289 lines
34 KiB
XML
289 lines
34 KiB
XML
<document id="B73C2E4BAE81AC8C1B8FEDF1BB306929" ID-DOI="10.1093/zoolinnean/zlad111" ID-ISSN="0024-4082" ID-PMC="PMC10983082" ID-PubMed="38566915" ID-Zenodo-Dep="11240925" ID-ZooBank="E8CC81A3-E625-4C48-B783-29AA9BFC83C3C" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1716363317501" checkinUser="plazi" docAuthor="Moreno, Karina, Rico, Diane M., Middlebrooks, Michael, Medrano, Sabrina, Valdés, Ángel A. & Krug, Patrick J." docDate="2024" docId="03C287FBFFA6FFC0FF8A4636FED92235" docLanguage="en" docName="zlad111.pdf" docOrigin="Zoological Journal of the Linnean Society 200 (4)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad111" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Cyerce willetteorum Moreno & Rico & Middlebrooks & Medrano & Valdés & Krug 2024, sp.nov." docType="treatment" docVersion="3" lastPageNumber="970" masterDocId="FFFBFF83FFBAFFDEFC6F452FFFAF206B" masterDocTitle="A cryptic radiation of Caribbean sea slugs revealed by integrative analysis: Cyerce ‘ antillensis’ (Sacoglossa: Caliphyllidae) is six distinct species" masterLastPageNumber="979" masterPageNumber="940" pageNumber="968" updateTime="1716496456803" updateUser="carolina">
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<mods:title id="3868758FDB81E574A8BD0A7B09686810">A cryptic radiation of Caribbean sea slugs revealed by integrative analysis: Cyerce ‘ antillensis’ (Sacoglossa: Caliphyllidae) is six distinct species</mods:title>
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<mods:namePart id="507343F07C6F4C723D3E39B6AB714432">Moreno, Karina</mods:namePart>
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<mods:affiliation id="0D76EDB3807DA46A778B4BA07BF52772">Department of Biological Sciences, California State Polytechnic University Pomona, Pomona, CA 91768, USA</mods:affiliation>
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<mods:namePart id="45D4CE26F1385C8E3079142867CD8644">Rico, Diane M.</mods:namePart>
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<mods:affiliation id="A025480C6FCD33B5968FCEDD63085B34">Department of Biological Sciences, California State University, Los Angeles, CA 90032 - 8201, USA</mods:affiliation>
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<mods:namePart id="53024053DE03A8B660E9AFDF2C95AAEE">Middlebrooks, Michael</mods:namePart>
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<mods:affiliation id="8A94FC266702E7B9BAC7E5AF928B6E22">Department of Biology, University of Tampa, Tampa, FL 33609, USA</mods:affiliation>
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<mods:namePart id="5D042E05B35DD2DF9E7ED6D7C9F30499">Medrano, Sabrina</mods:namePart>
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<mods:affiliation id="D1DC29F3E6A8C275407ED9650E82F20D">Department of Biological Sciences, California State Polytechnic University Pomona, Pomona, CA 91768, USA</mods:affiliation>
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<mods:namePart id="EF060DFD7C47082B4BC886718673226F">Valdés, Ángel A.</mods:namePart>
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<mods:affiliation id="63B852E11D8EEEBE922C54C96E4D9A3E">Department of Biological Sciences, California State Polytechnic University Pomona, Pomona, CA 91768, USA</mods:affiliation>
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<mods:namePart id="883E8B02396E3212FC0765955ADB668A">Krug, Patrick J.</mods:namePart>
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<mods:affiliation id="40318A0D4E77AB901AE35566407ACE50">Department of Biological Sciences, California State University, Los Angeles, CA 90032 - 8201, USA</mods:affiliation>
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<mods:nameIdentifier id="689D1B9A686A7819F1086BE9C9A20B7A" type="email">pkrug@calstatela.edu</mods:nameIdentifier>
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<mods:title id="852818E19DF1C82E364EFB4045F2F8D7">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="DF2BC5573BBDFD208A7651D23E5034F7">2024</mods:date>
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<treatment id="03C287FBFFA6FFC0FF8A4636FED92235" LSID="urn:lsid:plazi:treatment:03C287FBFFA6FFC0FF8A4636FED92235" httpUri="http://treatment.plazi.org/id/03C287FBFFA6FFC0FF8A4636FED92235" lastPageId="30" lastPageNumber="970" pageId="28" pageNumber="968">
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<heading id="D09C8181FFA6FFC2FF8A4636FB572358" bold="true" box="[997,1272,792,819]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="968" reason="2">
