treatments-xml/data/7B/3B/87/7B3B87B83602EA32FF3614B90FFD019E.xml
2024-06-21 12:41:18 +02:00

235 lines
29 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="E1577697EC22ABA23A4DA623D5F2B049" ID-DOI="10.11646/zootaxa.4748.1.12" ID-GBIF-Dataset="43a5345c-7b56-42a8-9df6-cb6c2a6d089c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3697562" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583398581213" checkinUser="plazi" docAuthor="Gnezdilov, Vladimir M. &amp; Emeljanov, Alexandr F." docDate="2020" docId="7B3B87B83602EA32FF3614B90FFD019E" docLanguage="en" docName="zootaxa.4748.1.12.pdf" docOrigin="Zootaxa 4748 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cixiinae Spinola 1839" docType="treatment" docVersion="5" lastPageNumber="196" masterDocId="8702FFC03602EA33FFA1121D0B4D036C" masterDocTitle="On the taxonomic position of the genera Krundia, Breukoscelis, and Uphodato (Hemiptera, Auchenorrhyncha, Fulgoroidea) described from the Insect Limestone of the Isle of Wight" masterLastPageNumber="200" masterPageNumber="195" pageNumber="195" updateTime="1698724963647" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="72DE76BBCD1E628C0CC286115BB229C5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6A1DA0CC5168C15C716388BF0526C5A9">
<mods:title id="F594F41D11A8B84E154B00FB5A3B31F0">On the taxonomic position of the genera Krundia, Breukoscelis, and Uphodato (Hemiptera, Auchenorrhyncha, Fulgoroidea) described from the Insect Limestone of the Isle of Wight</mods:title>
</mods:titleInfo>
<mods:name id="397D7BCA91FE94F590FCDE0444C423E4" type="personal">
<mods:role id="6DABBDEFAA1DC9144AA777D0F6268C56">
<mods:roleTerm id="32A4F5976D6E628262BB5AEC5349F2D1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B06DA774921224023215477E316B707">Gnezdilov, Vladimir M.</mods:namePart>
</mods:name>
<mods:name id="FFCE78BAE7EB80F42873C4DAD10BF7D1" type="personal">
<mods:role id="A17FC5A1E85FC33B5BBBCDD9A65CB75D">
<mods:roleTerm id="AB48CFB958345C825A77072555867453">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2898A3E833D443581A0802901578D7FF">Emeljanov, Alexandr F.</mods:namePart>
</mods:name>
<mods:typeOfResource id="60C44295DF0B866D6AAD0F7B529E49E1">text</mods:typeOfResource>
<mods:relatedItem id="97AF4AA8A8B765AF7DBE5B38AD693126" type="host">
<mods:titleInfo id="659D522EF0C66F3AB9D7C261973F6CE5">
<mods:title id="93963ED274EFA402F6FB65679FD1142E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="ED0E52889DEA747CB41558DFEE4BDF3D">
<mods:date id="22AA0B06641C79A4F6399845B2EBB365">2020</mods:date>
<mods:detail id="AD6264FA87EA2B1D4996B84896D9AB87" type="pubDate">
<mods:number id="9C588E11673E7764CB00B3CB515F9F1E">2020-03-05</mods:number>
</mods:detail>
<mods:detail id="5B15933BAEE90F2C387D952F92D39A3D" type="volume">
<mods:number id="C91B22CB83E2697FC642ABA466DBF10B">4748</mods:number>
</mods:detail>
<mods:detail id="6EE80C54859C022CDCFD727708D7998F" type="issue">
<mods:number id="37E9C26137EC1892F5714EB5E68E3EDC">1</mods:number>
</mods:detail>
<mods:extent id="DBECC479F83A40EB7CA6CD8D368010FD" unit="page">
<mods:start id="0E1981F99EF36E8F0B471C46186DAD1B">195</mods:start>
<mods:end id="9A7EF8C25382B9DE41E808C3AFF299C1">200</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D6B60CC1E63408BF4B45F7D10A50BB52">journal article</mods:classification>
<mods:identifier id="2D3757CC38D8ECFA5CC9E4D0E9C1D4EE" type="DOI">10.11646/zootaxa.4748.1.12</mods:identifier>
<mods:identifier id="164EFB61DC0D7FBC8CDDAC70C1E6A275" type="GBIF-Dataset">43a5345c-7b56-42a8-9df6-cb6c2a6d089c</mods:identifier>
