279 lines
39 KiB
XML
279 lines
39 KiB
XML
<document id="436F421390DCAB52D2B393AE6A3AFE8D" ID-CLB-Dataset="54394" ID-DOI="10.11646/zootaxa.5278.1.2" ID-GBIF-Dataset="cabaaf5e-455a-4c57-84ca-aed0c83106ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894979" ID-ZooBank="9D0E61E2-298C-4248-BD76-A2C392372F90" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683182983640" checkinUser="plazi" docAuthor="Hansson, Christer" docDate="2023" docId="03C287F3A41BFFE0FF7BFAED1CA7FBBF" docLanguage="en" docName="zootaxa.5278.1.2.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Grotiusomyia guanacastensis Hansson 2023, sp. nov." docType="treatment" docUuid="A0C46708-A424-406E-AE86-689FC1AEA882" docUuidSource="ZooBank" docVersion="7" lastPageNumber="49" masterDocId="FFFBFF8BA412FFEAFFECFFD61D5BFFE6" masterDocTitle="Review of Grotiusomyia Girault (Chalcidoidea: Eulophidae), an exclusively New World genus, including 17 new species" masterLastPageNumber="77" masterPageNumber="39" pageNumber="48" updateTime="1699515298434" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E635598DC70D6DF50F48B2E526B0232D">Review of Grotiusomyia Girault (Chalcidoidea: Eulophidae), an exclusively New World genus, including 17 new species</mods:title>
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<mods:name id="B705CCB14DC31EDCAAAF098CF939C1D7" type="personal">
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<mods:namePart id="6D222409E65A74473FC598F95379FCB5">Hansson, Christer</mods:namePart>
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<mods:title id="C6BA06EA8F345AA203EF49E7497E1EA7">Zootaxa</mods:title>
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<mods:part id="F017025DACA603FEBEE22BB7A148E85E">
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<mods:date id="4208D08B382E729F081CF8430813E733">2023</mods:date>
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<mods:number id="DFD94042C5C60596A2A540F817F91A32">2023-05-04</mods:number>
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<mods:number id="C58EBCA67047E49EA3E1AC943BF87EAE">5278</mods:number>
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<treatment id="03C287F3A41BFFE0FF7BFAED1CA7FBBF" ID-DOI="http://doi.org/10.5281/zenodo.7894965" ID-GBIF-Taxon="209582908" ID-Zenodo-Dep="7894965" LSID="urn:lsid:zoobank.org:act:A0C46708-A424-406E-AE86-689FC1AEA882" httpUri="http://treatment.plazi.org/id/03C287F3A41BFFE0FF7BFAED1CA7FBBF" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48">
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<subSubSection id="C371656EA41BFFE3FF7BFAED1F3BFAB3" box="[151,608,1339,1365]" pageId="9" pageNumber="48" type="nomenclature">
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<paragraph id="8BD436E5A41BFFE3FF7BFAED1F3BFAB3" blockId="9.[151,608,1339,1401]" box="[151,608,1339,1365]" pageId="9" pageNumber="48">
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<heading id="D09C8189A41BFFE3FF7BFAED1F3BFAB3" bold="true" box="[151,608,1339,1365]" fontSize="11" level="1" pageId="9" pageNumber="48" reason="1">
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<emphasis id="B91FEAF7A41BFFE3FF7BFAED1F3BFAB3" bold="true" box="[151,608,1339,1365]" pageId="9" pageNumber="48">
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<taxonomicName id="4C6B4D66A41BFFE3FF7BFAED1CA0FAB3" authority="Hansson, 2023" authorityName="Hansson" authorityYear="2023" box="[151,507,1339,1365]" class="Insecta" family="Eulophidae" genus="Grotiusomyia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="guanacastensis" status="sp. nov.">