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<emphasis id="B91FEAFFFFA6FFC2FF8A4636FB572358" bold="true" box="[997,1272,792,819]" pageId="28" pageNumber="968">
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<taxonomicName id="4C6B4D6EFFA6FFC2FF8A4636FB072358" ID-CoL="C4SJS" authority="Moreno & Rico & Middlebrooks & Medrano & Valdés & Krug, 2024" authorityName="Moreno & Rico & Middlebrooks & Medrano & Valdés & Krug" authorityYear="2024" box="[997,1192,792,819]" class="Gastropoda" family="Hermaeidae" genus="Cyerce" kingdom="Animalia" order="Sacoglossa" pageId="28" pageNumber="968" phylum="Mollusca" rank="species" species="willetteorum" status="sp. nov.">
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<emphasis id="B91FEAFFFFA6FFC2FF8A4636FB072358" bold="true" box="[997,1192,792,819]" italics="true" pageId="28" pageNumber="968">Cyerce willetteorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22C5784FFA6FFC2F8C34636FB572358" box="[1196,1272,793,819]" pageId="28" pageNumber="968" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C3716566FFA6FFC0F8694667FED92235" lastPageId="30" lastPageNumber="970" pageId="28" pageNumber="968" type="description">
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<paragraph id="8BD436EDFFA6FFC2F8694667FB782309" blockId="28.[1030,1239,840,866]" box="[1030,1239,840,866]" pageId="28" pageNumber="968">
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<emphasis id="B91FEAFFFFA6FFC2F8694667FB782309" bold="true" box="[1030,1239,840,866]" pageId="28" pageNumber="968">
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(
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<figureCitation id="13502A68FFA6FFC2F87E4667FBC52309" box="[1041,1130,840,866]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[130,195,1841,1865]" captionStartId-1="8.[113,178,1844,1868]" captionStartId-2="9.[131,196,1848,1872]" captionTargetBox-0="[129,1473,145,1813]" captionTargetBox-1="[400,1174,149,1814]" captionTargetBox-2="[144,754,1402,1824]" captionTargetId-0="figure-64@3.[128,1475,144,1814]" captionTargetId-1="figure-7@8.[399,1174,148,1815]" captionTargetId-2="figure-918@9.[143,754,1401,1825]" captionTargetPageId-0="3" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="Figure 1. Western Atlantic sampling sites; for more details, see Table 1. Top panel shows southern Florida peninsula and eastern Caribbean; bottom panel shows expanded view of inset box." captionText-1="Figure 2. Consensus phylogenetic tree for Cyerce and Sohgenia based on maximum likelihood (ML) analysis of three concatenated loci. Percentages corresponding to significant bootstrap support are given below branches, and posterior probabilities from Bayesian inference (BI) analysis are shown in bold above branches. Capital letters label three key supported nodes. Haplotypes are coded by date–location–specimen number on terminals. Results of four species delimitation analyses are indicated by vertical bars; horizontal shaded bars enclose specimens grouped as species entities by all methods." captionText-2="Figure 3. Histogram of pairwise Tamura–Nei (TrN)-corrected COI distances, showing barcoding gap; white bars denote intraspecific distances and black bars interspecific distances, based on automated barcode gap discovery (ABGD) and assemble species by automatic partitioning (ASAP) analyses." figureDoi-0="http://doi.org/10.5281/zenodo.11240927" figureDoi-1="http://doi.org/10.5281/zenodo.11240929" figureDoi-2="http://doi.org/10.5281/zenodo.11240931" httpUri-0="https://zenodo.org/record/11240927/files/figure.png" httpUri-1="https://zenodo.org/record/11240929/files/figure.png" httpUri-2="https://zenodo.org/record/11240931/files/figure.png" pageId="28" pageNumber="968">Figs 1–3</figureCitation>
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,
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<figureCitation id="13502A68FFA6FFC2F81A4667FB2C2309" box="[1141,1155,840,866]" captionStart="Figure 5" captionStartId="13.[129,194,1102,1126]" captionTargetBox="[132,1471,147,1073]" captionTargetId="figure-464@13.[129,1473,144,1075]" captionTargetPageId="13" captionText="Figure 5. Haplotype network of phased histone 3 alleles colour-coded by candidate species, with the size of each circle denoting the frequency of the sampled allele. A small black dot indicates an unsampled allele. Each bar crossing a line denotes one nucleotide substitution." figureDoi="http://doi.org/10.5281/zenodo.11240935" httpUri="https://zenodo.org/record/11240935/files/figure.png" pageId="28" pageNumber="968">5</figureCitation>