<mods:identifier id="84D7D03B0C6EBE1AFDD09A784A335860" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A83229D0A821EE73AB5D70F90C755E8F" type="Zenodo-Dep">3697562</mods:identifier>
<mods:identifier id="E81646238A87E2E22022E54C3EC3EB66" type="ZooBank">F45EA07B-C354-437B-A60D-078C95328010</mods:identifier>
</mods:mods>
<treatment id="7B3B87B83602EA32FF3614B90FFD019E" ID-DOI="http://doi.org/10.5281/zenodo.3704276" ID-GBIF-Taxon="162443235" ID-Zenodo-Dep="3704276" LSID="urn:lsid:plazi:treatment:7B3B87B83602EA32FF3614B90FFD019E" httpUri="http://treatment.plazi.org/id/7B3B87B83602EA32FF3614B90FFD019E" lastPageId="1" lastPageNumber="196" pageId="0" pageNumber="195">
<subSubSection id="BB8865253602EA33FF3614B90972058E" pageId="0" pageNumber="195" type="nomenclature">
<paragraph id="F32D36AE3602EA33FF3614B9097A05D2" blockId="0.[151,1437,1629,2014]" box="[151,567,1700,1726]" pageId="0" pageNumber="195">
<heading id="A86581C23602EA33FF3614B9097A05D2" bold="true" box="[151,567,1700,1726]" fontSize="11" level="1" pageId="0" pageNumber="195" reason="1">
<emphasis id="C1E6EABC3602EA33FF3614B9097A05D2" bold="true" box="[151,567,1700,1726]" pageId="0" pageNumber="195">
Subfamily
<taxonomicName id="34924D2D3602EA33FEBE14B9097A05D2" authority="Spinola, 1839" authorityName="Spinola" authorityYear="1839" box="[287,567,1700,1726]" class="Insecta" family="Cixiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="195" phylum="Arthropoda" rank="family">Cixiinae Spinola, 1839</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="F32D36AE3602EA33FF3614D50972058E" blockId="0.[151,1437,1629,2014]" box="[151,575,1736,1762]" pageId="0" pageNumber="195">
<heading id="A86581C23602EA33FF3614D50972058E" bold="true" box="[151,575,1736,1762]" fontSize="11" level="1" pageId="0" pageNumber="195" reason="1">
<emphasis id="C1E6EABC3602EA33FF3614D50972058E" bold="true" box="[151,575,1736,1762]" pageId="0" pageNumber="195">Tribe Eucarpiini Emeljanov, 2002</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BB8865253602EA32FF3614F10FFD019E" lastPageId="1" lastPageNumber="196" pageId="0" pageNumber="195" type="description">
<paragraph id="F32D36AE3602EA33FF3614F10958046A" blockId="0.[151,1437,1629,2014]" box="[151,533,1772,1799]" pageId="0" pageNumber="195">
<emphasis id="C1E6EABC3602EA33FF3614F10958046A" bold="true" box="[151,533,1772,1799]" pageId="0" pageNumber="195">
Genus
<emphasis id="C1E6EABC3602EA33FF4F14F10A28046B" bold="true" box="[238,357,1772,1799]" italics="true" pageId="0" pageNumber="195">Uphodato</emphasis>
<bibRefCitation id="97034B5F3602EA33FECC14F10958046A" author="Szwedo, J. &amp; Drohojowska, J. &amp; Popov, Yu. A. &amp; Simon, E. &amp; Wegierek, P." box="[365,533,1772,1798]" pageId="0" pageNumber="195" pagination="331 - 396" refId="ref1881" refString="Szwedo, J., Drohojowska, J., Popov, Yu. A., Simon, E. &amp; Wegierek, P. (2019) Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 110, 331 - 396. https: // doi. org / 10.1017 / S 175569101900001 X" type="journal article" year="2019">Szwedo, 2019</bibRefCitation>
</emphasis>
</paragraph>
<paragraph id="F32D36AE3602EA33FF36150F0FFA0422" blockId="0.[151,1437,1629,2014]" pageId="0" pageNumber="195">
<emphasis id="C1E6EABC3602EA33FF36150F0B9E0446" bold="true" box="[151,211,1810,1834]" pageId="0" pageNumber="195">Note.</emphasis>
We suggest placement of
<emphasis id="C1E6EABC3602EA33FE4A150F093F0446" box="[491,626,1810,1834]" italics="true" pageId="0" pageNumber="195">Breukoscelis</emphasis>
