrii remains" masterLastPageNumber="58" masterPageNumber="41" pageNumber="53" updateTime="1698737317490" updateUser="ExternalLinkService">
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<mods:title id="E3ED6DFC15716C9955DE77810073486E">A new species of the late Miocene charadriiform bird (Aves: Charadriiformes), with a summary of all Paleogene and Miocene Charadrii remains</mods:title>
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<mods:namePart id="80541F591462E9CB814791805EC14719">Bochenski, Zbigniew M.</mods:namePart>
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<mods:namePart id="FDB5C90CE5A58703710EA070951F3DDC">Wertz, Krzysztof</mods:namePart>
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<mods:namePart id="BE0B2D41139F99FE7F684DD430DE03F0">Tomek, Teresa</mods:namePart>
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<mods:namePart id="19D1AE3FC01D8E94B8D2DAA323EE5789">Gorobets, Leonid</mods:namePart>
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<treatment id="842887D0EF30813FFF3EFE223F9E128F" ID-DOI="http://doi.org/10.5281/zenodo.5931128" ID-GBIF-Taxon="158407528" ID-Zenodo-Dep="5931128" LSID="urn:lsid:plazi:treatment:842887D0EF30813FFF3EFE223F9E128F" httpUri="http://treatment.plazi.org/id/842887D0EF30813FFF3EFE223F9E128F" lastPageNumber="53" pageId="12" pageNumber="53">
<subSubSection id="449B654DEF30813FFF3EFE223CC517BF" box="[151,664,477,504]" pageId="12" pageNumber="53" type="nomenclature">
<paragraph id="0C3E36C6EF30813FFF3EFE223CC517BF" blockId="12.[151,664,477,504]" box="[151,664,477,504]" pageId="12" pageNumber="53">
<heading id="577681AAEF30813FFF3EFE223CC517BF" bold="true" box="[151,664,477,504]" fontSize="11" level="1" pageId="12" pageNumber="53" reason="1">
<emphasis id="3EF5EAD4EF30813FFF3EFE223CC517BF" bold="true" box="[151,664,477,504]" pageId="12" pageNumber="53">
<taxonomicName id="CB814D45EF30813FFF3EFE223F8617B0" authority="Bochenski &amp; Wertz &amp; Tomek &amp; Gorobets, 2019" authorityName="Bochenski &amp; Wertz &amp; Tomek &amp; Gorobets" authorityYear="2019" box="[151,475,477,504]" class="Aves" family="Haematopodidae" genus="Cherevychnavis" kingdom="Animalia" order="Charadriiformes" pageId="12" pageNumber="53" phylum="Chordata" rank="species" species="umanskae" status="gen. et sp. nov.">
<emphasis id="3EF5EAD4EF30813FFF3EFE223F8617B0" bold="true" box="[151,475,477,504]" italics="true" pageId="12" pageNumber="53">Cherevychnavis umanskae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25C657AFEF30813FFE4BFE223CC517BF" box="[482,664,477,503]" pageId="12" pageNumber="53" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="449B654DEF30813FFF3EFDDA3FE714CF" pageId="12" pageNumber="53" type="etymology">
<paragraph id="0C3E36C6EF30813FFF3EFDDA3FE714CF" blockId="12.[151,1437,549,1223]" pageId="12" pageNumber="53">
<emphasis id="3EF5EAD4EF30813FFF3EFDDA3F431477" bold="true" box="[151,286,549,575]" pageId="12" pageNumber="53">Etymology.</emphasis>
The species is named after Alla Semenivna Umanska (Umanskaya—in publications in Russian), a female Ukrainian palaeontologist, who first realized that the specimens represent an extinct species, but passed away before describing it.
</paragraph>
</subSubSection>
<subSubSection id="449B654DEF30813FFF6EFD6E3FE4125B" pageId="12" pageNumber="53" type="materials_examined">
<paragraph id="0C3E36C6EF30813FFF6EFD6E3AF81487" blockId="12.[151,1437,549,1223]" pageId="12" pageNumber="53">
<materialsCitation id="BCE93C9BEF30813FFF6EFD6E3AF81487" ID-GBIF-Occurrence="2270703908" collectionCode="NMNHU-P" country="Ukraine" location="National Academy of Sciences" municipality="National Museum of Natural History" pageId="12" pageNumber="53" specimenCount="1" stateProvince="Kyiv" typeStatus="holotype">
<emphasis id="3EF5EAD4EF30813FFF6EFD6E3F6614E3" bold="true" box="[199,315,657,683]" pageId="12" pageNumber="53">
<typeStatus id="D33A8864EF30813FFF6EFD6E3F6B14E3" box="[199,310,657,683]" pageId="12" pageNumber="53" type="holotype">Holotype</typeStatus>
.