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<emphasis id="B91FEAF7A41BFFE3FF7BFAED1CA0FAB3" bold="true" box="[151,507,1339,1365]" italics="true" pageId="9" pageNumber="48">Grotiusomyia guanacastensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22C578CA41BFFE3FDEFFAED1F3BFAB3" box="[515,608,1339,1365]" pageId="9" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C371656EA41BFFE3FF7BFA891CDFFA9F" box="[151,388,1374,1401]" pageId="9" pageNumber="48" type="description">
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<paragraph id="8BD436E5A41BFFE3FF7BFA891CDFFA9F" blockId="9.[151,608,1339,1401]" box="[151,388,1374,1401]" pageId="9" pageNumber="48">
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<figureCitation id="13502A60A41BFFE3FF7BFA891D85FA9F" box="[151,222,1375,1401]" captionStart="FIGURES 7–12" captionStartId="22.[151,264,1882,1906]" captionTargetBox="[151,1435,181,1858]" captionTargetId="figure-35@22.[151,1435,181,1858]" captionTargetPageId="22" captionText="FIGURES 7–12. Grotiusomyia spp. nov., body in lateral view, females; 7 G. guanacastensis, holotype; 8 G. hansoni, holotype; 9 G. jimenezi, holotype; 10 G. lascrucensis, holotype; 11 G. leucaena, holotype; 12 G. nicoyae, holotype." figureDoi="http://doi.org/10.5281/zenodo.7894985" httpUri="https://zenodo.org/record/7894985/files/figure.png" pageId="9" pageNumber="48">Figs 7</figureCitation>
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,
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<figureCitation id="13502A60A41BFFE3FF06FA891C53FA9F" box="[234,264,1375,1401]" captionStart="FIGURES 25–30" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[159,1424,181,1920]" captionTargetId="figure-17@25.[159,1428,181,1920]" captionTargetPageId="25" captionText="FIGURES 25–30. Grotiusomyia spp., body in dorsal view; 25–26. G. azofeifai sp. nov.; 25 female paratype; 26 male paratype; 27 G. flavicornis Girault, female non-type; 28 G. flavimacula sp. nov., female holotype; 29 G. guanacastensis sp. nov., female holotype; 30 G. hansoni sp. nov., female holotype." figureDoi="http://doi.org/10.5281/zenodo.7894993" httpUri="https://zenodo.org/record/7894993/files/figure.png" pageId="9" pageNumber="48">29</figureCitation>
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,
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<figureCitation id="13502A60A41BFFE3FEF8FA881C69FA9F" box="[276,306,1374,1401]" captionStart="FIGURES 51–57" captionStartId="29.[151,264,1879,1903]" captionTargetBox="[151,1435,191,1855]" captionTargetId="figure-17@29.[151,1435,181,1855]" captionTargetPageId="29" captionText="FIGURES 51–57. Grotiusomyia spp. nov., head+antenna in lateral view; 51–52. G. annesmithae; 51 female holotype; 52 male paratype; 53–54. G. azofeifai; 53 female paratype; 54 male paratype; 55 G. flavimacula, female holotype; 56 G. guanacastensis, female holotype; 57 G. jimenezi, female holotype." figureDoi="http://doi.org/10.5281/zenodo.7895005" httpUri="https://zenodo.org/record/7895005/files/figure.png" pageId="9" pageNumber="48">56</figureCitation>
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,