<bibRefCitation id="97034B5F3602EA33FDD8150F085D0446" author="Szwedo, J. &amp; Drohojowska, J. &amp; Popov, Yu. A. &amp; Simon, E. &amp; Wegierek, P." box="[633,784,1810,1834]" pageId="0" pageNumber="195" pagination="331 - 396" refId="ref1881" refString="Szwedo, J., Drohojowska, J., Popov, Yu. A., Simon, E. &amp; Wegierek, P. (2019) Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 110, 331 - 396. https: // doi. org / 10.1017 / S 175569101900001 X" type="journal article" year="2019">
Szwedo,
<quantity id="346A9B4B3602EA33FD7A150F085D0446" box="[731,784,1810,1834]" metricMagnitude="1" metricUnit="m" metricValue="5.12826" pageId="0" pageNumber="195" unit="in" value="2019.0">2019</quantity>
</bibRefCitation>
in synonymy under
<emphasis id="C1E6EABC3602EA33FC4D150F0F1A0446" box="[1004,1111,1810,1834]" italics="true" pageId="0" pageNumber="195">Uphodato</emphasis>
<bibRefCitation id="97034B5F3602EA33FBFF150F0FB80446" author="Szwedo, J. &amp; Drohojowska, J. &amp; Popov, Yu. A. &amp; Simon, E. &amp; Wegierek, P." box="[1118,1269,1810,1834]" pageId="0" pageNumber="195" pagination="331 - 396" refId="ref1881" refString="Szwedo, J., Drohojowska, J., Popov, Yu. A., Simon, E. &amp; Wegierek, P. (2019) Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 110, 331 - 396. https: // doi. org / 10.1017 / S 175569101900001 X" type="journal article" year="2019">Szwedo, 2019</bibRefCitation>
as indicated by the basally concaved costal margin of forewing and peculiar venation of the wing discussed below.
</paragraph>
<paragraph id="F32D36AE3602EA33FF6615470ED104B1" blockId="0.[151,1437,1629,2014]" pageId="0" pageNumber="195">
A basal concavity in the costal margin of the forewing is a characteristic feature of the tribe Eucarpiini Emeljanov of
<taxonomicName id="34924D2D3602EA33FF3615630BBC04FA" authorityName="Spinola" authorityYear="1839" box="[151,241,1918,1942]" class="Insecta" family="Cixiidae" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="195" phylum="Arthropoda" rank="family">Cixiidae</taxonomicName>
and some
<taxonomicName id="34924D2D3602EA33FEC515630A8104FA" authorityName="Stal" authorityYear="1866" box="[356,460,1918,1942]" class="Insecta" family="Achilidae" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="195" phylum="Arthropoda" rank="family">Achilidae</taxonomicName>
(the genera
<taxonomicName id="34924D2D3602EA33FDF01563082B04FA" authority="Muir, 1924" authorityName="Muir" authorityYear="1924" box="[593,870,1918,1942]" class="Insecta" family="Achilidae" genus="Tropiphlepsia" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6EABC3602EA33FDF0156309A904FA" box="[593,740,1918,1942]" italics="true" pageId="0" pageNumber="195">Tropiphlepsia</emphasis>
Muir, 1924
</taxonomicName>
,
<taxonomicName id="34924D2D3602EA33FCD015630F2604FA" authority="Fowler, 1900" authorityName="Fowler" authorityYear="1900" box="[881,1131,1918,1942]" class="Insecta" family="Achilidae" genus="Apateson" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6EABC3602EA33FCD01563089904F9" box="[881,980,1918,1941]" italics="true" pageId="0" pageNumber="195">Apateson</emphasis>
Fowler, 1900
</taxonomicName>
, and
<taxonomicName id="34924D2D3602EA33FB0515630EDD04FA" authority="Muir, 1923" authorityName="Muir" authorityYear="1923" box="[1188,1424,1918,1942]" class="Insecta" family="Achilixiidae" genus="Achilixius" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6EABC3602EA33FB0515630E4204FA" box="[1188,1295,1918,1942]" italics="true" pageId="0" pageNumber="195">Achilixius</emphasis>
Muir, 1923
</taxonomicName>
), with their wings held steeply tectiform in repose. This concavity serves for optimal position of the hind legs before jumping.