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,
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<figureCitation id="13502A68FFA6FFC2F8E34667FB082309" box="[1164,1191,840,866]" captionStart="Figure 8" captionStartId="16.[115,180,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-341@16.[114,1458,144,1176]" captionTargetPageId="16" captionText="Figure 8. Stained and mounted penial stylets of Cyerce species from the Western Atlantic. Black arrow indicates stylet.A, Cyerce nicholasi, isolate 10Bim03. B, Cyerce piercei, isolate KM66. C, Cyerce ellingsonorum, isolate 06Jam05. D, Cyerce antillensis, isolate 10Bim01.E, Cyerce browneveorum, isolate 09Gei01.F, Cyerce willetteorum, isolate 07Stir01.G, Cyerce cf. cristallina, isolate B101." figureDoi="http://doi.org/10.5281/zenodo.11240943" httpUri="https://zenodo.org/record/11240943/files/figure.png" pageId="28" pageNumber="968">8F</figureCitation>
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,
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<figureCitation id="13502A68FFA6FFC2F8DE4667FB622309" box="[1201,1229,840,866]" captionStart="Figure 17" captionStartId="31.[129,194,1546,1570]" captionTargetBox="[129,1473,144,1519]" captionTargetId="figure-249@31.[129,1473,144,1519]" captionTargetPageId="31" captionText="Figure 17. Cyerce willetteorum, radular scanning electron micrographs of LACM 3844 (isolate 07Stir01). A, complete radula, with ascus. B, close-up view of tissue-free teeth from descending limb, showing irregularly shaped denticles.C, close-up view of ascus." figureDoi="http://doi.org/10.5281/zenodo.11240963" httpUri="https://zenodo.org/record/11240963/files/figure.png" pageId="28" pageNumber="968">17</figureCitation>
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)
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</emphasis>
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<paragraph id="8BD436EDFFA6FFC2FF4546A6FAFD23AB" blockId="28.[810,1460,905,960]" pageId="28" pageNumber="968">
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<emphasis id="B91FEAFFFFA6FFC2FF4546A6FB3E23CB" box="[810,1169,905,929]" italics="true" pageId="28" pageNumber="968">ZooBank registration:</emphasis>
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lsid:zoobank. org:act:
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<uuid id="FFCD0C38FFA6FFC2FF1C4687FAFD23AB" box="[883,1362,936,960]" pageId="28" pageNumber="968">B2309FF2-4BF7-4317-BE5F-5398AB298F30</uuid>
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</paragraph>
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<paragraph id="8BD436EDFFA6FFC2F84146CFFB002392" blockId="28.[1070,1199,991,1017]" box="[1070,1199,991,1017]" pageId="28" pageNumber="968">
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<emphasis id="B91FEAFFFFA6FFC2F84146CFFB002392" box="[1070,1199,991,1017]" italics="true" pageId="28" pageNumber="968">Type material</emphasis>
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</paragraph>
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<paragraph id="8BD436EDFFA6FFC2FF454129FB172456" blockId="28.[810,1459,1030,1085]" pageId="28" pageNumber="968">
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<materialsCitation id="3B033CB0FFA6FFC2FF454129FB172456" collectingDate="2001-07" collectionCode="LACM" country="Bahamas" location="Stirrup Cay" pageId="28" pageNumber="968" specimenCode="LACM 3844" specimenCount="1" typeStatus="holotype">
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<emphasis id="B91FEAFFFFA6FFC2FF454129FC252475" box="[810,906,1030,1054]" italics="true" pageId="28" pageNumber="968">
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<typeStatus id="54D0884FFFA6FFC2FF454129FC2A2475" box="[810,901,1030,1054]" pageId="28" pageNumber="968" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<location id="8EB46036FFA6FFC2FFFA4129FBBE2475" LSID="urn:lsid:plazi:treatment:03C287FBFFA6FFC0FF8A4636FED92235:8EB46036FFA6FFC2FFFA4129FBBE2475" box="[917,1041,1030,1054]" country="Bahamas" name="Stirrup Cay" pageId="28" pageNumber="968">Stirrup Cay</location>
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,
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||
<collectingCountry id="F37C767DFFA6FFC2F84F4129FBD12475" box="[1056,1150,1030,1054]" name="Bahamas" pageId="28" pageNumber="968">Bahamas</collectingCountry>