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<figureCitation id="13502A60A41BFFE3FED2FA891C07FA9F" box="[318,348,1375,1401]" captionStart="FIGURES 76–83" captionStartId="33.[151,264,1764,1788]" captionTargetBox="[151,1435,181,1740]" captionTargetId="figure-17@33.[151,1435,181,1740]" captionTargetPageId="33" captionText="FIGURES 76–83. Grotiusomyia spp., vertex, females; 76 G. flavicornis Girault, non-type; 77 G. nigricans (Howard), non-type; 78 G. albiclava sp. nov., holotype; 79 G. annesmithae sp. nov., holotype; 80 G. azofeifai sp. nov., paratype; 81 G. flavimacula sp. nov., holotype; 82 G. guanacastensis sp. nov., holotype; 83 G. hansoni sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.7895013" httpUri="https://zenodo.org/record/7895013/files/figure.png" pageId="9" pageNumber="48">82</figureCitation>
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,
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<figureCitation id="13502A60A41BFFE3FE84FA891CDFFA9F" box="[360,388,1375,1401]" captionStart="FIGURES 92–102" captionStartId="35.[151,264,1828,1852]" captionTargetBox="[151,1435,181,1804]" captionTargetId="figure-17@35.[151,1435,181,1804]" captionTargetPageId="35" captionText="FIGURES 92–102. Grotiusomyia spp., females; 92–93. Vertex; 92 G. rubii sp. nov., holotype; 93 G. septentrionalis sp. nov., holotype; 94–102. Head in frontal view; 94 G. nigricans (Howard), non-type specimen; 95 G. albiclava sp. nov., holotype; 96 G. annesmithae sp. nov., holotype; 97 G. azofeifai sp. nov., paratype; 98 G. flavimacula sp. nov., holotype; 99 G. guanacastensis sp. nov., paratype; 100 G. hansoni sp. nov., holotype; 101 G. lascrucensis sp. nov., holotype; 102 G. jimenezi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.7895017" httpUri="https://zenodo.org/record/7895017/files/figure.png" pageId="9" pageNumber="48">99</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C371656EA41BFFE3FF7BFA701CF0F907" pageId="9" pageNumber="48" type="diagnosis">
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<paragraph id="8BD436E5A41BFFE3FF7BFA701CF0F907" blockId="9.[151,1437,1446,2019]" pageId="9" pageNumber="48">
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<emphasis id="B91FEAF7A41BFFE3FF7BFA701C51FA26" bold="true" box="[151,266,1446,1472]" pageId="9" pageNumber="48">Diagnosis</emphasis>
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. Mandibles dark brown with apex yellowish-brown, with a row of small teeth at apex (as in
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<figureCitation id="13502A60A41BFFE3FAC1FA7118D4FA27" box="[1325,1423,1447,1473]" captionStart="FIGURES 103–112" captionStartId="36.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-35@36.[151,1435,181,1819]" captionTargetPageId="36" captionText="FIGURES 103–112. Grotiusomyia spp. nov., females; 103–110. Head in frontal view; 103 G. leucaena, paratype; 104 G. nicoyae, paratype; 105 G. noyesi, holotype; 106 G. phalaros, paratype; 107 G. pilosa, paratype; 108 G. reticulata, paratype; 109 G. rubii, holotype; 110 G. septentrionalis, paratype; 111–112. Mandibles; 111 G. phalaros, paratype; 112 G. lascrucensis, holotype." figureDoi="http://doi.org/10.5281/zenodo.7895021" httpUri="https://zenodo.org/record/7895021/files/figure.png" pageId="9" pageNumber="48">Fig. 112</figureCitation>
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); flagellum dark brown, 0.7× as long as width of mesoscutum, F4 0.6× as long as wide, apex of clava pointed (
