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<document id="A42AE32BDF4A92344A62F3F8D3C89DAE" ID-DOI="10.11646/zootaxa.5506.3.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13747177" ID-ZooBank="0B48C57C-402F-4231-9B8D-55C5F85FB0B0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726054442374" checkinUser="plazi" docAuthor="Lepeco, Anderson" docDate="2024" docId="03B57538577DFFFACBE4FF66C381CCDE" docLanguage="en" docName="zootaxa.5506.3.4.pdf" docOrigin="Zootaxa 5506 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.3.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxytrigona Cockerell 1917" docType="key" docVersion="2" lastPageNumber="392" masterDocId="FF8C0D405768FFEDCB73FFF1C61DCE5C" masterDocTitle="Contribution to the taxonomy of Oxytrigona Cockerell, 1917 (Apidae, Meliponini)" masterLastPageNumber="395" masterPageNumber="369" pageNumber="390" updateTime="1726055285633" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="77A83E437B78221A38470F716053A44F">Contribution to the taxonomy of Oxytrigona Cockerell, 1917 (Apidae, Meliponini)</mods:title>
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<mods:namePart id="78408CF597E40BA84A4177327A0A5AD2">Lepeco, Anderson</mods:namePart>
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<treatment id="03B57538577DFFFACBE4FF66C381CCDE" LSID="urn:lsid:plazi:treatment:03B57538577DFFFACBE4FF66C381CCDE" httpUri="http://treatment.plazi.org/id/03B57538577DFFFACBE4FF66C381CCDE" lastPageId="23" lastPageNumber="392" pageId="21" pageNumber="390">
<subSubSection id="C30697A5577DFFF8CBE4FF66C58DCEED" box="[151,912,151,177]" pageId="21" pageNumber="390" type="nomenclature">
<paragraph id="8BA3C42E577DFFF8CBE4FF66C58DCEED" blockId="21.[151,912,151,177]" box="[151,912,151,177]" pageId="21" pageNumber="390">
<heading id="D0EB7342577DFFF8CBE4FF66C58DCEED" bold="true" box="[151,912,151,177]" fontSize="11" level="1" pageId="21" pageNumber="390" reason="1">
<emphasis id="B968183C577DFFF8CBE4FF66C58DCEED" bold="true" box="[151,912,151,177]" pageId="21" pageNumber="390">
Key to the species of
<taxonomicName id="4C1CBFAD577DFFF8CAEEFF66C4BACEED" ID-CoL="6BJH" authority="Cockerell" authorityName="Cockerell" authorityYear="1917" box="[413,679,151,177]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B968183C577DFFF8CAEEFF66C43BCEED" bold="true" box="[413,550,151,177]" italics="true" pageId="21" pageNumber="390">Oxytrigona</emphasis>
Cockerell
</taxonomicName>
, based on workers
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30697A5577DFFFACBE4FF2DC381CCDE" lastPageId="23" lastPageNumber="392" pageId="21" pageNumber="390" type="key">
<keyStep id="30E8DC8B577DFFF8CBE4FF2DC381CF15" pageId="21" pageNumber="390">
<paragraph id="8BA3C42E577DFFF8CBE4FF2DC381CF52" blockId="21.[151,1437,220,330]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FF2DC381CF52" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4FF2DC6B7CEAD" bold="true" box="[151,170,220,241]" pageId="21" pageNumber="390">1.</emphasis>
Forewing with distal portion of radial cell and entire marginal cell distinctly darkened; veins and pterostigma dark brown (
<figureCitation id="1327D8AB577DFFF8CB9BFF08C751CF53" box="[232,332,249,271]" captionStart="FIGURE 8" captionStartId="22.[151,250,1452,1476]" captionTargetBox="[151,1435,180,1427]" captionTargetId="figure-25@22.[151,1435,180,1427]" captionTargetPageId="22" captionText="FIGURE 8. A) forewing of Oxytrigona obscura (Friese, 1900) from Cuiabá (Mato Grosso, Brazil, RPSP), scale bar: 1 mm; B) scape of Oxytrigona mulfordi (Schwarz, 1948) from Porto Acre (Acre, Brazil, RPSP), scale bar: 0.5 mm; C) anterior view of hind basitarsus of Oxytrigona daemoniaca Camargo, 1984, from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; D) anterior view of hind basitarsus of Oxytrigona ignis Camargo, 1984, from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm; E) scape of O. daemoniaca from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; F) scape of O. ignis from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747201" httpUri="https://zenodo.org/record/13747201/files/figure.png" pageId="21" pageNumber="390">Figure 8A</figureCitation>
).........................................................................................
<emphasis id="B968183C577DFFF8CEE3FF08C381CF52" bold="true" box="[1424,1436,249,270]" pageId="21" pageNumber="390">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577DFFF8CBE4FEE6C381CF15" blockId="21.[151,1437,220,330]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FEE6C381CF15" pageId="21" pageNumber="390">
- Forewing with membrane completely or mostly hyaline, distal portion of radial and marginal cells not strongly darkened,
<emphasis id="B968183C577DFFF8CB91FEC4C75BCF16" box="[226,326,309,330]" italics="true" pageId="21" pageNumber="390">sometimes</emphasis>
yellowish or brownish infumate (as in
<figureCitation id="1327D8AB577DFFF8C9CDFEC5C55ECF15" box="[702,835,308,330]" captionStart="FIGURE 3" captionStartId="6.[151,250,1583,1607]" captionTargetBox="[151,1436,180,1558]" captionTargetId="figure-25@6.[151,1436,180,1558]" captionTargetPageId="6" captionText="FIGURE 3. A) Oxytrigona cagafogo (Müller, 1874), worker from São João de Petrópolis (Espírio Santo, Brazil, RPSP), dorsal view of mesosoma, scale bar: 0.5 mm; B) Oxytrigona flaveola (Friese, 1900), worker from Costa Marques (Rondônia, Brazil, RPSP), anterior view of dorsal portion of the head, scale bar: 0.5 mm; C) O. cagafogo, worker from São Sebastião (São Paulo, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; D) O. flaveola, worker from Costa Marques (Rodônia, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; E) O. cagafogo, worker from Salvador (Bahia, Brazil, RPSP), forewing, scale bar: 1 mm;; F) O. flaveola, worker from Pimenteira (Rodônia, Brazil, RPSP), forewing, scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13747185" httpUri="https://zenodo.org/record/13747185/files/figure.png" pageId="21" pageNumber="390">Figures 3E, F</figureCitation>
); veins and pterostigma yellow to brown...............
<emphasis id="B968183C577DFFF8CEE3FEC5C381CF15" bold="true" box="[1424,1436,308,329]" pageId="21" pageNumber="390">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E8DC8B577DFFF8CBE4FE9EC380CFA6" pageId="21" pageNumber="390">
<paragraph id="8BA3C42E577DFFF8CBE4FE9EC380CFFE" blockId="21.[151,1437,367,506]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FE9EC380CFFE" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4FE9EC6B4CFD8" bold="true" box="[151,169,367,388]" pageId="21" pageNumber="390">2.</emphasis>
Body predominantly yellow to orange; pubescence of head and mesosoma ferruginous; antennal scape with erect black setae (between 0.5 and 0.8 MOD in length) restricted to its distal portion (
<figureCitation id="1327D8AB577DFFF8C801FE7DC5C9CFFD" box="[882,980,396,418]" captionStart="FIGURE 8" captionStartId="22.[151,250,1452,1476]" captionTargetBox="[151,1435,180,1427]" captionTargetId="figure-25@22.[151,1435,180,1427]" captionTargetPageId="22" captionText="FIGURE 8. A) forewing of Oxytrigona obscura (Friese, 1900) from Cuiabá (Mato Grosso, Brazil, RPSP), scale bar: 1 mm; B) scape of Oxytrigona mulfordi (Schwarz, 1948) from Porto Acre (Acre, Brazil, RPSP), scale bar: 0.5 mm; C) anterior view of hind basitarsus of Oxytrigona daemoniaca Camargo, 1984, from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; D) anterior view of hind basitarsus of Oxytrigona ignis Camargo, 1984, from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm; E) scape of O. daemoniaca from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; F) scape of O. ignis from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747201" httpUri="https://zenodo.org/record/13747201/files/figure.png" pageId="21" pageNumber="390">Figure 8B</figureCitation>
)....
<taxonomicName id="4C1CBFAD577DFFF8CF55FE7DC380CFFE" ID-CoL="4BLV6" authority="(Schwarz, 1948)" baseAuthorityName="Schwarz" baseAuthorityYear="1948" box="[1062,1437,396,418]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="species" species="mulfordi">
<emphasis id="B968183C577DFFF8CF55FE7DC380CFFE" bold="true" box="[1062,1437,396,418]" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CF55FE7DC2EECFFE" bold="true" box="[1062,1267,396,418]" italics="true" pageId="21" pageNumber="390">Oxytrigona mulfordi</emphasis>
(
<bibRefCitation id="EF8DB9DF577DFFF8CF8CFE7DC388CFFE" author="Schwarz, H. F." box="[1279,1429,396,418]" pageId="21" pageNumber="390" pagination="1 - 546" refId="ref27900" refString="Schwarz, H. F. (1948) Stingless bees (Meliponidae) of the Western Hemisphere. Bulletin of American Museum of Natural History, 90, 1 - 546." type="journal article" year="1948">Schwarz, 1948</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577DFFF8CBE4FE5BC380CFA6" blockId="21.[151,1437,367,506]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FE5BC380CFA6" pageId="21" pageNumber="390">
- Body predominantly dark brown to black, only lower half of head pale; pubescence of head and mesosoma black; antennal scape
<emphasis id="B968183C577DFFF8CA6EFE36C77FCF80" box="[285,354,455,476]" italics="true" pageId="21" pageNumber="390">usually</emphasis>
with erect black setae (between 0.8 and 1.5 MOD in length) scattered throughout...............................................................................................