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,
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||
<date id="FFD5102DFFA6FFC2F8E34129FB5A2475" box="[1164,1269,1030,1054]" pageId="28" pageNumber="968" value="2001-07">
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<collectingDate id="EF91E9C5FFA6FFC2F8E34129FB5A2475" box="[1164,1269,1030,1054]" pageId="28" pageNumber="968" value="2001-07">July 2001</collectingDate>
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</date>
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,
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<quantity id="4C939B08FFA6FFC2F96C4129FAEB2476" box="[1283,1348,1030,1054]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="968" unit="mm" value="2.0">2 mm</quantity>
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preserved length (
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<emphasis id="B91FEAFFFFA6FFC2FF144109FC542456" box="[891,1019,1061,1085]" italics="true" pageId="28" pageNumber="968">
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<specimenCode id="DBCD9E96FFA6FFC2FF144109FC542456" box="[891,1019,1061,1085]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="28" pageNumber="968" type="Museum">LACM 3844</specimenCode>
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||
</emphasis>
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; isolate 07Stir01).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD436EDFFA6FFC2F83E4171FB22241C" blockId="28.[810,1262,1118,1179]" box="[1105,1165,1118,1143]" pageId="28" pageNumber="968">
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<emphasis id="B91FEAFFFFA6FFC2F83E4171FB22241C" box="[1105,1165,1118,1143]" italics="true" pageId="28" pageNumber="968">Range</emphasis>
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</paragraph>
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<paragraph id="8BD436EDFFA6FFC2FF4541ACFB4124F0" blockId="28.[810,1262,1118,1179]" box="[810,1262,1155,1179]" pageId="28" pageNumber="968">
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||
Great Stirrup Cay,
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<collectingCountry id="F37C767DFFA6FFC2FF8741ACFBEB24F0" box="[1000,1092,1155,1179]" name="Bahamas" pageId="28" pageNumber="968">Bahamas</collectingCountry>
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(present study).
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</paragraph>
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<paragraph id="8BD436EDFFA6FFC2F8574193FB0B24BE" blockId="28.[810,1459,1212,1336]" box="[1080,1188,1212,1237]" pageId="28" pageNumber="968">
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<emphasis id="B91FEAFFFFA6FFC2F8574193FB0B24BE" box="[1080,1188,1212,1237]" italics="true" pageId="28" pageNumber="968">Description</emphasis>
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</paragraph>
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<paragraph id="8BD436EDFFA6FFC2FF4541CEFC0D2553" blockId="28.[810,1459,1212,1336]" pageId="28" pageNumber="968">
|
||
<emphasis id="B91FEAFFFFA6FFC2FF4541CEFBAE2492" box="[810,1025,1249,1273]" italics="true" pageId="28" pageNumber="968">External morphology:</emphasis>
|
||
No photograph was taken of the live specimen, hence no description of external morphology is possible at this time.
|
||
</paragraph>
|
||
<paragraph id="8BD436EDFFA6FFC2FF454076FC0E2607" blockId="28.[810,1460,1369,1707]" pageId="28" pageNumber="968">
|
||
<emphasis id="B91FEAFFFFA6FFC2FF454076FBA9251A" box="[810,1030,1369,1393]" italics="true" pageId="28" pageNumber="968">Internal morphology:</emphasis>
|
||
Radula of
|
||
<typeStatus id="54D0884FFFA6FFC2F8EE4076FB4E251A" box="[1153,1249,1369,1393]" pageId="28" pageNumber="968" type="holotype">holotype</typeStatus>
|
||
(
|
||
<emphasis id="B91FEAFFFFA6FFC2F8994075FAD2251A" box="[1270,1405,1369,1393]" italics="true" pageId="28" pageNumber="968">LACM 3844</emphasis>
|
||
, isolate 07Stir01) with six teeth on ascending limb and seven teeth on descending limb (
|
||