<taxonomicName id="34924D2D3602EA33FF6615DB0A5C04B2" authorityName="Spinola" authorityYear="1839" box="[199,273,1990,2014]" class="Insecta" family="Issidae" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="195" phylum="Arthropoda" rank="family">Issidae</taxonomicName>
have no such concavity, as their legs are covered by forewings before jumping and are angled differently at
</paragraph>
<paragraph id="F32D36AE3603EA32FF361287091D020A" blockId="1.[151,1437,154,754]" pageId="1" pageNumber="196">
jumping (
<bibRefCitation id="97034B5F3603EA32FF5C1287097903DE" author="Anufriev, G. A. &amp; Emeljanov, A. F." box="[253,564,154,178]" pageId="1" pageNumber="196" pagination="12 - 495" refId="ref1568" refString="Anufriev, G. A. &amp; Emeljanov, A. F. (1988) Suborder Cicadinea (Auchenorrhyncha). In: Lehr, P. A. (Ed.), Opredelitel' nasekomykh Dal'nego Vostoka SSSR. Vol. 2. Nauka, Leningrad, pp. 12 - 495." type="book chapter" year="1988">Anufriev &amp; Emeljanov, 1988</bibRefCitation>
, fig. 11: 7). The forewing in
<taxonomicName id="34924D2D3603EA32FCFE128708E403DE" authorityName="Spinola" authorityYear="1839" box="[863,937,154,178]" class="Insecta" family="Issidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="family">Issidae</taxonomicName>
has the bulla in basal half under
<emphasis id="C1E6EABC3603EA32FAA112870E5503DB" box="[1280,1304,154,183]" italics="true" pageId="1" pageNumber="196">
R
<subScript id="6F1634EB3603EA32FAB112B40E5503DB" attach="left" box="[1296,1304,169,183]" fontSize="6" pageId="1" pageNumber="196">1</subScript>
</emphasis>
close to first furcation of the radius. Most issid species have convex forewings except, for example,
<taxonomicName id="34924D2D3603EA32FB8D12A30E6203BA" authority="Fennah, 1947" authorityName="Fennah" authorityYear="1947" box="[1068,1327,190,214]" class="Insecta" family="Issidae" genus="Oronoqua" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6EABC3603EA32FB8D12A30FD403B9" box="[1068,1177,190,213]" italics="true" pageId="1" pageNumber="196">Oronoqua</emphasis>
Fennah, 1947
</taxonomicName>
and
<taxonomicName id="34924D2D3603EA32FAC012A30A090395" authority="Walker, 1857" authorityName="Walker" authorityYear="1857" class="Insecta" family="Issidae" genus="Eupilis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6EABC3603EA32FAC012A30BFD0396" italics="true" pageId="1" pageNumber="196">Eupilis</emphasis>
Walker, 1857
</taxonomicName>
with flat wings; however, these genera have forewings with convex costal margins, without a concavity. The forewing imprint of
<taxonomicName id="34924D2D3603EA32FE3C131B086F0272" authority="Szwedo, 2019" authorityName="Szwedo" authorityYear="2019" box="[413,802,262,286]" class="Insecta" family="Cixiidae" genus="Uphodato" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="garwoterus">
<emphasis id="C1E6EABC3603EA32FE3C131B09CB0271" box="[413,646,262,286]" italics="true" pageId="1" pageNumber="196">Uphodato garwoterus</emphasis>