<taxonomicName id="4C1CBFAD577DFFF8CF34FE15C380CFA6" ID-CoL="4BLV7" authority="(Friese, 1900)" baseAuthorityName="Friese" baseAuthorityYear="1900" box="[1095,1437,484,506]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B968183C577DFFF8CF34FE15C380CFA6" bold="true" box="[1095,1437,484,506]" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CF34FE15C311CFA6" bold="true" box="[1095,1292,484,506]" italics="true" pageId="21" pageNumber="390">Oxytrigona obscura</emphasis>
(Friese, 1900)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E8DC8B577DFFF8CBE4FDEEC381CCF5" pageId="21" pageNumber="390">
<paragraph id="8BA3C42E577DFFF8CBE4FDEEC381CC0D" blockId="21.[151,1437,543,682]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FDEEC381CC0D" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4FDEEC6B4CC68" bold="true" box="[151,169,543,564]" pageId="21" pageNumber="390">3.</emphasis>
Head mostly dark brown to black,
<emphasis id="B968183C577DFFF8C954FDEEC44ACC68" box="[551,599,543,564]" italics="true" pageId="21" pageNumber="390">often</emphasis>
becoming lighter ventral to antennal socket level; mesoscutum entirely black; scutellum dark brown to black; pubescence mostly black..............................................................
<emphasis id="B968183C577DFFF8CEE3FDCDC381CC0D" bold="true" box="[1424,1436,572,593]" pageId="21" pageNumber="390">4</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577DFFF8CBE4FDABC381CCF5" blockId="21.[151,1437,543,682]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FDABC381CCF5" pageId="21" pageNumber="390">
- Head mostly orange or yellow,
<emphasis id="B968183C577DFFF8C979FDABC452CC33" box="[522,591,602,623]" italics="true" pageId="21" pageNumber="390">at most</emphasis>
with dark spot near ocelli; mesoscutum
<emphasis id="B968183C577DFFF8C8BEFDA8C20FCC32" box="[973,1042,601,622]" italics="true" pageId="21" pageNumber="390">usually</emphasis>
yellowish to brown, at least medially and on surface adjacent to tegula (except for
<taxonomicName id="4C1CBFAD577DFFF8C917FD86C4C4CCD0" baseAuthorityName="Muller" baseAuthorityYear="1874" box="[612,729,631,652]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="species" species="cagafogo">
<emphasis id="B968183C577DFFF8C917FD86C4C4CCD0" box="[612,729,631,652]" italics="true" pageId="21" pageNumber="390">O. cagafogo</emphasis>
</taxonomicName>
, in which it
<emphasis id="B968183C577DFFF8C83CFD86C5AECCD0" box="[847,947,631,652]" italics="true" pageId="21" pageNumber="390">sometimes</emphasis>
is entirely black, but the head is orange); scutellum yellow to brown; color of pubescence
<emphasis id="B968183C577DFFF8C921FD65C4BFCCF5" box="[594,674,660,681]" italics="true" pageId="21" pageNumber="390">variable</emphasis>
.............................................................
<emphasis id="B968183C577DFFF8CEE3FD65C381CCF5" bold="true" box="[1424,1436,660,681]" pageId="21" pageNumber="390">7</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E8DC8B577DFFF8CBE4FD3EC381CDED" pageId="21" pageNumber="390">
<paragraph id="8BA3C42E577DFFF8CBE4FD3EC381CD60" blockId="21.[151,1437,719,946]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FD3EC381CD60" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4FD3EC6B4CCB8" bold="true" box="[151,169,719,740]" pageId="21" pageNumber="390">4.</emphasis>
Mesoscutum with long erect setae becoming virtually absent posterior to mid-level of tegula; scutellum with erect setae becoming distinctly shorter or absent on anterior half; scape mostly without erect black setae, only with very short erect setae near distal end; distal angle of dorsal margin of hind basitarsus sharply angled (
<figureCitation id="1327D8AB577DFFF8C89BFCF8C24DCD43" box="[1000,1104,777,799]" captionStart="FIGURE 6" captionStartId="18.[151,250,1298,1322]" captionTargetBox="[151,1436,181,1274]" captionTargetId="figure-25@18.[151,1436,181,1274]" captionTargetPageId="18" captionText="FIGURE 6. Holotype worker of Oxytrigona sauronica sp. nov. (RPSP 09.0361) from Tamara (Casanare, Colombia).A) habitus, lateral view, scale bar: 2 mm; B) head and mesosoma, dorsal view, scale bar: 0.5 mm; C) head, frontal view, scale bar: 0.5 mm; D) hind leg, scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747195" httpUri="https://zenodo.org/record/13747195/files/figure.png" pageId="21" pageNumber="390">Figure 6D</figureCitation>
); distal corner of dorsal margin of hind tibia with dense tuft of densely branched brown setae (
<figureCitation id="1327D8AB577DFFF8C86CFCD6C59ECD61" box="[799,899,807,829]" captionStart="FIGURE 6" captionStartId="18.[151,250,1298,1322]" captionTargetBox="[151,1436,181,1274]" captionTargetId="figure-25@18.[151,1436,181,1274]" captionTargetPageId="18" captionText="FIGURE 6. Holotype worker of Oxytrigona sauronica sp. nov. (RPSP 09.0361) from Tamara (Casanare, Colombia).A) habitus, lateral view, scale bar: 2 mm; B) head and mesosoma, dorsal view, scale bar: 0.5 mm; C) head, frontal view, scale bar: 0.5 mm; D) hind leg, scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747195" httpUri="https://zenodo.org/record/13747195/files/figure.png" pageId="21" pageNumber="390">Figure 6D</figureCitation>
)..................
<emphasis id="B968183C577DFFF8CF02FCD6C381CD60" bold="true" box="[1137,1436,807,828]" pageId="21" pageNumber="390">
<taxonomicName id="4C1CBFAD577DFFF8CF02FCD6C357CD60" authorityName="Lepeco" authorityYear="2024" box="[1137,1354,807,828]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="species" species="sauronica" status="sp. nov.">
<emphasis id="B968183C577DFFF8CF02FCD6C357CD60" bold="true" box="[1137,1354,807,828]" italics="true" pageId="21" pageNumber="390">Oxytrigona sauronica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BA547577DFFF8CE23FCD6C381CD60" box="[1360,1436,807,828]" pageId="21" pageNumber="390" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577DFFF8CBE4FCB5C381CDED" blockId="21.[151,1437,719,946]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FCB5C381CDED" pageId="21" pageNumber="390">
- Mesoscutum with long erect setae evenly distributed throughout sclerite; scutellum with erect setae evenly long, distributed throughout sclerite; scape with black erect setae with at least 0.5 MOD in length,
<emphasis id="B968183C577DFFF8C89EFC90C200CD2A" box="[1005,1053,865,886]" italics="true" pageId="21" pageNumber="390">often</emphasis>
restricted to distal end; distal portion of dorsal margin of hind basitarsus widely angled or rounded; distal corner of dorsal margin of hind tibia with relatively sparse black setae with few apical ramifications..................................................................
<emphasis id="B968183C577DFFF8CEE3FC6DC381CDED" bold="true" box="[1424,1436,924,945]" pageId="21" pageNumber="390">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E8DC8B577DFFF8CBE4FC26C381CAA8" pageId="21" pageNumber="390">
<paragraph id="8BA3C42E577DFFF8CBE4FC26C381CA3D" blockId="21.[151,1437,983,1268]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FC26C381CA3D" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4FC26C6B4CDB0" bold="true" box="[151,169,983,1004]" pageId="21" pageNumber="390">5.</emphasis>
Hind basitarsus conspicuously narrower proximally, distal corner of dorsal margin rounded (as in
<figureCitation id="1327D8AB577DFFF8CFF3FC26C2FFCDB0" box="[1152,1250,983,1005]" captionStart="FIGURE 8" captionStartId="22.[151,250,1452,1476]" captionTargetBox="[151,1435,180,1427]" captionTargetId="figure-25@22.[151,1435,180,1427]" captionTargetPageId="22" captionText="FIGURE 8. A) forewing of Oxytrigona obscura (Friese, 1900) from Cuiabá (Mato Grosso, Brazil, RPSP), scale bar: 1 mm; B) scape of Oxytrigona mulfordi (Schwarz, 1948) from Porto Acre (Acre, Brazil, RPSP), scale bar: 0.5 mm; C) anterior view of hind basitarsus of Oxytrigona daemoniaca Camargo, 1984, from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; D) anterior view of hind basitarsus of Oxytrigona ignis Camargo, 1984, from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm; E) scape of O. daemoniaca from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; F) scape of O. ignis from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747201" httpUri="https://zenodo.org/record/13747201/files/figure.png" pageId="21" pageNumber="390">Figure 8C</figureCitation>
); smaller size, head width below
<quantity id="4CE469CB577DFFF8CA2CFC05C7B1CA56" box="[351,428,1012,1034]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="390" unit="mm" value="2.3">2.3 mm</quantity>
; scape with erect black setae (0.51.2 × F2 width) scattered throughout (as in
<figureCitation id="1327D8AB577DFFF8CFE8FC05C2E3CA55" box="[1179,1278,1012,1034]" captionStart="FIGURE 8" captionStartId="22.[151,250,1452,1476]" captionTargetBox="[151,1435,180,1427]" captionTargetId="figure-25@22.[151,1435,180,1427]" captionTargetPageId="22" captionText="FIGURE 8. A) forewing of Oxytrigona obscura (Friese, 1900) from Cuiabá (Mato Grosso, Brazil, RPSP), scale bar: 1 mm; B) scape of Oxytrigona mulfordi (Schwarz, 1948) from Porto Acre (Acre, Brazil, RPSP), scale bar: 0.5 mm; C) anterior view of hind basitarsus of Oxytrigona daemoniaca Camargo, 1984, from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; D) anterior view of hind basitarsus of Oxytrigona ignis Camargo, 1984, from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm; E) scape of O. daemoniaca from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; F) scape of O. ignis from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747201" httpUri="https://zenodo.org/record/13747201/files/figure.png" pageId="21" pageNumber="390">Figure 8E</figureCitation>
); pubescence on posterior surface of T3T5 relatively sparse; short setae on posterior margin of T3 and T4 scarce, shorter than 0.2 MOD; F2 with distinctly long setae (longer than 0.7 F2 width, these setae can be absent in some specimens); surface between mid and lateral ocelli with short branched setae....................................................................