<figureCitation id="13502A68FFA6FFC2F83F40B7FB0325DB" box="[1104,1196,1432,1456]" captionStart="Figure 17" captionStartId="31.[129,194,1546,1570]" captionTargetBox="[129,1473,144,1519]" captionTargetId="figure-249@31.[129,1473,144,1519]" captionTargetPageId="31" captionText="Figure 17. Cyerce willetteorum, radular scanning electron micrographs of LACM 3844 (isolate 07Stir01). A, complete radula, with ascus. B, close-up view of tissue-free teeth from descending limb, showing irregularly shaped denticles.C, close-up view of ascus." figureDoi="http://doi.org/10.5281/zenodo.11240963" httpUri="https://zenodo.org/record/11240963/files/figure.png" pageId="28" pageNumber="968">Fig. 17A</figureCitation>
|
||
). Teeth shallow, bent at an angle of 18°, tapering to pointed tip; 97 µm in length (
|
||
<figureCitation id="13502A68FFA6FFC2F9E44098FCFB2584" captionStart="Figure 17" captionStartId="31.[129,194,1546,1570]" captionTargetBox="[129,1473,144,1519]" captionTargetId="figure-249@31.[129,1473,144,1519]" captionTargetPageId="31" captionText="Figure 17. Cyerce willetteorum, radular scanning electron micrographs of LACM 3844 (isolate 07Stir01). A, complete radula, with ascus. B, close-up view of tissue-free teeth from descending limb, showing irregularly shaped denticles.C, close-up view of ascus." figureDoi="http://doi.org/10.5281/zenodo.11240963" httpUri="https://zenodo.org/record/11240963/files/figure.png" pageId="28" pageNumber="968">Fig. 17B</figureCitation>
|
||
). Row of nine denticles along each cutting edge, irregular in shape and width; narrow at base, widening to a curved edge, ≤ 2.5 μm wide. Denticles narrower, more rectangular near tip (
|
||
<figureCitation id="13502A68FFA6FFC2FFFE431AFC452626" box="[913,1002,1589,1613]" captionStart="Figure 17" captionStartId="31.[129,194,1546,1570]" captionTargetBox="[129,1473,144,1519]" captionTargetId="figure-249@31.[129,1473,144,1519]" captionTargetPageId="31" captionText="Figure 17. Cyerce willetteorum, radular scanning electron micrographs of LACM 3844 (isolate 07Stir01). A, complete radula, with ascus. B, close-up view of tissue-free teeth from descending limb, showing irregularly shaped denticles.C, close-up view of ascus." figureDoi="http://doi.org/10.5281/zenodo.11240963" httpUri="https://zenodo.org/record/11240963/files/figure.png" pageId="28" pageNumber="968">Fig. 17B</figureCitation>
|
||
). Partial ascus with four pre-radular teeth (
|
||
<figureCitation id="13502A68FFA6FFC2FF5A437BFC3F2607" box="[821,912,1620,1644]" captionStart="Figure 17" captionStartId="31.[129,194,1546,1570]" captionTargetBox="[129,1473,144,1519]" captionTargetId="figure-249@31.[129,1473,144,1519]" captionTargetPageId="31" captionText="Figure 17. Cyerce willetteorum, radular scanning electron micrographs of LACM 3844 (isolate 07Stir01). A, complete radula, with ascus. B, close-up view of tissue-free teeth from descending limb, showing irregularly shaped denticles.C, close-up view of ascus." figureDoi="http://doi.org/10.5281/zenodo.11240963" httpUri="https://zenodo.org/record/11240963/files/figure.png" pageId="28" pageNumber="968">Fig. 17C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD436EDFFA6FFC2FF2A435CFA3526C0" blockId="28.[810,1460,1369,1707]" pageId="28" pageNumber="968">
|
||
Penis with curved, wide stylet, narrowing gradually from embedded base, 120 µm long; oval opening at wide tip (
|
||
<figureCitation id="13502A68FFA6FFC2F92B43BCFA2526C0" box="[1348,1418,1683,1707]" captionStart="Figure 8" captionStartId="16.[115,180,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-341@16.[114,1458,144,1176]" captionTargetPageId="16" captionText="Figure 8. Stained and mounted penial stylets of Cyerce species from the Western Atlantic. Black arrow indicates stylet.A, Cyerce nicholasi, isolate 10Bim03. B, Cyerce piercei, isolate KM66. C, Cyerce ellingsonorum, isolate 06Jam05. D, Cyerce antillensis, isolate 10Bim01.E, Cyerce browneveorum, isolate 09Gei01.F, Cyerce willetteorum, isolate 07Stir01.G, Cyerce cf. cristallina, isolate B101." figureDoi="http://doi.org/10.5281/zenodo.11240943" httpUri="https://zenodo.org/record/11240943/files/figure.png" pageId="28" pageNumber="968">Fig. 8F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD436EDFFA6FFC2F82543E5FB3B268F" blockId="28.[1098,1172,1738,1764]" box="[1098,1172,1738,1764]" pageId="28" pageNumber="968">
|
||
<emphasis id="B91FEAFFFFA6FFC2F82543E5FB3B268F" box="[1098,1172,1738,1764]" italics="true" pageId="28" pageNumber="968">Ecology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD436EDFFA6FFC2FF4543DEFBE62763" blockId="28.[810,1097,1777,1801]" box="[810,1097,1777,1801]" pageId="28" pageNumber="968">
|
||
Collected from
|
||
<emphasis id="B91FEAFFFFA6FFC2FFA543DEFBEC2763" box="[970,1091,1777,1800]" italics="true" pageId="28" pageNumber="968">
|
||
Pe.