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<figureCitation id="13502A60A41BFFE3FA86FA1D1DFCF9EF" captionStart="FIGURES 7–12" captionStartId="22.[151,264,1882,1906]" captionTargetBox="[151,1435,181,1858]" captionTargetId="figure-35@22.[151,1435,181,1858]" captionTargetPageId="22" captionText="FIGURES 7–12. Grotiusomyia spp. nov., body in lateral view, females; 7 G. guanacastensis, holotype; 8 G. hansoni, holotype; 9 G. jimenezi, holotype; 10 G. lascrucensis, holotype; 11 G. leucaena, holotype; 12 G. nicoyae, holotype." figureDoi="http://doi.org/10.5281/zenodo.7894985" httpUri="https://zenodo.org/record/7894985/files/figure.png" pageId="9" pageNumber="48">Figs 7</figureCitation>
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,
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<figureCitation id="13502A60A41BFFE3FF58FA381D8FF9EF" box="[180,212,1518,1545]" captionStart="FIGURES 51–57" captionStartId="29.[151,264,1879,1903]" captionTargetBox="[151,1435,191,1855]" captionTargetId="figure-17@29.[151,1435,181,1855]" captionTargetPageId="29" captionText="FIGURES 51–57. Grotiusomyia spp. nov., head+antenna in lateral view; 51–52. G. annesmithae; 51 female holotype; 52 male paratype; 53–54. G. azofeifai; 53 female paratype; 54 male paratype; 55 G. flavimacula, female holotype; 56 G. guanacastensis, female holotype; 57 G. jimenezi, female holotype." figureDoi="http://doi.org/10.5281/zenodo.7895005" httpUri="https://zenodo.org/record/7895005/files/figure.png" pageId="9" pageNumber="48">56</figureCitation>
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); frons between scrobes and eyes and vertex with strong reticulation (
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<figureCitation id="13502A60A41BFFE3FC02FA39191DF9EF" box="[1006,1094,1519,1545]" captionStart="FIGURES 76–83" captionStartId="33.[151,264,1764,1788]" captionTargetBox="[151,1435,181,1740]" captionTargetId="figure-17@33.[151,1435,181,1740]" captionTargetPageId="33" captionText="FIGURES 76–83. Grotiusomyia spp., vertex, females; 76 G. flavicornis Girault, non-type; 77 G. nigricans (Howard), non-type; 78 G. albiclava sp. nov., holotype; 79 G. annesmithae sp. nov., holotype; 80 G. azofeifai sp. nov., paratype; 81 G. flavimacula sp. nov., holotype; 82 G. guanacastensis sp. nov., holotype; 83 G. hansoni sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.7895013" httpUri="https://zenodo.org/record/7895013/files/figure.png" pageId="9" pageNumber="48">Figs 82</figureCitation>
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,
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<figureCitation id="13502A60A41BFFE3FBBFFA391928F9EF" box="[1107,1139,1519,1545]" captionStart="FIGURES 92–102" captionStartId="35.[151,264,1828,1852]" captionTargetBox="[151,1435,181,1804]" captionTargetId="figure-17@35.[151,1435,181,1804]" captionTargetPageId="35" captionText="FIGURES 92–102. Grotiusomyia spp., females; 92–93. Vertex; 92 G. rubii sp. nov., holotype; 93 G. septentrionalis sp. nov., holotype; 94–102. Head in frontal view; 94 G. nigricans (Howard), non-type specimen; 95 G. albiclava sp. nov., holotype; 96 G. annesmithae sp. nov., holotype; 97 G. azofeifai sp. nov., paratype; 98 G. flavimacula sp. nov., holotype; 99 G. guanacastensis sp. nov., paratype; 100 G. hansoni sp. nov., holotype; 101 G. lascrucensis sp. nov., holotype; 102 G. jimenezi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.7895017" httpUri="https://zenodo.org/record/7895017/files/figure.png" pageId="9" pageNumber="48">99</figureCitation>
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); occipital margin with an edge behind ocellar triangle (