<bibRefCitation id="97034B5F3603EA32FD2D131B086F0272" author="Szwedo, J. &amp; Drohojowska, J. &amp; Popov, Yu. A. &amp; Simon, E. &amp; Wegierek, P." box="[652,802,262,286]" pageId="1" pageNumber="196" pagination="331 - 396" refId="ref1881" refString="Szwedo, J., Drohojowska, J., Popov, Yu. A., Simon, E. &amp; Wegierek, P. (2019) Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 110, 331 - 396. https: // doi. org / 10.1017 / S 175569101900001 X" type="journal article" year="2019">Szwedo, 2019</bibRefCitation>
</taxonomicName>
exhibits a distinct “cixiid clavus”, with
<emphasis id="C1E6EABC3603EA32FB69131B0FBC0271" box="[1224,1265,262,285]" italics="true" pageId="1" pageNumber="196">Pcu</emphasis>
+
<emphasis id="C1E6EABC3603EA32FAAD131B0E68024F" box="[1292,1317,262,291]" italics="true" pageId="1" pageNumber="196">
A
<subScript id="6F1634EB3603EA32FABC13080E68024F" attach="left" box="[1309,1317,277,291]" fontSize="6" pageId="1" pageNumber="196">1</subScript>
</emphasis>
joining the wing margin before the claval apex, and with tubercles on setal bases on the veins (
<figureCitation id="6BA92A2B3603EA32FC5D13370F75022E" box="[1020,1080,298,322]" captionStart="FIGURES 14" captionStartId="2.[151,258,1960,1982]" captionTargetBox="[315,1269,197,1917]" captionTargetId="figure@2.[302,1285,182,1933]" captionTargetPageId="2" captionText="FIGURES 14. Eucarpiini, forewing. 1. Uphodato garwoterus Szwedo; 2. U. vadimgratshevi (Szwedo); 3. U. phrikkosus (Szwedo); 4. Dilacreon orpheus (Fennah). 13 after Szwedo et al. (2019), modified, 4 after Fennah (1956)." figureDoi="http://doi.org/10.5281/zenodo.3697564" httpUri="https://zenodo.org/record/3697564/files/figure.png" pageId="1" pageNumber="196">Fig. 1</figureCitation>
) which are characteristic features of
<taxonomicName id="34924D2D3603EA32FF1513530A5C020A" authorityName="Spinola" authorityYear="1839" box="[180,273,334,358]" class="Insecta" family="Cixiidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="family">Cixiidae</taxonomicName>
, but not known for
<taxonomicName id="34924D2D3603EA32FE4013530901020A" authorityName="Stal" authorityYear="1866" box="[481,588,334,358]" class="Insecta" family="Achilidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="family">Achilidae</taxonomicName>
.
</paragraph>
<paragraph id="F32D36AE3603EA32FF66136F0FFD019E" blockId="1.[151,1437,154,754]" pageId="1" pageNumber="196">
<emphasis id="C1E6EABC3603EA32FF66136F0A7F02E6" box="[199,306,370,394]" italics="true" pageId="1" pageNumber="196">Uphodato</emphasis>
<bibRefCitation id="97034B5F3603EA32FE99136F0A8202E6" author="Szwedo, J. &amp; Drohojowska, J. &amp; Popov, Yu. A. &amp; Simon, E. &amp; Wegierek, P." box="[312,463,370,394]" pageId="1" pageNumber="196" pagination="331 - 396" refId="ref1881" refString="Szwedo, J., Drohojowska, J., Popov, Yu. A., Simon, E. &amp; Wegierek, P. (2019) Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 110, 331 - 396. https: // doi. org / 10.1017 / S 175569101900001 X" type="journal article" year="2019">Szwedo, 2019</bibRefCitation>
and
<emphasis id="C1E6EABC3603EA32FDA2136F09C702E6" box="[515,650,370,394]" italics="true" pageId="1" pageNumber="196">Breukoscelis</emphasis>