|
||
<taxonomicName id="4C6B4D6EFFA6FFC2FF8443DEFBEC2763" box="[1003,1091,1777,1800]" class="Bivalvia" family="Penicillidae" genus="Penicillus" kingdom="Animalia" pageId="28" pageNumber="968" phylum="Mollusca" rank="species" species="capitatus">capitatus</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD436EDFFA6FFC2F8534207FB0D2729" blockId="28.[1084,1186,1832,1858]" box="[1084,1186,1832,1858]" pageId="28" pageNumber="968">
|
||
<emphasis id="B91FEAFFFFA6FFC2F8534207FB0D2729" box="[1084,1186,1832,1858]" italics="true" pageId="28" pageNumber="968">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD436EDFFA6FFC2FF454260FB6D27AE" blockId="28.[810,1459,1871,1989]" pageId="28" pageNumber="968">Named in joint honour of Dr Demian Willette, who contributed specimens to this work and helped P.J.K. to initiate studies of sacoglossan biodiversity, and his brother Darian Willette, a devoted naturalist lost at too young an age.</paragraph>
|
||
<caption id="DF146665FFA7FFC3FCEE43B2FB3826BA" ID-DOI="http://doi.org/10.5281/zenodo.11240961" ID-Zenodo-Dep="11240961" httpUri="https://zenodo.org/record/11240961/files/figure.png" pageId="29" pageNumber="969" startId="29.[129,194,1693,1717]" targetBox="[129,1473,144,1665]" targetPageId="29" targetType="figure">
|
||
<paragraph id="8BD436EDFFA7FFC3FCEE43B2FB3826BA" blockId="29.[129,1463,1693,1745]" pageId="29" pageNumber="969">
|
||
<emphasis id="B91FEAFFFFA7FFC3FCEE43B2FE1726DE" bold="true" box="[129,440,1693,1717]" pageId="29" pageNumber="969">
|
||
Figure 16.
|
||
<taxonomicName id="4C6B4D6EFFA7FFC3FC8543B2FE1726DE" authorityName="Moreno & Rico & Middlebrooks & Medrano & Valdés & Krug" authorityYear="2024" box="[234,440,1693,1717]" class="Gastropoda" family="Hermaeidae" genus="Cyerce" kingdom="Animalia" pageId="29" pageNumber="969" phylum="Mollusca" rank="species" species="ellingsonorum">
|
||
<emphasis id="B91FEAFFFFA7FFC3FC8543B2FE1726DE" bold="true" box="[234,440,1693,1717]" italics="true" pageId="29" pageNumber="969">Cyerce ellingsonorum</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
, radular scanning electron micrographs of
|
||
<emphasis id="B91FEAFFFFA7FFC3FF2743B1FC1126DE" box="[840,958,1693,1717]" italics="true" pageId="29" pageNumber="969">LACM 3841</emphasis>
|
||
(isolate 06JAM09). A, complete radula with ascus. B, close-up view of tissue-free teeth from ascending limb, showing rectangular denticles. C, close-up view of ascus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD436EDFFA7FFC3FDF4422EFE402770" blockId="29.[411,495,1793,1819]" box="[411,495,1793,1819]" pageId="29" pageNumber="969">
|
||
<emphasis id="B91FEAFFFFA7FFC3FDF4422EFE402770" box="[411,495,1793,1819]" italics="true" pageId="29" pageNumber="969">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD436EDFFA7FFC0FCEE4207FDAD208D" blockId="29.[129,778,1832,1981]" lastBlockId="30.[113,764,144,606]" lastPageId="30" lastPageNumber="970" pageId="29" pageNumber="969">
|
||
Only a single exemplar of
|
||
<taxonomicName id="4C6B4D6EFFA7FFC3FDE04207FD8A272B" authorityName="Moreno & Rico & Middlebrooks & Medrano & Valdés & Krug" authorityYear="2024" box="[399,549,1832,1856]" class="Gastropoda" family="Hermaeidae" genus="Cyerce" kingdom="Animalia" pageId="29" pageNumber="969" phylum="Mollusca" rank="species" species="willetteorum">
|
||
<emphasis id="B91FEAFFFFA7FFC3FDE04207FD8A272B" box="[399,549,1832,1856]" italics="true" pageId="29" pageNumber="969">C. willetteorum</emphasis>
|
||
</taxonomicName>
|
||