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<figureCitation id="13502A60A41BFFE3FE31F9C51F6AF9CB" box="[477,561,1555,1581]" captionStart="FIGURES 76–83" captionStartId="33.[151,264,1764,1788]" captionTargetBox="[151,1435,181,1740]" captionTargetId="figure-17@33.[151,1435,181,1740]" captionTargetPageId="33" captionText="FIGURES 76–83. Grotiusomyia spp., vertex, females; 76 G. flavicornis Girault, non-type; 77 G. nigricans (Howard), non-type; 78 G. albiclava sp. nov., holotype; 79 G. annesmithae sp. nov., holotype; 80 G. azofeifai sp. nov., paratype; 81 G. flavimacula sp. nov., holotype; 82 G. guanacastensis sp. nov., holotype; 83 G. hansoni sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.7895013" httpUri="https://zenodo.org/record/7895013/files/figure.png" pageId="9" pageNumber="48">Fig. 82</figureCitation>
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); mesoscutum and mesoscutellum with same colour and reticulation (
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<figureCitation id="13502A60A41BFFE3FAD7F9C518D4F9CB" box="[1339,1423,1555,1581]" captionStart="FIGURES 25–30" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[159,1424,181,1920]" captionTargetId="figure-17@25.[159,1428,181,1920]" captionTargetPageId="25" captionText="FIGURES 25–30. Grotiusomyia spp., body in dorsal view; 25–26. G. azofeifai sp. nov.; 25 female paratype; 26 male paratype; 27 G. flavicornis Girault, female non-type; 28 G. flavimacula sp. nov., female holotype; 29 G. guanacastensis sp. nov., female holotype; 30 G. hansoni sp. nov., female holotype." figureDoi="http://doi.org/10.5281/zenodo.7894993" httpUri="https://zenodo.org/record/7894993/files/figure.png" pageId="9" pageNumber="48">Fig. 29</figureCitation>
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); mesoscutellum convex; dorsellum ±flat, smooth and shiny, posterior margin ±evenly rounded; fore wing with 4–5 setae in apico-dorsal part of costal cell, completely hyaline (
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<figureCitation id="13502A60A41BFFE3FCDFF98D1E2DF993" box="[819,886,1627,1653]" captionStart="FIGURES 7–12" captionStartId="22.[151,264,1882,1906]" captionTargetBox="[151,1435,181,1858]" captionTargetId="figure-35@22.[151,1435,181,1858]" captionTargetPageId="22" captionText="FIGURES 7–12. Grotiusomyia spp. nov., body in lateral view, females; 7 G. guanacastensis, holotype; 8 G. hansoni, holotype; 9 G. jimenezi, holotype; 10 G. lascrucensis, holotype; 11 G. leucaena, holotype; 12 G. nicoyae, holotype." figureDoi="http://doi.org/10.5281/zenodo.7894985" httpUri="https://zenodo.org/record/7894985/files/figure.png" pageId="9" pageNumber="48">Fig. 7</figureCitation>
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); coxae black with metallic tinges (
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<figureCitation id="13502A60A41BFFE3FAECF98D181FF993" box="[1280,1348,1627,1653]" captionStart="FIGURES 7–12" captionStartId="22.[151,264,1882,1906]" captionTargetBox="[151,1435,181,1858]" captionTargetId="figure-35@22.[151,1435,181,1858]" captionTargetPageId="22" captionText="FIGURES 7–12. Grotiusomyia spp. nov., body in lateral view, females; 7 G. guanacastensis, holotype; 8 G. hansoni, holotype; 9 G. jimenezi, holotype; 10 G. lascrucensis, holotype; 11 G. leucaena, holotype; 12 G. nicoyae, holotype." figureDoi="http://doi.org/10.5281/zenodo.7894985" httpUri="https://zenodo.org/record/7894985/files/figure.png" pageId="9" pageNumber="48">Fig. 7</figureCitation>
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); gaster (
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<figureCitation id="13502A60A41BFFE3FF73F9A91DABF97F" box="[159,240,1663,1689]" captionStart="FIGURES 25–30" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[159,1424,181,1920]" captionTargetId="figure-17@25.[159,1428,181,1920]" captionTargetPageId="25" captionText="FIGURES 25–30. Grotiusomyia spp., body in dorsal view; 25–26. G. azofeifai sp. nov.; 25 female paratype; 26 male paratype; 27 G. flavicornis Girault, female non-type; 28 G. flavimacula sp. nov., female holotype; 29 G. guanacastensis sp. nov., female holotype; 30 G. hansoni sp. nov., female holotype." figureDoi="http://doi.org/10.5281/zenodo.7894993" httpUri="https://zenodo.org/record/7894993/files/figure.png" pageId="9" pageNumber="48">Fig. 29</figureCitation>
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) 1.8× as long as wide; Gt
|
||
<subScript id="17EF34A0A41BFFE3FDE1F9471F4EF979" attach="left" box="[525,533,1681,1695]" fontSize="6" pageId="9" pageNumber="48">1</subScript>
|
||
0.3× as long as length of gaster, dorsal surface smooth, posterior margin sinuate; Gt
|
||
<subScript id="17EF34A0A41BFFE3FF5FF9631DE0F925" attach="left" box="[179,187,1717,1731]" fontSize="6" pageId="9" pageNumber="48">6</subScript>
|
||
0.1× as long as length of gaster with anterior part 1.7× as wide as width of posterior part, posterior margin straight with sharp edge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371656EA41BFFE0FF2BF93C1CD3FD2B" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" type="description">
|
||
<paragraph id="8BD436E5A41BFFE3FF2BF93C1E6EF83B" blockId="9.[151,1437,1446,2019]" pageId="9" pageNumber="48">
|
||
<emphasis id="B91FEAF7A41BFFE3FF2BF93C1F62F8E2" bold="true" box="[199,569,1770,1796]" pageId="9" pageNumber="48">
|
||
Description (female
|
||
<typeStatus id="54D08847A41BFFE3FE2EF93C1F77F8E2" box="[450,556,1770,1796]" pageId="9" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Length
|
||
<quantity id="4C939B00A41BFFE3FD4CF93D1FA6F8E3" box="[672,765,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="48" unit="mm" value="2.1">2.1 mm</quantity>
|
||
. Frons between scrobes and eyes and vertex with strong reticulation, scrobes smooth. Mandibles with a row of small teeth at apex. Occipital margin with an edge behind ocellar triangle. Dorsellum ±flat, smooth and shiny, posterior margin ±evenly rounded. Propodeal callus with 18 setae. Fore wing with seven setae on dorsal surface of submarginal vein; with 16 admarginal setae; costal cell with 18 setae on ventral surface in a single row, apico-dorsal margin with five setae; speculum distinct and closed below. Gaster long ovate; Gt
|
||
<subScript id="17EF34A0A41BFFE3FE7DF8671CC2F859" attach="left" box="[401,409,1969,1983]" fontSize="6" pageId="9" pageNumber="48">1</subScript>
|
||
smooth and shiny without sculpture, remaining tergites with weak reticulation; Gt
|
||
<subScript id="17EF34A0A41BFFE3FAB4F867183BF859" attach="left" box="[1368,1376,1969,1983]" fontSize="6" pageId="9" pageNumber="48">1</subScript>
|
||
with posterior margin sinuate, Gt
|
||
<subScript id="17EF34A0A41BFFE3FE3DF8031C82F805" attach="left" box="[465,473,2005,2019]" fontSize="6" pageId="9" pageNumber="48">6</subScript>
|
||
with posterior margin straight.