</emphasis>
An almost complete left coracoid,
<collectionCode id="6A90AE03EF30813FFD7DFD6E3D0A14E3" box="[724,855,657,683]" pageId="12" pageNumber="53">NMNHU-P</collectionCode>
, no.45-2453 (
<figureCitation id="94BA2A43EF30813FFC5EFD6E3A3814E4" box="[1015,1125,657,684]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,250,1834,1859]" captionStartId-1="4.[151,250,1834,1859]" captionStartId-2="5.[151,250,1690,1715]" captionStartId-3="6.[151,250,1834,1859]" captionStartId-4="7.[151,250,1798,1823]" captionStartId-5="8.[151,250,1722,1747]" captionTargetBox-0="[151,1435,417,1815]" captionTargetBox-1="[151,1435,417,1815]" captionTargetBox-2="[151,1435,273,1671]" captionTargetBox-3="[151,1435,417,1815]" captionTargetBox-4="[151,1435,381,1779]" captionTargetBox-5="[151,1435,669,1702]" captionTargetId-0="figure@3.[151,1435,417,1815]" captionTargetId-1="figure@4.[151,1435,417,1815]" captionTargetId-2="figure@5.[151,1435,273,1671]" captionTargetId-3="figure@6.[151,1435,417,1815]" captionTargetId-4="figure@7.[151,1435,381,1779]" captionTargetId-5="figure@8.[151,1435,669,1702]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1. Holotype of Cherevychnavis umanskae sp. nov. (NMNHU-P, no.45-2453, left coracoid) in comparison to extant Charadrii species. Scale bars: 10 mm." captionText-1="FIGURE 2. Coracoid, characters #17 (see text for details) that differentiate fossil Cherevychnavis umanskae from extant Charadrii genera." captionText-2="FIGURE 3. Coracoid, characters #811 (see text for details) that differentiate fossil Cherevychnavis umanskae from extant Charadrii genera." captionText-3="FIGURE 4. Coracoid, characters #1215 (see text for details) that differentiate fossil Cherevychnavis umanskae from extant Charadrii genera." captionText-4="FIGURE 5. Coracoid, characters #1619 (see text for details) that differentiate fossil Cherevychnavis umanskae from extant Charadrii genera." captionText-5="FIGURE 6. Coracoid, characters #2022 (see text for details) that differentiate fossil Cherevychnavis umanskae from extant Charadrii genera." figureDoi-0="http://doi.org/10.5281/zenodo.3416779" figureDoi-1="http://doi.org/10.5281/zenodo.3416781" figureDoi-2="http://doi.org/10.5281/zenodo.3419390" figureDoi-3="http://doi.org/10.5281/zenodo.3419392" figureDoi-4="http://doi.org/10.5281/zenodo.3419396" httpUri-0="https://zenodo.org/record/3416779/files/figure.png" httpUri-1="https://zenodo.org/record/3416781/files/figure.png" httpUri-2="https://zenodo.org/record/3419390/files/figure.png" httpUri-3="https://zenodo.org/record/3419392/files/figure.png" httpUri-4="https://zenodo.org/record/3419396/files/figure.png" httpUri-5="https://zenodo.org/record/3258460/files/figure.png" pageId="12" pageNumber="53">Figs. 16</figureCitation>
), deposited at the National Museum of Natural History at the National Academy of Sciences of Ukraine, Kyiv, Ukraine.
</materialsCitation>
</paragraph>
<paragraph id="0C3E36C6EF30813FFF6EFD273BDD15CB" blockId="12.[151,1437,549,1223]" pageId="12" pageNumber="53">
<emphasis id="3EF5EAD4EF30813FFF6EFD273FFA14BB" bold="true" box="[199,423,728,755]" pageId="12" pageNumber="53">Referred material.</emphasis>
Proximal left humerus with shaft,
<collectionCode id="6A90AE03EF30813FFC9EFD263DE714BB" box="[823,954,729,755]" pageId="12" pageNumber="53">NMNHU-P</collectionCode>
, no.45-2452 (
<figureCitation id="94BA2A43EF30813FFBFEFD263A8914BB" box="[1111,1236,729,755]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="9.[151,250,1816,1841]" captionStartId-1="10.[151,250,1738,1763]" captionStartId-2="11.[151,250,1978,2003]" captionStartId-3="13.[151,250,1798,1823]" captionTargetBox-0="[151,1435,360,1779]" captionTargetBox-1="[151,1435,288,1707]" captionTargetBox-2="[151,1435,561,1959]" captionTargetBox-3="[151,1435,381,1779]" captionTargetId-0="figure@9.[151,1435,381,1779]" captionTargetId-1="figure@10.[151,1435,309,1707]" captionTargetId-2="figure@11.[151,1435,561,1959]" captionTargetId-3="figure@13.[151,1435,381,1779]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="13" captionText-0="FIGURE 7. Referred material of Cherevychnavis umanskae sp. nov. (NMNHU-P, no.45-2452, proximal left humerus) in comparison to extant Charadrii species. Scale bars: 10mm." captionText-1="FIGURE 8. Humerus, characters #17 (see text for details) that differentiate fossil Cherevychnavis umanskae from extant Charadrii genera." captionText-2="FIGURE 9. Humerus, characters #89 (see text for details) that differentiate fossil Cherevychnavis umanskae from extant Charadrii genera." captionText-3="FIGURE 10. Humerus, characters #1014 (see text for details) that differentiate fossil Cherevychnavis umanskae from extant Charadrii genera." figureDoi-2="http://doi.org/10.5281/zenodo.3419400" figureDoi-3="http://doi.org/10.5281/zenodo.3419404" httpUri-0="https://zenodo.org/record/3258462/files/figure.png" httpUri-1="https://zenodo.org/record/3258464/files/figure.png" httpUri-2="https://zenodo.org/record/3419400/files/figure.png" httpUri-3="https://zenodo.org/record/3419404/files/figure.png" pageId="12" pageNumber="53">
<figureCitation id="94BA2A43EF30813FFBFEFD263AFB14BB" box="[1111,1190,729,755]" captionStart="FIGURE 7" captionStartId="9.[151,250,1816,1841]" captionTargetBox="[151,1435,360,1779]" captionTargetId="figure@9.[151,1435,381,1779]" captionTargetPageId="9" captionText="FIGURE 7. Referred material of Cherevychnavis umanskae sp. nov. (NMNHU-P, no.45-2452, proximal left humerus) in comparison to extant Charadrii species. Scale bars: 10mm." httpUri="https://zenodo.org/record/3258462/files/figure.png" pageId="12" pageNumber="53">Figs. 7</figureCitation>
10
</figureCitation>
), deposited at the National Museum of Natural History at the National Academy of Sciences of
<collectingCountry id="74967656EF30813FFBA6FD023A31155F" box="[1039,1132,765,791]" name="Ukraine" pageId="12" pageNumber="53">Ukraine</collectingCountry>
,
<collectingRegion id="CE45F824EF30813FFBD0FD023AEF155F" box="[1145,1202,765,791]" country="Ukraine" name="Kyiv" pageId="12" pageNumber="53">Kyiv</collectingRegion>
,
<collectingCountry id="74967656EF30813FFB17FD023B41155F" box="[1214,1308,765,791]" name="Ukraine" pageId="12" pageNumber="53">Ukraine</collectingCountry>
. It is likely that the humerus represents the same extinct taxon as the
<typeStatus id="D33A8864EF30813FFC8CFCDE3DDA1573" box="[805,903,801,827]" pageId="12" pageNumber="53" type="holotype">holotype</typeStatus>
because the two bones were found at the same site, they match in size, and can be assigned to the same suborder
<taxonomicName id="CB814D45EF30813FFC28FCBA3B511517" authority="sensu Paton et al. (2003)" authorityName="Paton" authorityYear="2003" box="[897,1292,837,863]" class="Aves" higherTaxonomySource="GBIF" kingdom="Animalia" order="Charadriiformes" pageId="12" pageNumber="53" phylum="Chordata" rank="subOrder" subOrder="Charadrii">
Charadrii sensu
<bibRefCitation id="68104B37EF30813FFB91FCBA3B511517" author="Paton, T. A. &amp; Baker, A. J. &amp; Groth, J. G. &amp; Barrowclough, G. F." box="[1080,1292,837,863]" pageId="12" pageNumber="53" pagination="268 - 278" refId="ref7684" refString="Paton, T. A., Baker, A. J., Groth, J. G. &amp; Barrowclough, G. F. (2003) RAG- 1 sequences resolve phylogenetic relationships within Charadriiform birds. Molecular phylogenetics and evolution, 29 (2), 268 - 278. https: // doi. org / 10.1016 / S 1055 - 7903 (03) 00098 - 8" type="journal article" year="2003">
Paton
<emphasis id="3EF5EAD4EF30813FFBD6FCB93AE51517" box="[1151,1208,837,863]" italics="true" pageId="12" pageNumber="53">et al.</emphasis>
(2003)
</bibRefCitation>
</taxonomicName>
, but because they were not found in articulation, it cannot be conclusively demonstrated that they belong to the same species.
</paragraph>
<paragraph id="0C3E36C6EF30813FFF6EFC723FE4125B" blockId="12.[151,1437,549,1223]" pageId="12" pageNumber="53">
<emphasis id="3EF5EAD4EF30813FFF6EFC723C5415EF" bold="true" box="[199,521,908,935]" pageId="12" pageNumber="53">Type locality and horizon.</emphasis>
Cherevychne 3, located on the left banks of Khadzhibey Estuary, between the villages of Cherevychne and Morozovka (Bilyayivka District,
<collectingRegion id="CE45F824EF30813FFCC3FC4E3A441583" box="[874,1049,945,971]" country="Ukraine" name="Odesa" pageId="12" pageNumber="53">Odessa Region</collectingRegion>
,
<collectingCountry id="74967656EF30813FFB8EFC4E3AD81583" box="[1063,1157,945,971]" name="Ukraine" pageId="12" pageNumber="53">Ukraine</collectingCountry>
;
<geoCoordinate id="69B55001EF30813FFB3AFC4E3B671583" box="[1171,1338,944,971]" degrees="46" direction="north" minutes="38" orientation="latitude" pageId="12" pageNumber="53" precision="15" seconds="35" value="46.64306">46° 38 35″ N</geoCoordinate>
,
<geoCoordinate id="69B55001EF30813FFAE0FC4E3E8315A7" degrees="30" direction="east" minutes="37" orientation="longitude" pageId="12" pageNumber="53" precision="15" seconds="12" value="30.62">30° 37 12″ E</geoCoordinate>
); late Miocene, lower part of the Mammal Neogene Zone 12 (
<collectionCode id="6A90AE03EF30813FFC17FC2A3DB015A7" box="[958,1005,981,1007]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="12" pageNumber="53">MN</collectionCode>
12) of the late Miocene (Tortonian), approximately 8-7.5 mya.
</paragraph>
</subSubSection>
<subSubSection id="449B654DEF30813FFF6EFBE33C5A127F" box="[199,519,1052,1079]" pageId="12" pageNumber="53" type="diagnosis">
<paragraph id="0C3E36C6EF30813FFF6EFBE33C5A127F" blockId="12.[151,1437,549,1223]" box="[199,519,1052,1079]" pageId="12" pageNumber="53">
<emphasis id="3EF5EAD4EF30813FFF6EFBE33F1F127F" bold="true" box="[199,322,1052,1079]" pageId="12" pageNumber="53">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="449B654DEF30813FFF6EFBBE3F9E128F" pageId="12" pageNumber="53" type="description">
<paragraph id="0C3E36C6EF30813FFF6EFBBE3F9E128F" blockId="12.[151,1437,549,1223]" pageId="12" pageNumber="53">
<emphasis id="3EF5EAD4EF30813FFF6EFBBE3F231213" bold="true" box="[199,382,1089,1115]" pageId="12" pageNumber="53">Measurements.</emphasis>
Coracoid: total length, measured between omal tip of processus acrocoracoideus and angulus lateralis:
<quantity id="CB799B23EF30813FFF56FB9A3F3A1237" box="[255,359,1125,1151]" metricMagnitude="-2" metricUnit="m" metricValue="3.38" pageId="12" pageNumber="53" unit="mm" value="33.8">33.8 mm</quantity>
; width of the sternal facet, measured between angulus medialis and angulus lateralis:
<quantity id="CB799B23EF30813FFA98FB9A3BC41237" box="[1329,1433,1125,1151]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="12" pageNumber="53" unit="mm" value="12.6">12.6 mm</quantity>
. Humerus: length as preserved:
<quantity id="CB799B23EF30813FFE54FB773C3A12EB" box="[509,615,1160,1187]" metricMagnitude="-2" metricUnit="m" metricValue="5.64" pageId="12" pageNumber="53" unit="mm" value="56.4">56.4 mm</quantity>
; proximal width, measured from tuberculum dorsale to outer margin of crista bicipitalis:
<quantity id="CB799B23EF30813FFEF1FB533FE2128F" box="[344,447,1196,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="12" pageNumber="53" unit="mm" value="17.5">17.5 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>