was sampled over the 15-year duration of this study of Western Atlantic sacoglossans. However, the lone specimen was supported as a distinct species by all methods of molecular species delimitation. Although the external morphology could not be described at this time, the penial stylet was distinct from the other complex members, less curved and shorter than
|
||
<taxonomicName id="4C6B4D6EFFA7FFC3F85D420EFB0D2753" authorityName="Moreno & Rico & Middlebrooks & Medrano & Valdés & Krug" authorityYear="2024" box="[1074,1186,1824,1848]" class="Gastropoda" family="Hermaeidae" genus="Cyerce" kingdom="Animalia" pageId="29" pageNumber="969" phylum="Mollusca" rank="species" species="nicholasi">
|
||
<emphasis id="B91FEAFFFFA7FFC3F85D420EFB0D2753" box="[1074,1186,1824,1848]" italics="true" pageId="29" pageNumber="969">C. nicholasi</emphasis>
|
||
</taxonomicName>
|
||
but more curved and longer than any other newly described species. The radular denticles were also distinctive, with an irregular shape and curved edge not observed in any other species. Given the genetic support for the
|
||
<typeStatus id="54D0884FFFA7FFC3FF5642B1FC3A27DD" box="[825,917,1950,1974]" pageId="29" pageNumber="969" type="holotype">holotype</typeStatus>
|
||
as a separate species and the distinctive characteristics of the internal anatomy, we elected to name the species to clarify the diversity of the Caribbean
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FDC9459FFDBA20AC" box="[422,533,176,199]" class="Gastropoda" family="Hermaeidae" genus="Cyerce" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B91FEAFFFFA4FFC0FDC9459FFE4920AC" box="[422,486,176,199]" italics="true" pageId="30" pageNumber="970">Cyerce</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
and to facilitate future studies of the biology of this rare taxon.
|
||
</paragraph>
|
||
<paragraph id="8BD436EDFFA4FFC0FCE245C1FED92235" blockId="30.[113,764,144,606]" pageId="30" pageNumber="970">
|
||
This was the only species collected from
|
||
<emphasis id="B91FEAFFFFA4FFC0FE7145C0FD39216D" box="[542,662,239,262]" italics="true" pageId="30" pageNumber="970">
|
||
Pe.
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FE5145C0FD39216D" box="[574,662,239,262]" class="Bivalvia" family="Penicillidae" genus="Penicillus" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="capitatus">capitatus</taxonomicName>
|
||
</emphasis>
|
||
, although
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FC1E4421FEA1214E" authorityName="Trinchese" authorityYear="1881" box="[113,270,269,293]" class="Gastropoda" family="Hermaeidae" genus="Cyerce" isUncertain="true" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="cristallina">
|
||
<emphasis id="B91FEAFFFFA4FFC0FC1E4421FF27214E" box="[113,136,270,293]" italics="true" pageId="30" pageNumber="970">C.</emphasis>
|
||
cf.
|
||
<emphasis id="B91FEAFFFFA4FFC0FCC14422FEA1214E" box="[174,270,269,293]" italics="true" pageId="30" pageNumber="970">cristallina</emphasis>
|
||
</taxonomicName>
|
||
associates transiently with
|
||
<emphasis id="B91FEAFFFFA4FFC0FE474421FD0B214E" box="[552,676,270,293]" italics="true" pageId="30" pageNumber="970">
|
||
Pe.
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FE234421FD0B214E" box="[588,676,270,293]" class="Bivalvia" family="Penicillidae" genus="Penicillus" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="capitatus">capitatus</taxonomicName>
|
||
</emphasis>
|
||
at night (M. Charteris, unpublished data). Algal congeners
|
||
<emphasis id="B91FEAFFFFA4FFC0FE1B4402FD55212E" box="[628,762,301,325]" italics="true" pageId="30" pageNumber="970">
|
||
Pe.