<bibRefCitation id="97034B5F3603EA32FD30136F086502E6" author="Szwedo, J. &amp; Drohojowska, J. &amp; Popov, Yu. A. &amp; Simon, E. &amp; Wegierek, P." box="[657,808,370,394]" pageId="1" pageNumber="196" pagination="331 - 396" refId="ref1881" refString="Szwedo, J., Drohojowska, J., Popov, Yu. A., Simon, E. &amp; Wegierek, P. (2019) Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 110, 331 - 396. https: // doi. org / 10.1017 / S 175569101900001 X" type="journal article" year="2019">Szwedo, 2019</bibRefCitation>
are distinguished by the stem
<emphasis id="C1E6EABC3603EA32FBCC136F0F3002E5" box="[1133,1149,370,393]" italics="true" pageId="1" pageNumber="196">R</emphasis>
+
<emphasis id="C1E6EABC3603EA32FB38136F0FE202E5" box="[1177,1199,370,393]" italics="true" pageId="1" pageNumber="196">M</emphasis>
on the forewing (
<figureCitation id="6BA92A2B3603EA32FACC136F0B8E02C2" captionStart="FIGURES 14" captionStartId="2.[151,258,1960,1982]" captionTargetBox="[315,1269,197,1917]" captionTargetId="figure@2.[302,1285,182,1933]" captionTargetPageId="2" captionText="FIGURES 14. Eucarpiini, forewing. 1. Uphodato garwoterus Szwedo; 2. U. vadimgratshevi (Szwedo); 3. U. phrikkosus (Szwedo); 4. Dilacreon orpheus (Fennah). 13 after Szwedo et al. (2019), modified, 4 after Fennah (1956)." figureDoi="http://doi.org/10.5281/zenodo.3697564" httpUri="https://zenodo.org/record/3697564/files/figure.png" pageId="1" pageNumber="196">Figs 1, 2</figureCitation>
). Such a condition is known for extant
<taxonomicName id="34924D2D3603EA32FDCA138B089202C1" authority="(Fennah, 1956)" baseAuthorityName="Fennah" baseAuthorityYear="1956" box="[619,991,405,430]" class="Insecta" family="Cixiidae" genus="Dilacreon" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="orpheus">
<emphasis id="C1E6EABC3603EA32FDCA138B087902C2" box="[619,820,406,430]" italics="true" pageId="1" pageNumber="196">Dilacreon orpheus</emphasis>
(
<bibRefCitation id="97034B5F3603EA32FCE3138B089A02C1" author="Fennah, R. G." box="[834,983,405,430]" pageId="1" pageNumber="196" pagination="39 - 211" refId="ref1623" refString="Fennah, R. G. (1956) Homoptera: Fulgoroidea. Insects of Micronesia, 6 (3), 39 - 211." type="journal article" year="1956">Fennah, 1956</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="6BA92A2B3603EA32FC4F138B0F7D02C2" box="[1006,1072,406,430]" captionStart="FIGURES 14" captionStartId="2.[151,258,1960,1982]" captionTargetBox="[315,1269,197,1917]" captionTargetId="figure@2.[302,1285,182,1933]" captionTargetPageId="2" captionText="FIGURES 14. Eucarpiini, forewing. 1. Uphodato garwoterus Szwedo; 2. U. vadimgratshevi (Szwedo); 3. U. phrikkosus (Szwedo); 4. Dilacreon orpheus (Fennah). 13 after Szwedo et al. (2019), modified, 4 after Fennah (1956)." figureDoi="http://doi.org/10.5281/zenodo.3697564" httpUri="https://zenodo.org/record/3697564/files/figure.png" pageId="1" pageNumber="196">Fig. 4</figureCitation>
), described from
<collectingCountry id="8B85763E3603EA32FB4C138B0ED002C2" box="[1261,1437,406,430]" name="Micronesia" pageId="1" pageNumber="196">Caroline Islands</collectingCountry>
(
<bibRefCitation id="97034B5F3603EA32FF3E13A70A7A02BD" author="Fennah, R. G." box="[159,311,441,466]" pageId="1" pageNumber="196" pagination="39 - 211" refId="ref1623" refString="Fennah, R. G. (1956) Homoptera: Fulgoroidea. Insects of Micronesia, 6 (3), 39 - 211." type="journal article" year="1956">Fennah, 1956</bibRefCitation>
), within the Eucarpiini.
<emphasis id="C1E6EABC3603EA32FDE113A709E602BE" box="[576,683,442,466]" italics="true" pageId="1" pageNumber="196">Uphodato</emphasis>
and
<emphasis id="C1E6EABC3603EA32FD4313A7082402BE" box="[738,873,442,466]" italics="true" pageId="1" pageNumber="196">Breukoscelis</emphasis>
are also characterized by a unique vein which may be treated as a transverse
<emphasis id="C1E6EABC3603EA32FE0B13C30AA00299" box="[426,493,478,501]" italics="true" pageId="1" pageNumber="196">m-cua</emphasis>
crossvein based on its position on the wing (
<figureCitation id="6BA92A2B3603EA32FC6F13C30F62029A" box="[974,1071,478,502]" captionStart="FIGURES 14" captionStartId="2.[151,258,1960,1982]" captionTargetBox="[315,1269,197,1917]" captionTargetId="figure@2.[302,1285,182,1933]" captionTargetPageId="2" captionText="FIGURES 14. Eucarpiini, forewing. 1. Uphodato garwoterus Szwedo; 2. U. vadimgratshevi (Szwedo); 3. U. phrikkosus (Szwedo); 4. Dilacreon orpheus (Fennah). 13 after Szwedo et al. (2019), modified, 4 after Fennah (1956)." figureDoi="http://doi.org/10.5281/zenodo.3697564" httpUri="https://zenodo.org/record/3697564/files/figure.png" pageId="1" pageNumber="196">Figs 13</figureCitation>
), but very likely it is the “root” of
<emphasis id="C1E6EABC3603EA32FF36101F0B8B0175" box="[151,198,514,537]" italics="true" pageId="1" pageNumber="196">CuA</emphasis>
, becoming a medio-cubital vein as a result of “interception” of the
<emphasis id="C1E6EABC3603EA32FC03101F089C0175" box="[930,977,514,537]" italics="true" pageId="1" pageNumber="196">CuA</emphasis>
by the
<emphasis id="C1E6EABC3603EA32FB85101F0F770175" box="[1060,1082,514,537]" italics="true" pageId="1" pageNumber="196">M</emphasis>
. In favor of this interpretation is the wing venation in members of the genus
<taxonomicName id="34924D2D3603EA32FDCA103B08230152" authority="Fennah, 1980" authorityName="Fennah" authorityYear="1980" box="[619,878,550,574]" class="Insecta" family="Cixiidae" genus="Dilacreon" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6EABC3603EA32FDCA103B099B0152" box="[619,726,550,574]" italics="true" pageId="1" pageNumber="196">Dilacreon</emphasis>
<bibRefCitation id="97034B5F3603EA32FD7C103B08230152" author="Fennah, R. G." box="[733,878,550,574]" pageId="1" pageNumber="196" pagination="237 - 328" refId="ref1649" refString="Fennah, R. G. (1980) The genus Bajauana and two allied new genera in New Guinea (Fulgoroidea: Cixiidae). Pacific Insects, 22 (3 - 4), 237 - 328." type="journal article" year="1980">Fennah, 1980</bibRefCitation>
</taxonomicName>
illustrated by
<bibRefCitation id="97034B5F3603EA32FBA9103B0FD30151" author="Fennah, R. G." box="[1032,1182,549,574]" pageId="1" pageNumber="196" pagination="39 - 211" refId="ref1623" refString="Fennah, R. G. (1956) Homoptera: Fulgoroidea. Insects of Micronesia, 6 (3), 39 - 211." type="journal article" year="1956">Fennah (1956</bibRefCitation>
,
<bibRefCitation id="97034B5F3603EA32FB09103B0FAC0152" author="Fennah, R. G." box="[1192,1249,550,574]" pageId="1" pageNumber="196" pagination="237 - 328" refId="ref1649" refString="Fennah, R. G. (1980) The genus Bajauana and two allied new genera in New Guinea (Fulgoroidea: Cixiidae). Pacific Insects, 22 (3 - 4), 237 - 328." type="journal article" year="1980">1980</bibRefCitation>
), in particular,
<taxonomicName id="34924D2D3603EA32FA23103B0BA0010E" baseAuthorityName="Fennah" baseAuthorityYear="1956" class="Insecta" family="Cixiidae" genus="Dilacreon" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="orpheus">
<emphasis id="C1E6EABC3603EA32FA23103B0BA0010E" italics="true" pageId="1" pageNumber="196">D. orpheus</emphasis>
</taxonomicName>
, which has a short stem
<emphasis id="C1E6EABC3603EA32FE5410570948010D" box="[501,517,586,609]" italics="true" pageId="1" pageNumber="196">R</emphasis>
+
<emphasis id="C1E6EABC3603EA32FD831057090D010D" box="[546,576,586,609]" italics="true" pageId="1" pageNumber="196">M.</emphasis>
However, on the drawings for
<emphasis id="C1E6EABC3603EA32FC2D105708BA010E" box="[908,1015,586,610]" italics="true" pageId="1" pageNumber="196">Uphodato</emphasis>
and
<emphasis id="C1E6EABC3603EA32FB8A10570FFF010E" box="[1067,1202,586,610]" italics="true" pageId="1" pageNumber="196">Breukoscelis</emphasis>
(
<bibRefCitation id="97034B5F3603EA32FB6110570ED5010E" author="Szwedo, J. &amp; Drohojowska, J. &amp; Popov, Yu. A. &amp; Simon, E. &amp; Wegierek, P." box="[1216,1432,586,610]" pageId="1" pageNumber="196" pagination="331 - 396" refId="ref1881" refString="Szwedo, J., Drohojowska, J., Popov, Yu. A., Simon, E. &amp; Wegierek, P. (2019) Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 110, 331 - 396. https: // doi. org / 10.1017 / S 175569101900001 X" type="journal article" year="2019">
Szwedo
<emphasis id="C1E6EABC3603EA32FABC10570E01010E" box="[1309,1356,586,610]" italics="true" pageId="1" pageNumber="196">et al</emphasis>
., 2019
</bibRefCitation>
, figs 4648)
<emphasis id="C1E6EABC3603EA32FEB610730A0B01E9" box="[279,326,622,645]" italics="true" pageId="1" pageNumber="196">CuA</emphasis>
is not furcating, which is not typical for
<taxonomicName id="34924D2D3603EA32FD551073081F01EA" authorityName="Spinola" authorityYear="1839" box="[756,850,622,646]" class="Insecta" family="Cixiidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="family">Cixiidae</taxonomicName>
. If our interpretation of vein “interception” is true, then
<emphasis id="C1E6EABC3603EA32FF36108F0B8B01C5" box="[151,198,658,681]" italics="true" pageId="1" pageNumber="196">CuA</emphasis>
acquires the usual character of furcating. Thus, the species of the genus
<taxonomicName id="34924D2D3603EA32FC77108F0F0C01C6" authorityName="Fennah" authorityYear="1980" box="[982,1089,658,682]" class="Insecta" family="Cixiidae" genus="Dilacreon" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="C1E6EABC3603EA32FC77108F0F0C01C6" box="[982,1089,658,682]" italics="true" pageId="1" pageNumber="196">Dilacreon</emphasis>
</taxonomicName>
have the basal part of
<emphasis id="C1E6EABC3603EA32FA96108F0E2201C3" box="[1335,1391,658,687]" italics="true" pageId="1" pageNumber="196">
CuA
<subScript id="6F1634EB3603EA32FAC610BC0E2201C3" attach="left" box="[1383,1391,673,687]" fontSize="6" pageId="1" pageNumber="196">1</subScript>
</emphasis>
and prenodal vein
<emphasis id="C1E6EABC3603EA32FE9110AB0A3E01A1" box="[304,371,694,717]" italics="true" pageId="1" pageNumber="196">m-cua</emphasis>
situated symmetrically as though ready for “interception”. The extravenal pterostigma is not visible on the imprints of fossil taxa; however, within extant Eucarpiini it is small or completely obsolete.
</paragraph>
</subSubSection>
</treatment>
</document>