<emphasis id="B968183C577DFFF8CEE3FBBDC381CA3D" bold="true" box="[1424,1436,1100,1121]" pageId="21" pageNumber="390">6</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577DFFF8CBE4FB9BC381CAA8" blockId="21.[151,1437,983,1268]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FB9BC381CAA8" pageId="21" pageNumber="390">
- Hind basitarsus almost as wide proximally as on its distal portion, distal corner of dorsal margin widely angled (
<figureCitation id="1327D8AB577DFFF8CE5AFB98C38DCA22" box="[1321,1424,1129,1151]" captionStart="FIGURE 8" captionStartId="22.[151,250,1452,1476]" captionTargetBox="[151,1435,180,1427]" captionTargetId="figure-25@22.[151,1435,180,1427]" captionTargetPageId="22" captionText="FIGURE 8. A) forewing of Oxytrigona obscura (Friese, 1900) from Cuiabá (Mato Grosso, Brazil, RPSP), scale bar: 1 mm; B) scape of Oxytrigona mulfordi (Schwarz, 1948) from Porto Acre (Acre, Brazil, RPSP), scale bar: 0.5 mm; C) anterior view of hind basitarsus of Oxytrigona daemoniaca Camargo, 1984, from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; D) anterior view of hind basitarsus of Oxytrigona ignis Camargo, 1984, from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm; E) scape of O. daemoniaca from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; F) scape of O. ignis from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747201" httpUri="https://zenodo.org/record/13747201/files/figure.png" pageId="21" pageNumber="390">Figure 8D</figureCitation>
); larger size, head as wide as
<quantity id="4CE469CB577DFFF8CA9BFB76C42DCAC0" box="[488,560,1159,1181]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="390" unit="mm" value="2.5">2.5 mm</quantity>
or more; scape with erect black setae (0.51.2 × F2 width) restricted to its distal end (
<figureCitation id="1327D8AB577DFFF8CE2FFB76C6E3CAE5" captionStart="FIGURE 8" captionStartId="22.[151,250,1452,1476]" captionTargetBox="[151,1435,180,1427]" captionTargetId="figure-25@22.[151,1435,180,1427]" captionTargetPageId="22" captionText="FIGURE 8. A) forewing of Oxytrigona obscura (Friese, 1900) from Cuiabá (Mato Grosso, Brazil, RPSP), scale bar: 1 mm; B) scape of Oxytrigona mulfordi (Schwarz, 1948) from Porto Acre (Acre, Brazil, RPSP), scale bar: 0.5 mm; C) anterior view of hind basitarsus of Oxytrigona daemoniaca Camargo, 1984, from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; D) anterior view of hind basitarsus of Oxytrigona ignis Camargo, 1984, from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm; E) scape of O. daemoniaca from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; F) scape of O. ignis from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747201" httpUri="https://zenodo.org/record/13747201/files/figure.png" pageId="21" pageNumber="390">Figure 8F</figureCitation>
); T3T5 with dense pubescence on posterior surface; short setae on posterior margin of T3 and T4 relatively abundant, longer than 0.2 MOD; F2
<emphasis id="B968183C577DFFF8CAAEFB33C40FCA8B" box="[477,530,1218,1239]" italics="true" pageId="21" pageNumber="390">never</emphasis>
with distinctly long setae; surface between mid and lateral ocelli with simple setae................................................................................
<taxonomicName id="4C1CBFAD577DFFF8CF27FB2EC381CAA8" ID-CoL="4BLV2" authority="Camargo, 1984" authorityName="Camargo" authorityYear="1984" box="[1108,1436,1246,1268]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="species" species="ignis">
<emphasis id="B968183C577DFFF8CF27FB2EC381CAA8" bold="true" box="[1108,1436,1246,1268]" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CF27FB2EC2E7CAA8" bold="true" box="[1108,1274,1247,1268]" italics="true" pageId="21" pageNumber="390">Oxytrigona ignis</emphasis>
<bibRefCitation id="EF8DB9DF577DFFF8CE73FB2FC381CAA8" author="Camargo, J. M. F." box="[1280,1436,1246,1268]" pageId="21" pageNumber="390" pagination="115 - 124" refId="ref26579" refString="Camargo, J. M. F. (1984) Notas sobre o genero Oxytrigona (Meliponinae, Apidae, Hymenoptera). Boletim do Museu Paraense Emilio Goeldi Zoologia, 1 (1), 115 - 124." type="journal article" year="1984">Camargo, 1984</bibRefCitation>
</emphasis>
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<paragraph id="8BA3C42E577DFFF8CBE4FAE8C381CB10" blockId="21.[151,1436,1305,1415]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FAE8C381CB10" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4FAE8C6B4CB72" bold="true" box="[151,169,1305,1326]" pageId="21" pageNumber="390">6.</emphasis>
Metasomal terga yellow to light ferruginous; scape light to dark brown; ventral half of face with orange hue................................................................................
<taxonomicName id="4C1CBFAD577DFFF8CF7FFAC6C381CB10" ID-CoL="6TH5D" authority="Camargo, 1984" authorityName="Camargo" authorityYear="1984" box="[1036,1436,1334,1356]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="species" species="daemoniaca">
<emphasis id="B968183C577DFFF8CF7FFAC6C381CB10" bold="true" box="[1036,1436,1334,1356]" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CF7FFAC6C2E7CB10" bold="true" box="[1036,1274,1334,1356]" italics="true" pageId="21" pageNumber="390">Oxytrigona daemoniaca</emphasis>
<bibRefCitation id="EF8DB9DF577DFFF8CE73FAC7C381CB10" author="Camargo, J. M. F." box="[1280,1436,1334,1356]" pageId="21" pageNumber="390" pagination="115 - 124" refId="ref26579" refString="Camargo, J. M. F. (1984) Notas sobre o genero Oxytrigona (Meliponinae, Apidae, Hymenoptera). Boletim do Museu Paraense Emilio Goeldi Zoologia, 1 (1), 115 - 124." type="journal article" year="1984">Camargo, 1984</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BA3C42E577DFFF8CBE4FAA5C381CBDA" blockId="21.[151,1436,1305,1415]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FAA5C381CBDA" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4FAA5C682CB35" bold="true" box="[151,159,1364,1385]" pageId="21" pageNumber="390">-</emphasis>
Metasomal terga dark brown; scape
<emphasis id="B968183C577DFFF8C94DFAA5C49ECB35" box="[574,643,1364,1385]" italics="true" pageId="21" pageNumber="390">usually</emphasis>
dark brown to black; ventral half of face with yellowish to light brown hue...............................................................
<taxonomicName id="4C1CBFAD577DFFF8C8D4FA80C381CBDA" ID-CoL="6TGTD" authority="Gonzalez and Roubik, 2008" authorityName="Gonzalez and Roubik" authorityYear="2008" box="[935,1436,1393,1415]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="species" species="chocoana">
<emphasis id="B968183C577DFFF8C8D4FA80C381CBDA" bold="true" box="[935,1436,1393,1415]" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8C8D4FA80C260CBDB" bold="true" box="[935,1149,1393,1415]" italics="true" pageId="21" pageNumber="390">Oxytrigona chocoana</emphasis>
<bibRefCitation id="EF8DB9DF577DFFF8CFF0FA80C381CBDA" author="Gonzalez, V. H. &amp; Roubik, D. W." box="[1155,1436,1393,1415]" pageId="21" pageNumber="390" pagination="43 - 71" refId="ref27261" refString="Gonzalez, V. H. &amp; Roubik, D. W. (2008) Especies nuevas y filogenia de las abejas de fuego, Oxytrigona (Hymenoptera: Apidae, Meliponini). Acta Zoologica Mexicana, 24 (1), 43 - 71. https: // doi. org / 10.21829 / azm. 2008.241615" type="journal article" year="2008">Gonzalez and Roubik, 2008</bibRefCitation>
</emphasis>
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<paragraph id="8BA3C42E577DFFF8CBE4FA5DC381CB82" blockId="21.[151,1437,1452,1562]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FA5DC381CB82" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4FA5DC6B4CB9D" bold="true" box="[151,169,1452,1473]" pageId="21" pageNumber="390">7.</emphasis>
Surface between ocellar triangle and antennal sockets with dense coverage of short setae, punctation relatively dense, especially near ocellar triangle level (as in
<figureCitation id="1327D8AB577DFFF8C964FA38C49ECB83" box="[535,643,1481,1503]" captionStart="FIGURE 4" captionStartId="14.[151,250,1198,1222]" captionTargetBox="[151,1435,180,1173]" captionTargetId="figure-25@14.[151,1435,180,1173]" captionTargetPageId="14" captionText="FIGURE 4. Holotype worker of Oxytrigona mellicolor (Packard, 1889) from Ecuador (MCZ 559). A) habitus, lateral view; B) head, frontal view; C) habitus, dorsal view; D) forewing; E) lateral view of mesosoma. Credits of the images: Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College (http://creativecommons.org/licences/ by-nc-sa/4.0/legalcode). No editions made over the original images." figureDoi="http://doi.org/10.5281/zenodo.13747189" httpUri="https://zenodo.org/record/13747189/files/figure.png" pageId="21" pageNumber="390">Figures 4B</figureCitation>
,
<figureCitation id="1327D8AB577DFFF8C9FEFA38C4B7CB83" box="[653,682,1481,1503]" captionStart="FIGURE 5" captionStartId="15.[151,250,1685,1709]" captionTargetBox="[151,1436,200,1660]" captionTargetId="figure-15@15.[151,1436,200,1660]" captionTargetPageId="15" captionText="FIGURE 5. Oxytrigona pallida sp. nov. A) worker holotype (RPSP 11295) from San José (Costa Rica), lateral view, scale bar: 2 mm; B) worker holotype, frontal view of dorsal portion of head, scale bar: 0.5 mm; C) worker holotype, dorsal view of mesosoma, scale bar: 1 mm; D) worker holotype, lateral view of legs and mesosoma, scale bar: 1 mm; E) male from Quepos (Costa Rica, RPSP), frontal view of head, scale bar: 1 mm; F) male from Quepos (Costa Rica, RPSP), lateral view, scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13747191" httpUri="https://zenodo.org/record/13747191/files/figure.png" pageId="21" pageNumber="390">5B</figureCitation>
)............................................................
<emphasis id="B968183C577DFFF8CEE3FA38C381CB82" bold="true" box="[1424,1436,1481,1502]" pageId="21" pageNumber="390">8</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577DFFF8CBE4FA16C380C845" blockId="21.[151,1437,1452,1562]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4FA16C380C845" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4FA16C682CBA0" bold="true" box="[151,159,1511,1532]" pageId="21" pageNumber="390">-</emphasis>
Surface between ocellar triangle and antennal sockets glabrous or with scattered tiny setae, punctation sparse (as in
<figureCitation id="1327D8AB577DFFF8CE2DFA16C6E3C845" captionStart="FIGURE 3" captionStartId="6.[151,250,1583,1607]" captionTargetBox="[151,1436,180,1558]" captionTargetId="figure-25@6.[151,1436,180,1558]" captionTargetPageId="6" captionText="FIGURE 3. A) Oxytrigona cagafogo (Müller, 1874), worker from São João de Petrópolis (Espírio Santo, Brazil, RPSP), dorsal view of mesosoma, scale bar: 0.5 mm; B) Oxytrigona flaveola (Friese, 1900), worker from Costa Marques (Rondônia, Brazil, RPSP), anterior view of dorsal portion of the head, scale bar: 0.5 mm; C) O. cagafogo, worker from São Sebastião (São Paulo, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; D) O. flaveola, worker from Costa Marques (Rodônia, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; E) O. cagafogo, worker from Salvador (Bahia, Brazil, RPSP), forewing, scale bar: 1 mm;; F) O. flaveola, worker from Pimenteira (Rodônia, Brazil, RPSP), forewing, scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13747185" httpUri="https://zenodo.org/record/13747185/files/figure.png" pageId="21" pageNumber="390">Figure 3B</figureCitation>
)...............................................................................................
<emphasis id="B968183C577DFFF8CEF5F9F5C380C845" bold="true" box="[1414,1437,1540,1561]" pageId="21" pageNumber="390">11</emphasis>
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<paragraph id="8BA3C42E577DFFF8CBE4F9CEC380C8F0" blockId="21.[151,1437,1599,1826]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4F9CEC380C8F0" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4F9CEC6B4C808" bold="true" box="[151,169,1599,1620]" pageId="21" pageNumber="390">8.</emphasis>
Frons with evenly dense punctation between compound eye and frontal line, interspaces about as long as puncture diameter; long setae on frons absent ventrad ocellar triangle level; antennal sockets separated by nearly one socket diameter (
<figureCitation id="1327D8AB577DFFF8CE5FF9ADC38DC82D" box="[1324,1424,1628,1650]" captionStart="FIGURE 4" captionStartId="14.[151,250,1198,1222]" captionTargetBox="[151,1435,180,1173]" captionTargetId="figure-25@14.[151,1435,180,1173]" captionTargetPageId="14" captionText="FIGURE 4. Holotype worker of Oxytrigona mellicolor (Packard, 1889) from Ecuador (MCZ 559). A) habitus, lateral view; B) head, frontal view; C) habitus, dorsal view; D) forewing; E) lateral view of mesosoma. Credits of the images: Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College (http://creativecommons.org/licences/ by-nc-sa/4.0/legalcode). No editions made over the original images." figureDoi="http://doi.org/10.5281/zenodo.13747189" httpUri="https://zenodo.org/record/13747189/files/figure.png" pageId="21" pageNumber="390">Figure 4B</figureCitation>
); postero-dorsal surface of propodeum striate; ventral margin of fore femur carinate; mid basitarsus relatively short, as long as 2 × its maximum width..................................................
<taxonomicName id="4C1CBFAD577DFFF8CF69F966C380C8F0" ID-CoL="4BLV5" authority="(Packard, 1869)" baseAuthorityName="Packard" baseAuthorityYear="1869" box="[1050,1437,1686,1708]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="species" species="mellicolor">
<emphasis id="B968183C577DFFF8CF69F966C380C8F0" bold="true" box="[1050,1437,1686,1708]" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CF69F966C2E9C8F0" bold="true" box="[1050,1268,1686,1708]" italics="true" pageId="21" pageNumber="390">Oxytrigona mellicolor</emphasis>
(
<bibRefCitation id="EF8DB9DF577DFFF8CE73F967C388C8F0" author="Packard, A. S. Jr." box="[1280,1429,1686,1708]" pageId="21" pageNumber="390" pagination="56 - 59" refId="ref27689" refString="Packard, A. S. Jr. (1869) List of hymenopterous and lepidopterous insects collected by the Smithsonian expedition to South America, under Prof. James Orton. Annual Report of the Trustees of the Peabody Academy of Science, 1, 56 - 59." type="journal article" year="1869">Packard, 1869</bibRefCitation>
)
</emphasis>
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</paragraph>
<paragraph id="8BA3C42E577DFFF8CBE4F945C381C97D" blockId="21.[151,1437,1599,1826]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4F945C381C97D" pageId="21" pageNumber="390">
- Frons with denser punctation near ocellar triangle level, becoming sparser near antennal sockets, interspaces far longer than puncture diameter; long setae
<emphasis id="B968183C577DFFF8C973F920C458C8BA" box="[512,581,1745,1766]" italics="true" pageId="21" pageNumber="390">usually</emphasis>
present ventrad ocellar triangle level; antennal sockets separated by 0.8 × socket diameter or less; postero-dorsal surface of propodeum smooth; ventral margin of fore femur not carinate, rounded; mid basitarsus relatively long, longer than 2.3 × its maximum width........................................................
<emphasis id="B968183C577DFFF8CEE3F8FDC381C97D" bold="true" box="[1424,1436,1804,1825]" pageId="21" pageNumber="390">9</emphasis>
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<paragraph id="8BA3C42E577DFFF8CBE4F8BCC381C9C1" blockId="21.[151,1436,1869,2009]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4F8BCC381C9C1" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8CBE4F8BCC6B4C93E" bold="true" box="[151,169,1869,1890]" pageId="21" pageNumber="390">9.</emphasis>
Fore femur less than 3.7 times longer than its maximum width; smaller size, head width equal
<quantity id="4CE469CB577DFFF8CF0AF8BFC2DEC93F" box="[1145,1219,1870,1891]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="390" unit="mm" value="2.4">2.4 mm</quantity>
or less; short setae on posterior margin of T3 relatively longer (0.5 × MOD) and densely branched...................................................................................................
<taxonomicName id="4C1CBFAD577DFFF8C8B0F879C381C9C1" ID-CoL="4BLV3" authority="Gonzalez &amp; Roubik, 2008" authorityName="Gonzalez &amp; Roubik" authorityYear="2008" box="[963,1436,1928,1950]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="390" phylum="Arthropoda" rank="species" species="isthmina">
<emphasis id="B968183C577DFFF8C8B0F879C381C9C1" bold="true" box="[963,1436,1928,1950]" pageId="21" pageNumber="390">
<emphasis id="B968183C577DFFF8C8B0F879C28DC9C2" bold="true" box="[963,1168,1928,1950]" italics="true" pageId="21" pageNumber="390">Oxytrigona isthmina</emphasis>
<bibRefCitation id="EF8DB9DF577DFFF8CFE5F879C381C9C1" author="Gonzalez, V. H. &amp; Roubik, D. W." box="[1174,1436,1928,1950]" pageId="21" pageNumber="390" pagination="43 - 71" refId="ref27261" refString="Gonzalez, V. H. &amp; Roubik, D. W. (2008) Especies nuevas y filogenia de las abejas de fuego, Oxytrigona (Hymenoptera: Apidae, Meliponini). Acta Zoologica Mexicana, 24 (1), 43 - 71. https: // doi. org / 10.21829 / azm. 2008.241615" type="journal article" year="2008">Gonzalez &amp; Roubik, 2008</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BA3C42E577DFFF8CBE4F857C381C984" blockId="21.[151,1436,1869,2009]" pageId="21" pageNumber="390">
<keyLead id="30ED671B577DFFF8CBE4F857C381C984" pageId="21" pageNumber="390">
- Fore femur longer than 4 × its maximum width; larger size, head width over
<quantity id="4CE469CB577DFFF8C8A4F854C238C9E7" box="[983,1061,1957,1979]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="390" unit="mm" value="2.5">2.5 mm</quantity>
; short setae on posterior margin of T3 shorter than 0.4 × MOD and mostly simple...............................................................
<emphasis id="B968183C577DFFF8CEF7F832C381C984" bold="true" box="[1412,1436,1987,2008]" pageId="21" pageNumber="390">10</emphasis>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF6394A6577EFFFBCBE4FA5DC74EC824" ID-DOI="http://doi.org/10.5281/zenodo.13747201" ID-Zenodo-Dep="13747201" httpUri="https://zenodo.org/record/13747201/files/figure.png" pageId="22" pageNumber="391" startId="22.[151,250,1452,1476]" targetBox="[151,1435,180,1427]" targetPageId="22" targetType="figure">
<paragraph id="8BA3C42E577EFFFBCBE4FA5DC74EC824" blockId="22.[151,1436,1452,1656]" pageId="22" pageNumber="391">
<emphasis id="B968183C577EFFFBCBE4FA5DC70ECB98" bold="true" box="[151,275,1452,1476]" pageId="22" pageNumber="391">FIGURE 8.</emphasis>
A) forewing of
<taxonomicName id="4C1CBFAD577EFFFBCAC4FA5CC50BCB98" authority="(Friese, 1900)" baseAuthorityName="Friese" baseAuthorityYear="1900" box="[439,790,1452,1476]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="391" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B968183C577EFFFBCAC4FA5CC49ECB98" box="[439,643,1452,1476]" italics="true" pageId="22" pageNumber="391">Oxytrigona obscura</emphasis>
(Friese, 1900)
</taxonomicName>
from Cuiabá (Mato Grosso, Brazil, RPSP), scale bar: 1 mm; B) scape of
<taxonomicName id="4C1CBFAD577EFFFBCB87FA20C468CBB4" authority="(Schwarz, 1948)" baseAuthorityName="Schwarz" baseAuthorityYear="1948" box="[244,629,1488,1512]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="391" phylum="Arthropoda" rank="species" species="mulfordi">
<emphasis id="B968183C577EFFFBCB87FA20C7D8CBB4" box="[244,453,1488,1512]" italics="true" pageId="22" pageNumber="391">Oxytrigona mulfordi</emphasis>
(
<bibRefCitation id="EF8DB9DF577EFFFBCAA7FA21C473CBB4" author="Schwarz, H. F." box="[468,622,1488,1512]" pageId="22" pageNumber="391" pagination="1 - 546" refId="ref27900" refString="Schwarz, H. F. (1948) Stingless bees (Meliponidae) of the Western Hemisphere. Bulletin of American Museum of Natural History, 90, 1 - 546." type="journal article" year="1948">Schwarz, 1948</bibRefCitation>
)
</taxonomicName>
from Porto Acre (Acre, Brazil, RPSP), scale bar: 0.5 mm; C) anterior view of hind basitarsus of
<taxonomicName id="4C1CBFAD577EFFFBCA22FA04C4F1C850" authority="Camargo, 1984" authorityName="Camargo" authorityYear="1984" box="[337,748,1524,1548]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="391" phylum="Arthropoda" rank="species" species="daemoniaca">
<emphasis id="B968183C577EFFFBCA22FA04C45AC850" box="[337,583,1524,1548]" italics="true" pageId="22" pageNumber="391">Oxytrigona daemoniaca</emphasis>
<bibRefCitation id="EF8DB9DF577EFFFBC93DFA05C4F1C850" author="Camargo, J. M. F." box="[590,748,1524,1548]" pageId="22" pageNumber="391" pagination="115 - 124" refId="ref26579" refString="Camargo, J. M. F. (1984) Notas sobre o genero Oxytrigona (Meliponinae, Apidae, Hymenoptera). Boletim do Museu Paraense Emilio Goeldi Zoologia, 1 (1), 115 - 124." type="journal article" year="1984">Camargo, 1984</bibRefCitation>
</taxonomicName>
, from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; D) anterior view of hind basitarsus of
<taxonomicName id="4C1CBFAD577EFFFBCAD4F9E8C4E4C86C" authority="Camargo, 1984" authorityName="Camargo" authorityYear="1984" box="[423,761,1560,1584]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="391" phylum="Arthropoda" rank="species" species="ignis">
<emphasis id="B968183C577EFFFBCAD4F9E8C44FC86C" box="[423,594,1561,1584]" italics="true" pageId="22" pageNumber="391">Oxytrigona ignis</emphasis>
<bibRefCitation id="EF8DB9DF577EFFFBC929F9E9C4E4C86C" author="Camargo, J. M. F." box="[602,761,1560,1584]" pageId="22" pageNumber="391" pagination="115 - 124" refId="ref26579" refString="Camargo, J. M. F. (1984) Notas sobre o genero Oxytrigona (Meliponinae, Apidae, Hymenoptera). Boletim do Museu Paraense Emilio Goeldi Zoologia, 1 (1), 115 - 124." type="journal article" year="1984">Camargo, 1984</bibRefCitation>
</taxonomicName>
, from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm; E) scape of
<taxonomicName id="4C1CBFAD577EFFFBCBE4F9CCC72EC808" authorityName="Camargo" authorityYear="1984" box="[151,307,1596,1620]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="391" phylum="Arthropoda" rank="species" species="daemoniaca">
<emphasis id="B968183C577EFFFBCBE4F9CCC72EC808" box="[151,307,1596,1620]" italics="true" pageId="22" pageNumber="391">O. daemoniaca</emphasis>
</taxonomicName>
from Puerto Pilon (Panama, RPSP), scale bar: 0.5 mm; F) scape of
<taxonomicName id="4C1CBFAD577EFFFBC898F9CCC221C808" authorityName="Camargo" authorityYear="1984" box="[1003,1084,1597,1620]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="391" phylum="Arthropoda" rank="species" species="ignis">
<emphasis id="B968183C577EFFFBC898F9CCC221C808" box="[1003,1084,1597,1620]" italics="true" pageId="22" pageNumber="391">O. ignis</emphasis>
</taxonomicName>
from Belém (Pará, Brazil, RPSP), scale bar: 0.5 mm.
</paragraph>
</caption>
<keyStep id="30E8DC8B577EFFFBCBE4F96CC381C919" pageId="22" pageNumber="391">
<paragraph id="8BA3C42E577EFFFBCBE4F96CC380C8B1" blockId="22.[151,1437,1693,1861]" pageId="22" pageNumber="391">
<keyLead id="30ED671B577EFFFBCBE4F96CC380C8B1" pageId="22" pageNumber="391">
<emphasis id="B968183C577EFFFBCBE4F96CC6ABC8EE" bold="true" box="[151,182,1693,1714]" pageId="22" pageNumber="391">10.</emphasis>
Longest setae on vertex about 1.7 × MOD; mesoscutum yellow to orange between notauli and dark brown to black laterad; legs brown; posterior margin of S6 widely rounded (see fig.
<quantity id="4CE469CB577EFFFBC986F94BC504C88C" box="[757,793,1722,1744]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="22" pageNumber="391" unit="in" value="9.0">9 in</quantity>
<bibRefCitation id="EF8DB9DF577EFFFBC86DF94BC20AC893" author="Gonzalez, V. H. &amp; Roubik, D. W." box="[798,1047,1722,1744]" pageId="22" pageNumber="391" pagination="43 - 71" refId="ref27261" refString="Gonzalez, V. H. &amp; Roubik, D. W. (2008) Especies nuevas y filogenia de las abejas de fuego, Oxytrigona (Hymenoptera: Apidae, Meliponini). Acta Zoologica Mexicana, 24 (1), 43 - 71. https: // doi. org / 10.21829 / azm. 2008.241615" type="journal article" year="2008">Gonzalez &amp; Roubik 2008</bibRefCitation>
); setae on distal end of scape longer than 0.7 × MOD.........................................................
<taxonomicName id="4C1CBFAD577EFFFBCF78F926C380C8B1" ID-CoL="4BLV4" authority="(Cockerell, 1913)" authorityName="EL" baseAuthorityName="Cockerell" baseAuthorityYear="1913" box="[1035,1437,1751,1773]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="391" phylum="Arthropoda" rank="species" species="mediorufa">
<emphasis id="B968183C577EFFFBCF78F926C380C8B1" bold="true" box="[1035,1437,1751,1773]" pageId="22" pageNumber="391">
<emphasis id="B968183C577EFFFBCF78F926C2F5C8B1" bold="true" box="[1035,1256,1751,1773]" italics="true" pageId="22" pageNumber="391">Oxytrigona mediorufa</emphasis>
(Cockerell, 1913)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577EFFFBCBE4F904C381C919" blockId="22.[151,1437,1693,1861]" pageId="22" pageNumber="391">
<keyLead id="30ED671B577EFFFBCBE4F904C381C919" pageId="22" pageNumber="391">
- Longest setae on vertex about 1.3 × MOD or shorter; mesoscutum entirely light brown or caramel yellow; legs yellow; posterior margin of S6 truncate (see fig.
<quantity id="4CE469CB577EFFFBC978F8E3C421C974" box="[523,572,1810,1832]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="22" pageNumber="391" unit="in" value="21.0">21 in</quantity>
<bibRefCitation id="EF8DB9DF577EFFFBC931F8E3C521C97B" author="Gonzalez, V. H. &amp; Roubik, D. W." box="[578,828,1810,1832]" pageId="22" pageNumber="391" pagination="43 - 71" refId="ref27261" refString="Gonzalez, V. H. &amp; Roubik, D. W. (2008) Especies nuevas y filogenia de las abejas de fuego, Oxytrigona (Hymenoptera: Apidae, Meliponini). Acta Zoologica Mexicana, 24 (1), 43 - 71. https: // doi. org / 10.21829 / azm. 2008.241615" type="journal article" year="2008">Gonzalez &amp; Roubik 2008</bibRefCitation>
); setae on distal end of scape shorter than 0.5 × MOD....................................................................................
<emphasis id="B968183C577EFFFBCFFCF8DEC381C919" bold="true" box="[1167,1436,1839,1861]" pageId="22" pageNumber="391">
<taxonomicName id="4C1CBFAD577EFFFBCFFCF8DEC357C919" authorityName="Lepeco" authorityYear="2024" box="[1167,1354,1839,1861]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="391" phylum="Arthropoda" rank="species" species="pallida" status="sp. nov.">
<emphasis id="B968183C577EFFFBCFFCF8DEC357C919" bold="true" box="[1167,1354,1839,1861]" italics="true" pageId="22" pageNumber="391">Oxytrigona pallida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BA547577EFFFBCE23F8C1C381C919" box="[1360,1436,1840,1861]" pageId="22" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E8DC8B577EFFFACBE4F89BC381CF7E" lastPageId="23" lastPageNumber="392" pageId="22" pageNumber="391">
<paragraph id="8BA3C42E577EFFFBCBE4F89BC380C9A9" blockId="22.[151,1437,1898,2037]" pageId="22" pageNumber="391">
<keyLead id="30ED671B577EFFFBCBE4F89BC380C9A9" pageId="22" pageNumber="391">
<emphasis id="B968183C577EFFFBCBE4F89BC6A9C923" bold="true" box="[151,180,1898,1919]" pageId="22" pageNumber="391">11.</emphasis>
Erect setae on vertex yellowish; hind basitarsus relatively slender, with a slightly curved dorsal margin, with the posterior distal corner somewhat angled towards a straight distal margin (
<figureCitation id="1327D8AB577EFFFBC867F876C566C9C1" box="[788,891,1927,1949]" captionStart="FIGURE 3" captionStartId="6.[151,250,1583,1607]" captionTargetBox="[151,1436,180,1558]" captionTargetId="figure-25@6.[151,1436,180,1558]" captionTargetPageId="6" captionText="FIGURE 3. A) Oxytrigona cagafogo (Müller, 1874), worker from São João de Petrópolis (Espírio Santo, Brazil, RPSP), dorsal view of mesosoma, scale bar: 0.5 mm; B) Oxytrigona flaveola (Friese, 1900), worker from Costa Marques (Rondônia, Brazil, RPSP), anterior view of dorsal portion of the head, scale bar: 0.5 mm; C) O. cagafogo, worker from São Sebastião (São Paulo, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; D) O. flaveola, worker from Costa Marques (Rodônia, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; E) O. cagafogo, worker from Salvador (Bahia, Brazil, RPSP), forewing, scale bar: 1 mm;; F) O. flaveola, worker from Pimenteira (Rodônia, Brazil, RPSP), forewing, scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13747185" httpUri="https://zenodo.org/record/13747185/files/figure.png" pageId="22" pageNumber="391">Figure 3D</figureCitation>
); distal half of wing membrane hyaline, at most slightly yellowish dorsally (
<figureCitation id="1327D8AB577EFFFBCAD3F854C41FC9E6" box="[416,514,1957,1979]" captionStart="FIGURE 3" captionStartId="6.[151,250,1583,1607]" captionTargetBox="[151,1436,180,1558]" captionTargetId="figure-25@6.[151,1436,180,1558]" captionTargetPageId="6" captionText="FIGURE 3. A) Oxytrigona cagafogo (Müller, 1874), worker from São João de Petrópolis (Espírio Santo, Brazil, RPSP), dorsal view of mesosoma, scale bar: 0.5 mm; B) Oxytrigona flaveola (Friese, 1900), worker from Costa Marques (Rondônia, Brazil, RPSP), anterior view of dorsal portion of the head, scale bar: 0.5 mm; C) O. cagafogo, worker from São Sebastião (São Paulo, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; D) O. flaveola, worker from Costa Marques (Rodônia, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; E) O. cagafogo, worker from Salvador (Bahia, Brazil, RPSP), forewing, scale bar: 1 mm;; F) O. flaveola, worker from Pimenteira (Rodônia, Brazil, RPSP), forewing, scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13747185" httpUri="https://zenodo.org/record/13747185/files/figure.png" pageId="22" pageNumber="391">Figure 3F</figureCitation>
); posterior margin of S6 somewhat pointed (see figure 65 of
<bibRefCitation id="EF8DB9DF577EFFFBCF22F854C351C9E6" author="Gonzalez, V. H. &amp; Roubik, D. W." box="[1105,1356,1957,1979]" pageId="22" pageNumber="391" pagination="43 - 71" refId="ref27261" refString="Gonzalez, V. H. &amp; Roubik, D. W. (2008) Especies nuevas y filogenia de las abejas de fuego, Oxytrigona (Hymenoptera: Apidae, Meliponini). Acta Zoologica Mexicana, 24 (1), 43 - 71. https: // doi. org / 10.21829 / azm. 2008.241615" type="journal article" year="2008">Gonzalez &amp; Roubik 2008</bibRefCitation>
); S4S6 with relatively denser coverage of setae, which are
<emphasis id="B968183C577EFFFBC9BDF833C510C98B" box="[718,781,1986,2007]" italics="true" pageId="22" pageNumber="391">mostly</emphasis>
yellowish; posterior margin of T4 mostly with sparse simple setae.......................................................................