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FEFA4402FD55212E" authorityName="Blainville" authorityYear="1830" baseAuthorityName="J. V. Lamouroux" box="[661,762,301,325]" class="Bivalvia" family="Penicillidae" genus="Penicillus" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="dumetosus">dumetosus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B91FEAFFFFA4FFC0FCCF4462FE93210F" box="[160,316,332,356]" italics="true" pageId="30" pageNumber="970">
|
||
Pe.
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FCAA4463FE93210F" authorityName="Decaisne" authorityYear="1842" box="[197,316,332,356]" class="Bivalvia" family="Penicillidae" genus="Penicillus" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="lamourouxii">lamourouxii</taxonomicName>
|
||
</emphasis>
|
||
were used as hosts by four other complex members and have thicker filaments than
|
||
<emphasis id="B91FEAFFFFA4FFC0FE404443FD0321E8" box="[559,684,364,387]" italics="true" pageId="30" pageNumber="970">
|
||
Pe.
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FE3B4443FD0321E8" box="[596,684,364,387]" class="Bivalvia" family="Penicillidae" genus="Penicillus" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="capitatus">capitatus</taxonomicName>
|
||
</emphasis>
|
||
on the ‘shaving brush’ portion of the algal thallus. However, extensive collections of
|
||
<emphasis id="B91FEAFFFFA4FFC0FD664484FE2721A9" box="[265,392,427,450]" italics="true" pageId="30" pageNumber="970">
|
||
Pe.
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FD5F4484FE2721A9" box="[304,392,427,450]" class="Bivalvia" family="Penicillidae" genus="Penicillus" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="capitatus">capitatus</taxonomicName>
|
||
</emphasis>
|
||
made elsewhere in the Caribbean did not yield specimens of
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FDEA44E5FE6A218A" authorityName="Bergh" authorityYear="1871" box="[389,453,458,481]" class="Gastropoda" family="Hermaeidae" genus="Cyerce" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91FEAFFFFA4FFC0FDEA44E5FE6A218A" box="[389,453,458,481]" italics="true" pageId="30" pageNumber="970">Cyerce</emphasis>
|
||
</taxonomicName>
|
||
, leaving it unclear whether
|
||
<emphasis id="B91FEAFFFFA4FFC0FEB144E5FF66226B" italics="true" pageId="30" pageNumber="970">
|
||
Pe.
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FC1E44C6FF66226B" box="[113,201,489,512]" class="Bivalvia" family="Penicillidae" genus="Penicillus" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="capitatus">capitatus</taxonomicName>
|
||
</emphasis>
|
||
is a preferred host of
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FDC644C6FD90226A" authorityName="Moreno & Rico & Middlebrooks & Medrano & Valdés & Krug" authorityYear="2024" box="[425,575,489,513]" class="Gastropoda" family="Hermaeidae" genus="Cyerce" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="willetteorum">
|
||
<emphasis id="B91FEAFFFFA4FFC0FDC644C6FD90226A" box="[425,575,489,513]" italics="true" pageId="30" pageNumber="970">C. willetteorum</emphasis>
|
||
</taxonomicName>
|
||
or was merely the more locally abundant
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FD304727FE732274" box="[351,476,520,544]" class="Bivalvia" family="Penicillidae" genus="Penicillus" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B91FEAFFFFA4FFC0FD304727FE15224B" box="[351,442,520,544]" italics="true" pageId="30" pageNumber="970">Penicillus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
at the
|
||
<typeStatus id="54D0884FFFA4FFC0FE4C4727FDFE224B" box="[547,593,520,544]" pageId="30" pageNumber="970">type</typeStatus>
|
||
locality. Further study is needed to understand the basis of host specificity within the
|
||
<taxonomicName id="4C6B4D6EFFA4FFC0FCF74768FEBC2235" authorityName="sensu Engel" authorityYear="1927" box="[152,275,582,606]" class="Gastropoda" family="Hermaeidae" genus="Cyerce" kingdom="Animalia" pageId="30" pageNumber="970" phylum="Mollusca" rank="species" species="antillensis">
|
||
<emphasis id="B91FEAFFFFA4FFC0FCF74768FEBC2235" box="[152,275,582,606]" italics="true" pageId="30" pageNumber="970">C. antillensis</emphasis>
|
||
</taxonomicName>
|
||
complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |