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<document id="36B0EDE38BC324193FAB2C99BA33B698" ID-CLB-Dataset="27441" ID-DOI="10.11646/zootaxa.4559.3.3" ID-GBIF-Dataset="f39d035d-0ca0-4d8a-888d-700de6678d8b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2627157" ID-ZooBank="04458AF0-1E51-4468-A875-9EEDCA34BE19" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554321924814" checkinUser="plazi" docAuthor="Dietrich, Christopher H. &amp; Zahniser, James N." docDate="2019" docId="7B3BEE3BFFE9FFA4EDA2A4A9FE40FF42" docLanguage="en" docName="zootaxa.4559.3.3.pdf" docOrigin="Zootaxa 4559 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Makilingia woodworthi Baker 1924" docType="treatment" docVersion="8" lastPageNumber="498" masterDocId="87029643FFFEFFBDED35A12BFFEEFFBA" masterDocTitle="Review of the leafhopper genus Makilingia Baker (Hemiptera: Cicadellidae: Mileewinae)" masterLastPageNumber="500" masterPageNumber="473" pageNumber="496" updateTime="1698717634450" updateUser="ExternalLinkService">
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<mods:title id="52B6AE69254E4CB9619B91213178379D">Review of the leafhopper genus Makilingia Baker (Hemiptera: Cicadellidae: Mileewinae)</mods:title>
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<mods:namePart id="8674E4563A1249C76F28C67DFFFE6FF6">Dietrich, Christopher H.</mods:namePart>
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<treatment id="7B3BEE3BFFE9FFA4EDA2A4A9FE40FF42" ID-DOI="http://doi.org/10.5281/zenodo.5936464" ID-GBIF-Taxon="154534483" ID-Zenodo-Dep="5936464" LSID="urn:lsid:plazi:treatment:7B3BEE3BFFE9FFA4EDA2A4A9FE40FF42" httpUri="http://treatment.plazi.org/id/7B3BEE3BFFE9FFA4EDA2A4A9FE40FF42" lastPageId="25" lastPageNumber="498" pageId="23" pageNumber="496">
<subSubSection id="BB880CA6FFE9FFAAEDA2A4A9FDEBFA27" box="[151,517,1410,1437]" pageId="23" pageNumber="496" type="nomenclature">
<paragraph id="F32D5F2DFFE9FFAAEDA2A4A9FDEBFA27" blockId="23.[151,517,1410,1470]" box="[151,517,1410,1437]" pageId="23" pageNumber="496">
<heading id="A865E841FFE9FFAAEDA2A4A9FDEBFA27" bold="true" box="[151,517,1410,1437]" fontSize="11" level="1" pageId="23" pageNumber="496" reason="1">
<taxonomicName id="349224AEFFE9FFAAEDA2A4A9FDEBFA27" ID-CoL="3XJZF" authority="Baker" authorityName="Baker" authorityYear="1924" box="[151,517,1410,1437]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="496" phylum="Arthropoda" rank="species" species="woodworthi">
<emphasis id="C1E6833FFFE9FFAAEDA2A4A9FDEBFA27" bold="true" box="[151,517,1410,1437]" pageId="23" pageNumber="496">
<emphasis id="C1E6833FFFE9FFAAEDA2A4A9FE5CFA26" bold="true" box="[151,434,1410,1436]" italics="true" pageId="23" pageNumber="496">Makilingia woodworthi</emphasis>
Baker
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BB880CA6FFE9FFAAEDA2A48EFE6FFA04" box="[151,385,1445,1470]" pageId="23" pageNumber="496" type="description">
<paragraph id="F32D5F2DFFE9FFAAEDA2A48EFE6FFA04" blockId="23.[151,517,1410,1470]" box="[151,385,1445,1470]" pageId="23" pageNumber="496">
(
<figureCitation id="6BA943A8FFE9FFAAEDAAA48EFF0BFA04" box="[159,229,1445,1470]" captionStart="FIGURE 2" captionStartId="4.[151,250,1890,1912]" captionTargetBox="[194,1359,197,1861]" captionTargetId="figure@4.[194,1394,193,1873]" captionTargetPageId="4" captionText="FIGURE 2. Makilingia spp., dorsal and lateral habitus: AB, M. paranigra, holotype male; CD, M. pruinosa, male; EF, M. siamensis, holotype male; GH, M. sibuyanensis lectotype male. IJ, M. similima, lectotype male; KL, M. speciosa, paralectotype female; MN, M. surigaoensis, lectotype male; OP, M. suturalis, lectotype male; QR, M. tenebrifrons, holotype male. ST, M. tettigonoides, lectotype male. UV, M. uncinata, holotype male; WX, M. viraktamathi, holotype male; YZ, M. woodworthi, holotype male. AABB, M. xanthopicta, holotype male." httpUri="https://zenodo.org/record/2627161/files/figure.png" pageId="23" pageNumber="496">Figs 2</figureCitation>
YZ, 7EEII)
</paragraph>
</subSubSection>
<subSubSection id="BB880CA6FFE9FFAAEDA2A4C6FDDDF89A" pageId="23" pageNumber="496" type="reference_group">
<paragraph id="F32D5F2DFFE9FFAAEDA2A4C6FD6FF9B9" blockId="23.[151,641,1517,1539]" box="[151,641,1517,1539]" pageId="23" pageNumber="496">
<treatmentCitationGroup id="D3827803FFE9FFAAEDA2A4C6FD6FF9B9" box="[151,641,1517,1539]" pageId="23" pageNumber="496">
<taxonomicName id="349224AEFFE9FFAAEDA2A4C6FDC0F9B9" ID-CoL="3XJZF" authority="Baker, 1924: 68" authorityName="Baker" authorityPageNumber="68" authorityYear="1924" box="[151,558,1517,1539]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="496" phylum="Arthropoda" rank="species" species="woodworthi">
<emphasis id="C1E6833FFFE9FFAAEDA2A4C6FE6BF9B8" box="[151,389,1517,1538]" italics="true" pageId="23" pageNumber="496">Makilingia woodworthi</emphasis>
<treatmentCitation id="7233793CFFE9FFAAECB9A4C6FDC0F9B9" author="Baker, C. F." box="[396,558,1517,1539]" page="68" pageId="23" pageNumber="496" year="1924">
<bibRefCitation id="970322DCFFE9FFAAECB9A4C6FDE7F9B9" author="Baker, C. F." box="[396,521,1517,1539]" pageId="23" pageNumber="496" pagination="57 - 71" refId="ref17954" refString="Baker, C. F. (1924) The genus Makilingia (Jassoidea) in the Philippines. Philippine Journal of Science, 24 (1), 57 - 71." type="journal article" year="1924">Baker, 1924</bibRefCitation>
: 68
</treatmentCitation>
</taxonomicName>
, Fig. 13
</treatmentCitationGroup>
</paragraph>
<paragraph id="F32D5F2DFFE9FFAAEDA2A704FDDDF89A" blockId="23.[151,1437,1583,2005]" pageId="23" pageNumber="496">Male genitalia. Pygofer appendage gradually tapered and evenly curved dorsad from base to apex, extended beyond lobe. Subgenital plate moderately long, curved gradually and slightly dorsad, distal section less darkly pigmented than basal section, dorsal callosity short and relatively tall. Style without preapical lobe, apex enlarged with prominent preapical heel, distal section relatively long, with several transverse rugae. Aedeagus in lateral view with preatrium as long as shaft, sclerotized part narrow and evenly curved dorsad; shaft moderately broad, curved slightly anterodorsad, apex bluntly rounded; in posterior view with preatrium slightly constricted, atrial processes broad and weakly bilobed.</paragraph>
</subSubSection>
<subSubSection id="BB880CA6FFE9FFAAEDF2A607FE7EF8FF" box="[199,400,1836,1861]" pageId="23" pageNumber="496" type="description">
<paragraph id="F32D5F2DFFE9FFAAEDF2A607FE7EF8FF" blockId="23.[151,1437,1583,2005]" box="[199,400,1836,1861]" pageId="23" pageNumber="496">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="BB880CA6FFE9FFA4EDF2A664FE40FF42" lastPageId="25" lastPageNumber="498" pageId="23" pageNumber="496" type="materials_examined">
<paragraph id="F32D5F2DFFE9FFAAEDF2A664FC78F86E" blockId="23.[151,1437,1583,2005]" pageId="23" pageNumber="496">
<emphasis id="C1E6833FFFE9FFAAEDF2A664FE5DF8D2" bold="true" box="[199,435,1871,1896]" pageId="23" pageNumber="496">Material examined.</emphasis>
<materialsCitation id="43FA5570FFE9FFAAEC88A664FC78F86E" ID-GBIF-Occurrence="2234227262" collectionCode="USNM" collectorName="Det. J. N. Zahniser" determinerName="Det. J. N. Zahniser" location="Other" pageId="23" pageNumber="496" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="2C29E18FFFE9FFAAEC88A664FDCBF8D2" box="[445,549,1871,1896]" pageId="23" pageNumber="496" type="holotype">Holotype</typeStatus>
male, labeled “20426; Mt. Makiling / Luzon, Baker;
<typeStatus id="2C29E18FFFE9FFAAE9ACA664FAEFF8D2" box="[1177,1281,1871,1896]" pageId="23" pageNumber="496" type="holotype">Holotype</typeStatus>
/ over / only spcmn. / in Baker / colln D.A.Y.;
<taxonomicName id="349224AEFFE9FFAAEF07A65FFD40F836" authorityName="Baker" authorityYear="1904" box="[562,686,1908,1932]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="496" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6833FFFE9FFAAEF07A65FFD40F836" box="[562,686,1908,1932]" italics="true" pageId="23" pageNumber="496">Makilingia</emphasis>
</taxonomicName>
/
<taxonomicName id="349224AEFFE9FFAAEFFFA65FFCBEF836" authorityName="Baker" authorityYear="1924" box="[714,848,1908,1932]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="496" phylum="Arthropoda" rank="species" species="woodworthi">
<emphasis id="C1E6833FFFE9FFAAEFFFA65FFCBEF836" box="[714,848,1908,1932]" italics="true" pageId="23" pageNumber="496">woodworthi</emphasis>
</taxonomicName>
/ Baker; JNZ_AA0091; USNMENT / 01513417”.
<location id="F64D09F6FFE9FFAAEDA2A6B3FF39F80B" LSID="urn:lsid:plazi:treatment:7B3BEE3BFFE9FFA4EDA2A4A9FE40FF42:F64D09F6FFE9FFAAEDA2A6B3FF39F80B" box="[151,215,1944,1969]" name="Other" pageId="23" pageNumber="496">Other</location>
material.
<specimenCount id="E59494A4FFE9FFAAEC79A6B3FE72F80B" box="[332,412,1944,1969]" pageId="23" pageNumber="496" type="male">1 male</specimenCount>
, labeled “Mt. Makiling / Luzon, Baker; St. 3-1862C /
<collectionCode id="9583C7E8FFE9FFAAE93AA6B3FB8AF80A" box="[1039,1124,1944,1968]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="496" type="Museum">USNM</collectionCode>
;
<taxonomicName id="349224AEFFE9FFAAE945A6B3FB02F80A" authorityName="Baker" authorityYear="1904" box="[1136,1260,1944,1968]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="496" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6833FFFE9FFAAE945A6B3FB02F80A" box="[1136,1260,1944,1968]" italics="true" pageId="23" pageNumber="496">Makilingia</emphasis>
</taxonomicName>
/
<taxonomicName id="349224AEFFE9FFAAE830A6B3FA65F80A" authorityName="Baker" authorityYear="1924" box="[1285,1419,1944,1968]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="496" phylum="Arthropoda" rank="species" species="woodworthi">
<emphasis id="C1E6833FFFE9FFAAE830A6B3FA65F80A" box="[1285,1419,1944,1968]" italics="true" pageId="23" pageNumber="496">woodworthi</emphasis>
</taxonomicName>
/ Baker /
<determinerName id="06F1B3E9FFE9FFAAEDDAA696FE52F86F" box="[239,444,1980,2005]" pageId="23" pageNumber="496">
<collectorName id="5E673AFBFFE9FFAAEDDAA696FE52F86F" box="[239,444,1980,2005]" pageId="23" pageNumber="496">Det. J.N. Zahniser</collectorName>
</determinerName>
2018; USNMENT / 01513418” (
<collectionCode id="9583C7E8FFE9FFAAEE07A697FC67F86E" box="[818,905,1980,2004]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="496" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="A7ED0FA5FFE6FFA5EDA2A644FDEBF85B" httpUri="https://zenodo.org/record/2627171/files/figure.png" pageId="24" pageNumber="497" startId="24.[151,250,1903,1925]" targetBox="[188,1388,202,1878]" targetPageId="24">
<paragraph id="F32D5F2DFFE6FFA5EDA2A644FDEBF85B" blockId="24.[151,1436,1903,2017]" pageId="24" pageNumber="497">
<emphasis id="C1E6833FFFE6FFA5EDA2A644FEFCF83E" bold="true" box="[151,274,1903,1925]" pageId="24" pageNumber="497">FIGURE 7.</emphasis>
<taxonomicName id="349224AEFFE6FFA5EC2DA644FE69F83E" authorityName="Baker" authorityYear="1904" box="[280,391,1903,1924]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6833FFFE6FFA5EC2DA644FE69F83E" box="[280,391,1903,1924]" italics="true" pageId="24" pageNumber="497">Makilingia</emphasis>
</taxonomicName>
spp. male genitalia: AE,
<taxonomicName id="349224AEFFE6FFA5EFA1A65BFCE2F83F" authorityName="Baker" authorityYear="1924" box="[660,780,1904,1925]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="497" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="C1E6833FFFE6FFA5EFA1A65BFCE2F83F" box="[660,780,1904,1925]" italics="true" pageId="24" pageNumber="497">M. speciosa</emphasis>
</taxonomicName>
, genital capsule in lateral view, subgenital plates in ventral view, genitalia in lateral and ventral views; FJ, same,
<taxonomicName id="349224AEFFE6FFA5EFA5A6A4FCDAF81E" authorityName="Baker" authorityYear="1924" box="[656,820,1935,1956]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="497" phylum="Arthropoda" rank="species" species="surigaoensis">
<emphasis id="C1E6833FFFE6FFA5EFA5A6A4FCDAF81E" box="[656,820,1935,1956]" italics="true" pageId="24" pageNumber="497">M. surigaoensis</emphasis>
</taxonomicName>
; KO, same,
<taxonomicName id="349224AEFFE6FFA5EEF1A6A4FB87F819" authorityName="Baker" authorityYear="1924" box="[964,1129,1934,1956]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="497" phylum="Arthropoda" rank="species" species="tettigonoides">
<emphasis id="C1E6833FFFE6FFA5EEF1A6A4FB87F819" box="[964,1129,1934,1956]" italics="true" pageId="24" pageNumber="497">M. tettigonoides</emphasis>
</taxonomicName>
; PT, same,
<taxonomicName id="349224AEFFE6FFA5E9C5A6A4FA7BF819" authorityName="Dietrich &amp; Zahniser" authorityYear="2019" box="[1264,1429,1934,1956]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="497" phylum="Arthropoda" rank="species" species="viraktamathi">
<emphasis id="C1E6833FFFE6FFA5E9C5A6A4FA7BF819" box="[1264,1429,1934,1956]" italics="true" pageId="24" pageNumber="497">M. viraktamathi</emphasis>
</taxonomicName>
; UY, same,
<taxonomicName id="349224AEFFE6FFA5EC21A686FE5EF87B" authorityName="Baker" authorityYear="1924" box="[276,432,1964,1986]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="497" phylum="Arthropoda" rank="species" species="woodworthi">
<emphasis id="C1E6833FFFE6FFA5EC21A686FE5EF87B" box="[276,432,1964,1986]" italics="true" pageId="24" pageNumber="497">M. woodworthi</emphasis>
</taxonomicName>
; ZDD,
<taxonomicName id="349224AEFFE6FFA5EF39A686FD49F87B" authorityName="Dietrich &amp; Zahniser" authorityYear="2019" box="[524,679,1964,1986]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="497" phylum="Arthropoda" rank="species" species="xanthopicta">
<emphasis id="C1E6833FFFE6FFA5EF39A686FD49F87B" box="[524,679,1964,1986]" italics="true" pageId="24" pageNumber="497">M. xanthopicta</emphasis>
</taxonomicName>
. Note: The genitalia of
<taxonomicName id="349224AEFFE6FFA5EEA8A686FBF9F878" authorityName="Baker" authorityYear="1924" box="[925,1047,1965,1986]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="497" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="C1E6833FFFE6FFA5EEA8A686FBF9F878" box="[925,1047,1965,1986]" italics="true" pageId="24" pageNumber="497">M. speciosa</emphasis>
</taxonomicName>
are shown with the dorsal connective still attached to the aedeagal atrium.
</paragraph>
</caption>
<paragraph id="F32D5F2DFFE7FFA4EDF2A1BCFE40FF42" blockId="25.[151,1436,151,248]" pageId="25" pageNumber="498">
<emphasis id="C1E6833FFFE7FFA4EDF2A1BCFEFFFF0A" bold="true" box="[199,273,151,176]" pageId="25" pageNumber="498">Notes.</emphasis>
This species is very similar to
<taxonomicName id="349224AEFFE7FFA4EF52A1B2FD19FF0A" authorityName="Melichar" authorityYear="1923" box="[615,759,152,176]" class="Insecta" family="Cicadellidae" genus="Makilingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="498" phylum="Arthropoda" rank="species" species="flavifrons">
<emphasis id="C1E6833FFFE7FFA4EF52A1B2FD19FF0A" box="[615,759,152,176]" italics="true" pageId="25" pageNumber="498">M. flavifrons</emphasis>
</taxonomicName>
in the color pattern of the head but the anterior margin of the head is more broadly rounded, the black markings of the crown are narrower and the margin of the gena is pale orange rather than black.
</paragraph>
</subSubSection>
</treatment>
</document>