<taxonomicName id="4C1CBFAD577EFFFBCF34F82EC380C9A9" ID-CoL="75GTD" authority="(Friese, 1900)" baseAuthorityName="Friese" baseAuthorityYear="1900" box="[1095,1437,2015,2037]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="391" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="B968183C577EFFFBCF34F82EC380C9A9" bold="true" box="[1095,1437,2015,2037]" pageId="22" pageNumber="391">
<emphasis id="B968183C577EFFFBCF34F82EC311C9A9" bold="true" box="[1095,1292,2015,2037]" italics="true" pageId="22" pageNumber="391">Oxytrigona flaveola</emphasis>
(Friese, 1900)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577FFFFACBE4FF69C381CF7E" blockId="23.[151,1436,151,291]" pageId="23" pageNumber="392">
<keyLead id="30ED671B577FFFFACBE4FF69C381CF7E" pageId="23" pageNumber="392">
- Erect setae on vertex light brown to black; hind basitarsus relatively broad, with a strongly curved dorsal margin, with the posterior distal corner somewhat rounded towards a curved distal margin (as in
<figureCitation id="1327D8AB577FFFFAC889FF44C274CE96" box="[1018,1129,181,203]" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1930]" captionTargetBox="[163,1435,846,1880]" captionTargetId="figure-360@3.[151,1435,846,1880]" captionTargetPageId="3" captionText="FIGURE 1. Holotype worker of Oxytrigona banana sp. nov. (RPSP 23.1802) from Ribeirão Preto (São Paulo, Brazil). A) habitus, lateral view, scale bar: 1 mm; B) dorsal view, scale bar: 1 mm; C) head, frontal view, scale bar: 1 mm; D) hind basitarsus, scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747179" httpUri="https://zenodo.org/record/13747179/files/figure.png" pageId="23" pageNumber="392">Figures 1D</figureCitation>
,
<figureCitation id="1327D8AB577FFFFACF05FF44C288CE96" box="[1142,1173,181,202]" captionStart="FIGURE 3" captionStartId="6.[151,250,1583,1607]" captionTargetBox="[151,1436,180,1558]" captionTargetId="figure-25@6.[151,1436,180,1558]" captionTargetPageId="6" captionText="FIGURE 3. A) Oxytrigona cagafogo (Müller, 1874), worker from São João de Petrópolis (Espírio Santo, Brazil, RPSP), dorsal view of mesosoma, scale bar: 0.5 mm; B) Oxytrigona flaveola (Friese, 1900), worker from Costa Marques (Rondônia, Brazil, RPSP), anterior view of dorsal portion of the head, scale bar: 0.5 mm; C) O. cagafogo, worker from São Sebastião (São Paulo, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; D) O. flaveola, worker from Costa Marques (Rodônia, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; E) O. cagafogo, worker from Salvador (Bahia, Brazil, RPSP), forewing, scale bar: 1 mm;; F) O. flaveola, worker from Pimenteira (Rodônia, Brazil, RPSP), forewing, scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13747185" httpUri="https://zenodo.org/record/13747185/files/figure.png" pageId="23" pageNumber="392">3C</figureCitation>
); color of wing membrane
<emphasis id="B968183C577FFFFACB91FF23C72FCEBB" box="[226,306,210,231]" italics="true" pageId="23" pageNumber="392">variable</emphasis>
,
<emphasis id="B968183C577FFFFACA4DFF23C773CEBB" box="[318,366,210,231]" italics="true" pageId="23" pageNumber="392">often</emphasis>
with light or dark brown spots (as in
<figureCitation id="1327D8AB577FFFFAC9A9FF23C55ACEBB" box="[730,839,210,232]" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1930]" captionTargetBox="[163,1435,846,1880]" captionTargetId="figure-360@3.[151,1435,846,1880]" captionTargetPageId="3" captionText="FIGURE 1. Holotype worker of Oxytrigona banana sp. nov. (RPSP 23.1802) from Ribeirão Preto (São Paulo, Brazil). A) habitus, lateral view, scale bar: 1 mm; B) dorsal view, scale bar: 1 mm; C) head, frontal view, scale bar: 1 mm; D) hind basitarsus, scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747179" httpUri="https://zenodo.org/record/13747179/files/figure.png" pageId="23" pageNumber="392">Figures 1A</figureCitation>
,
<figureCitation id="1327D8AB577FFFFAC821FF23C572CEBB" box="[850,879,210,231]" captionStart="FIGURE 3" captionStartId="6.[151,250,1583,1607]" captionTargetBox="[151,1436,180,1558]" captionTargetId="figure-25@6.[151,1436,180,1558]" captionTargetPageId="6" captionText="FIGURE 3. A) Oxytrigona cagafogo (Müller, 1874), worker from São João de Petrópolis (Espírio Santo, Brazil, RPSP), dorsal view of mesosoma, scale bar: 0.5 mm; B) Oxytrigona flaveola (Friese, 1900), worker from Costa Marques (Rondônia, Brazil, RPSP), anterior view of dorsal portion of the head, scale bar: 0.5 mm; C) O. cagafogo, worker from São Sebastião (São Paulo, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; D) O. flaveola, worker from Costa Marques (Rodônia, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; E) O. cagafogo, worker from Salvador (Bahia, Brazil, RPSP), forewing, scale bar: 1 mm;; F) O. flaveola, worker from Pimenteira (Rodônia, Brazil, RPSP), forewing, scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13747185" httpUri="https://zenodo.org/record/13747185/files/figure.png" pageId="23" pageNumber="392">3E</figureCitation>
); posterior margin of S6 widely rounded (see figure 62 of
<bibRefCitation id="EF8DB9DF577FFFFACB91FF1EC7FDCF59" author="Gonzalez, V. H. &amp; Roubik, D. W." box="[226,480,239,261]" pageId="23" pageNumber="392" pagination="43 - 71" refId="ref27261" refString="Gonzalez, V. H. &amp; Roubik, D. W. (2008) Especies nuevas y filogenia de las abejas de fuego, Oxytrigona (Hymenoptera: Apidae, Meliponini). Acta Zoologica Mexicana, 24 (1), 43 - 71. https: // doi. org / 10.21829 / azm. 2008.241615" type="journal article" year="2008">Gonzalez &amp; Roubik 2008</bibRefCitation>
); S4S6 with sparser coverage of setae, which are entirely or mostly dark; posterior margin of T4 with relatively denser coverage of branched setae..........................................................
<emphasis id="B968183C577FFFFACEF7FEFCC381CF7E" bold="true" box="[1412,1436,269,290]" pageId="23" pageNumber="392">12</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E8DC8B577FFFFACBE4FEB6C381CFB3" pageId="23" pageNumber="392">
<paragraph id="8BA3C42E577FFFFACBE4FEB6C380CFC4" blockId="23.[151,1437,327,496]" pageId="23" pageNumber="392">
<keyLead id="30ED671B577FFFFACBE4FEB6C380CFC4" pageId="23" pageNumber="392">
<emphasis id="B968183C577FFFFACBE4FEB6C6ABCF00" bold="true" box="[151,182,327,348]" pageId="23" pageNumber="392">12.</emphasis>
Mesoscutum entirely black,
<emphasis id="B968183C577FFFFACA9CFEB9C44ECF01" box="[495,595,328,349]" italics="true" pageId="23" pageNumber="392">sometimes</emphasis>
with a lighter stripe adjacent to tegula (
<figureCitation id="1327D8AB577FFFFAC8BBFEB6C233CF01" box="[968,1070,327,349]" captionStart="FIGURE 3" captionStartId="6.[151,250,1583,1607]" captionTargetBox="[151,1436,180,1558]" captionTargetId="figure-25@6.[151,1436,180,1558]" captionTargetPageId="6" captionText="FIGURE 3. A) Oxytrigona cagafogo (Müller, 1874), worker from São João de Petrópolis (Espírio Santo, Brazil, RPSP), dorsal view of mesosoma, scale bar: 0.5 mm; B) Oxytrigona flaveola (Friese, 1900), worker from Costa Marques (Rondônia, Brazil, RPSP), anterior view of dorsal portion of the head, scale bar: 0.5 mm; C) O. cagafogo, worker from São Sebastião (São Paulo, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; D) O. flaveola, worker from Costa Marques (Rodônia, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; E) O. cagafogo, worker from Salvador (Bahia, Brazil, RPSP), forewing, scale bar: 1 mm;; F) O. flaveola, worker from Pimenteira (Rodônia, Brazil, RPSP), forewing, scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13747185" httpUri="https://zenodo.org/record/13747185/files/figure.png" pageId="23" pageNumber="392">Figure 3A</figureCitation>
); forewing with distal portion of radial cell and entire marginal cell brownish infumate (
<figureCitation id="1327D8AB577FFFFAC9C7FE94C505CF26" box="[692,792,357,379]" captionStart="FIGURE 3" captionStartId="6.[151,250,1583,1607]" captionTargetBox="[151,1436,180,1558]" captionTargetId="figure-25@6.[151,1436,180,1558]" captionTargetPageId="6" captionText="FIGURE 3. A) Oxytrigona cagafogo (Müller, 1874), worker from São João de Petrópolis (Espírio Santo, Brazil, RPSP), dorsal view of mesosoma, scale bar: 0.5 mm; B) Oxytrigona flaveola (Friese, 1900), worker from Costa Marques (Rondônia, Brazil, RPSP), anterior view of dorsal portion of the head, scale bar: 0.5 mm; C) O. cagafogo, worker from São Sebastião (São Paulo, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; D) O. flaveola, worker from Costa Marques (Rodônia, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; E) O. cagafogo, worker from Salvador (Bahia, Brazil, RPSP), forewing, scale bar: 1 mm;; F) O. flaveola, worker from Pimenteira (Rodônia, Brazil, RPSP), forewing, scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13747185" httpUri="https://zenodo.org/record/13747185/files/figure.png" pageId="23" pageNumber="392">Figure 3E</figureCitation>
); labrum and mandible mostly dark brown to black..................................................................................
<taxonomicName id="4C1CBFAD577FFFFACF46FE73C380CFC4" authority="(Muller, 1874)" baseAuthorityName="Muller" baseAuthorityYear="1874" box="[1077,1437,386,408]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="392" phylum="Arthropoda" rank="species" species="cagafogo">
<emphasis id="B968183C577FFFFACF46FE73C380CFC4" bold="true" box="[1077,1437,386,408]" pageId="23" pageNumber="392">
<emphasis id="B968183C577FFFFACF46FE73C31BCFC4" bold="true" box="[1077,1286,386,408]" italics="true" pageId="23" pageNumber="392">Oxytrigona cagafogo</emphasis>
(
<bibRefCitation id="EF8DB9DF577FFFFACE61FE73C388CFC4" author="Muller, H." box="[1298,1429,386,408]" pageId="23" pageNumber="392" pagination="31 - 32" refId="ref27649" refString="Muller, H. (1874) Larvae of Membracis serving as milk-cattle to a Brazilian species of bee. Nature, 10, 31 - 32. https: // doi. org / 10.1038 / 010031 a 0" type="journal article" year="1874">Müller, 1874</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577FFFFACBE4FE51C381CFB3" blockId="23.[151,1437,327,496]" pageId="23" pageNumber="392">
<keyLead id="30ED671B577FFFFACBE4FE51C381CFB3" pageId="23" pageNumber="392">
<emphasis id="B968183C577FFFFACBE4FE51C682CFE9" bold="true" box="[151,159,416,437]" pageId="23" pageNumber="392">-</emphasis>
Mesoscutum
<emphasis id="B968183C577FFFFACA10FE6EC7B5CFE8" box="[355,424,415,436]" italics="true" pageId="23" pageNumber="392">usually</emphasis>
yellow to light brown between notauli and darker laterad,
<emphasis id="B968183C577FFFFAC896FE51C254CFE9" box="[997,1097,416,437]" italics="true" pageId="23" pageNumber="392">sometimes</emphasis>
entirely light brown (as in
<figureCitation id="1327D8AB577FFFFACE27FE6EC71DCF8E" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1930]" captionTargetBox="[163,1435,846,1880]" captionTargetId="figure-360@3.[151,1435,846,1880]" captionTargetPageId="3" captionText="FIGURE 1. Holotype worker of Oxytrigona banana sp. nov. (RPSP 23.1802) from Ribeirão Preto (São Paulo, Brazil). A) habitus, lateral view, scale bar: 1 mm; B) dorsal view, scale bar: 1 mm; C) head, frontal view, scale bar: 1 mm; D) hind basitarsus, scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747179" httpUri="https://zenodo.org/record/13747179/files/figure.png" pageId="23" pageNumber="392">Figures 1B</figureCitation>
,
<figureCitation id="1327D8AB577FFFFACA7FFE4CC731CF8E" box="[268,300,445,466]" captionStart="FIGURE 7" captionStartId="20.[151,250,1179,1203]" captionTargetBox="[151,1435,180,1155]" captionTargetId="figure-25@20.[151,1435,180,1155]" captionTargetPageId="20" captionText="FIGURE 7. Worker of Oxytrigona tataira (Smith, 1863), from Viçosa (Ceará, Brazil, RPSP). A) dorsal view of mesosoma; B) detail of mesoscutum; C) lateral view of mesosoma; D) detail of mesepisternum. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747199" httpUri="https://zenodo.org/record/13747199/files/figure.png" pageId="23" pageNumber="392">7A</figureCitation>
); forewing with distal portion of radial cell and entire marginal cell hyaline or yellowish (as in
<figureCitation id="1327D8AB577FFFFACF92FE4CC359CF8E" box="[1249,1348,445,467]" captionStart="FIGURE 3" captionStartId="6.[151,250,1583,1607]" captionTargetBox="[151,1436,180,1558]" captionTargetId="figure-25@6.[151,1436,180,1558]" captionTargetPageId="6" captionText="FIGURE 3. A) Oxytrigona cagafogo (Müller, 1874), worker from São João de Petrópolis (Espírio Santo, Brazil, RPSP), dorsal view of mesosoma, scale bar: 0.5 mm; B) Oxytrigona flaveola (Friese, 1900), worker from Costa Marques (Rondônia, Brazil, RPSP), anterior view of dorsal portion of the head, scale bar: 0.5 mm; C) O. cagafogo, worker from São Sebastião (São Paulo, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; D) O. flaveola, worker from Costa Marques (Rodônia, Brazil, RPSP), anterior view of hind basitarsus, scale bar: 0.5 mm; E) O. cagafogo, worker from Salvador (Bahia, Brazil, RPSP), forewing, scale bar: 1 mm;; F) O. flaveola, worker from Pimenteira (Rodônia, Brazil, RPSP), forewing, scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13747185" httpUri="https://zenodo.org/record/13747185/files/figure.png" pageId="23" pageNumber="392">Figure 3F</figureCitation>
); labrum yellowish to light brown; medial third of mandible
<emphasis id="B968183C577FFFFAC9BDFE2BC50ECFB3" box="[718,787,474,495]" italics="true" pageId="23" pageNumber="392">usually</emphasis>
yellowish to light brown (as in
<figureCitation id="1327D8AB577FFFFACF36FE2BC2BBCFB3" box="[1093,1190,474,496]" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1930]" captionTargetBox="[163,1435,846,1880]" captionTargetId="figure-360@3.[151,1435,846,1880]" captionTargetPageId="3" captionText="FIGURE 1. Holotype worker of Oxytrigona banana sp. nov. (RPSP 23.1802) from Ribeirão Preto (São Paulo, Brazil). A) habitus, lateral view, scale bar: 1 mm; B) dorsal view, scale bar: 1 mm; C) head, frontal view, scale bar: 1 mm; D) hind basitarsus, scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747179" httpUri="https://zenodo.org/record/13747179/files/figure.png" pageId="23" pageNumber="392">Figure 1C</figureCitation>
).................
<emphasis id="B968183C577FFFFACEF7FE2BC381CFB3" bold="true" box="[1412,1436,474,495]" pageId="23" pageNumber="392">13</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E8DC8B577FFFFACBE4FDE4C381CCDE" pageId="23" pageNumber="392">
<paragraph id="8BA3C42E577FFFFACBE4FDE4C380CC14" blockId="23.[151,1437,533,643]" pageId="23" pageNumber="392">
<keyLead id="30ED671B577FFFFACBE4FDE4C380CC14" pageId="23" pageNumber="392">
<emphasis id="B968183C577FFFFACBE4FDE4C6A8CC76" bold="true" box="[151,181,533,554]" pageId="23" pageNumber="392">13.</emphasis>
Mesoscutum with short branched setae reaching posterior level of tegula, coverage of branched setae relatively denser (
<figureCitation id="1327D8AB577FFFFACE2FFDE4C71CCC1B" captionStart="FIGURE 7" captionStartId="20.[151,250,1179,1203]" captionTargetBox="[151,1435,180,1155]" captionTargetId="figure-25@20.[151,1435,180,1155]" captionTargetPageId="20" captionText="FIGURE 7. Worker of Oxytrigona tataira (Smith, 1863), from Viçosa (Ceará, Brazil, RPSP). A) dorsal view of mesosoma; B) detail of mesoscutum; C) lateral view of mesosoma; D) detail of mesepisternum. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747199" httpUri="https://zenodo.org/record/13747199/files/figure.png" pageId="23" pageNumber="392">Figure 7B</figureCitation>
); mesepisternum with dense coverage of branched setae (
<figureCitation id="1327D8AB577FFFFAC85BFDC3C5AFCC1B" box="[808,946,562,584]" captionStart="FIGURE 7" captionStartId="20.[151,250,1179,1203]" captionTargetBox="[151,1435,180,1155]" captionTargetId="figure-25@20.[151,1435,180,1155]" captionTargetPageId="20" captionText="FIGURE 7. Worker of Oxytrigona tataira (Smith, 1863), from Viçosa (Ceará, Brazil, RPSP). A) dorsal view of mesosoma; B) detail of mesoscutum; C) lateral view of mesosoma; D) detail of mesepisternum. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747199" httpUri="https://zenodo.org/record/13747199/files/figure.png" pageId="23" pageNumber="392">Figures 7C, D</figureCitation>
)...........
<taxonomicName id="4C1CBFAD577FFFFACF25FDC3C380CC14" ID-CoL="4BLV8" authority="(Smith, 1863)" baseAuthorityName="Smith" baseAuthorityYear="1863" box="[1110,1437,562,584]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="392" phylum="Arthropoda" rank="species" species="tataira">
<emphasis id="B968183C577FFFFACF25FDC3C380CC14" bold="true" box="[1110,1437,562,584]" pageId="23" pageNumber="392">
<emphasis id="B968183C577FFFFACF25FDC3C310CC1B" bold="true" box="[1110,1293,562,583]" italics="true" pageId="23" pageNumber="392">Oxytrigona tataira</emphasis>
(Smith, 1863)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA3C42E577FFFFACBE4FDA1C381CCDE" blockId="23.[151,1437,533,643]" pageId="23" pageNumber="392">
<keyLead id="30ED671B577FFFFACBE4FDA1C381CCDE" pageId="23" pageNumber="392">
<emphasis id="B968183C577FFFFACBE4FDA1C682CC39" bold="true" box="[151,159,592,613]" pageId="23" pageNumber="392">-</emphasis>
Mesoscutum with short branched setae restricted to anterior portion of sclerite, not reaching posterior level of tegula, coverage of branched setae sparse (
<figureCitation id="1327D8AB577FFFFACAABFD9CC421CCDE" box="[472,572,621,643]" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1930]" captionTargetBox="[163,1435,846,1880]" captionTargetId="figure-360@3.[151,1435,846,1880]" captionTargetPageId="3" captionText="FIGURE 1. Holotype worker of Oxytrigona banana sp. nov. (RPSP 23.1802) from Ribeirão Preto (São Paulo, Brazil). A) habitus, lateral view, scale bar: 1 mm; B) dorsal view, scale bar: 1 mm; C) head, frontal view, scale bar: 1 mm; D) hind basitarsus, scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13747179" httpUri="https://zenodo.org/record/13747179/files/figure.png" pageId="23" pageNumber="392">Figure 1B</figureCitation>
); branched setae on mesepisternum relatively sparse......
<emphasis id="B968183C577FFFFACFFAFD9CC381CCDE" bold="true" box="[1161,1436,620,642]" pageId="23" pageNumber="392">
<taxonomicName id="4C1CBFAD577FFFFACFFAFD9CC357CCDE" authorityName="Lepeco" authorityYear="2024" box="[1161,1354,620,642]" class="Insecta" family="Apidae" genus="Oxytrigona" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="392" phylum="Arthropoda" rank="species" species="banana" status="sp. nov.">
<emphasis id="B968183C577FFFFACFFAFD9CC357CCDE" bold="true" box="[1161,1354,620,642]" italics="true" pageId="23" pageNumber="392">Oxytrigona banana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BA547577FFFFACE23FD9CC381CCDE" box="[1360,1436,621,642]" pageId="23" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>