|
||
</paragraph>
|
||
<paragraph id="8BD436E5A418FFE0FF2BFF411C18FEFB" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="49">Colour. Head black with metallic tinges. Mandibles dark brown with apex yellowish-brown. Antennae with scape yellowish-brown, pedicel and flagellum dark brown. Mesosoma black with metallic tinges. Legs with coxae black with metallic tinges, trochanters and femora dark brown, tibiae and tarsi yellowish-brown. Gaster black with metallic tinges.</paragraph>
|
||
<paragraph id="8BD436E5A418FFE0FF2BFEF11872FD87" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="49">
|
||
Relative measurements: head length, dorsal view 20; head length, frontal view 35; head width 47; POL 11.5; OOL 4.5; mouth width 13; malar space 10; eye length 23; lateral ocellus diameter 4; scape length 17; scape width 4; pedicel length 5.5; pedicel width. dorsal view 3.5; flagellum length 31; F1 length 4.5; F1 width 4; F2 length 4; F2 width 5; F3 length 4; F3 width 6; F4 length 4; F4 width 6.5; clava length 10; clava width 6; pronotum length 18; pronotum width 38; mesosoma length 66; mesosoma width 45; midlobe of mesoscutum length 23; mesoscutellum length 20; mesoscutellum width 22; propodeum length 10; hind tibia length 41; hind tibial spur1 length 5; hind tibial spur2 length 3; hind tarsus length 26; hind basitarsus length 8.5; fore wing length 106; fore wing width 48; costal cell length 36; costal cell width 3.5; marginal vein length 29; postmarginal vein length 14; stigmal vein length 11.5; Gt
|
||
<subScript id="17EF34A0A418FFE0FF5FFD8F1DE0FD81" attach="left" box="[179,187,601,615]" fontSize="6" pageId="10" pageNumber="49">1</subScript>
|
||
length 22; Gt
|
||
<subScript id="17EF34A0A418FFE0FEBAFD8F1C05FD81" attach="left" box="[342,350,601,615]" fontSize="6" pageId="10" pageNumber="49">6</subScript>
|
||
length 9; Gt
|
||
<subScript id="17EF34A0A418FFE0FE07FD8F1CA8FD81" attach="left" box="[491,499,601,615]" fontSize="6" pageId="10" pageNumber="49">6</subScript>
|
||
width at base 26; Gt
|
||
<subScript id="17EF34A0A418FFE0FD30FD8F1FBFFD81" attach="left" box="[732,740,601,615]" fontSize="6" pageId="10" pageNumber="49">6</subScript>
|
||
width at apex 15; gaster length 80; gaster width 44.
|
||
</paragraph>
|
||
<paragraph id="8BD436E5A418FFE0FF2BFDBD1C7FFD4F" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="49">
|
||
Variation. Length of body
|
||
<quantity id="4C939B00A418FFE0FE19FDBD1FDAFD63" box="[501,641,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" pageId="10" pageNumber="49" unit="mm" value="1.85" valueMax="2.1" valueMin="1.6">1.6–2.1 mm</quantity>
|
||
. Costal cell with 4–5 setae in apico-dorsal part. Propodeal callus with 10–18 setae.
|
||
</paragraph>
|
||
<paragraph id="8BD436E5A418FFE0FF2BFD651CD3FD2B" blockId="10.[151,1437,151,1113]" box="[199,392,691,717]" pageId="10" pageNumber="49">
|
||
<emphasis id="B91FEAF7A418FFE0FF2BFD651C5FFD2B" bold="true" box="[199,260,691,717]" pageId="10" pageNumber="49">Male</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371656EA418FFE0FF2BFD001CB2FD17" box="[199,489,726,753]" pageId="10" pageNumber="49" type="distribution">
|
||
<paragraph id="8BD436E5A418FFE0FF2BFD001CB2FD17" blockId="10.[151,1437,151,1113]" box="[199,489,726,753]" pageId="10" pageNumber="49">
|
||
<emphasis id="B91FEAF7A418FFE0FF2BFD001C02FD16" bold="true" box="[199,345,726,752]" pageId="10" pageNumber="49">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F37C7675A418FFE0FE8BFD011CBEFD17" box="[359,485,727,753]" name="Costa Rica" pageId="10" pageNumber="49">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371656EA418FFE0FF2BFD2D1E3EFCF3" box="[199,869,763,789]" pageId="10" pageNumber="49" type="etymology">
|
||
<paragraph id="8BD436E5A418FFE0FF2BFD2D1E3EFCF3" blockId="10.[151,1437,151,1113]" box="[199,869,763,789]" pageId="10" pageNumber="49">
|
||
<emphasis id="B91FEAF7A418FFE0FF2BFD2D1C13FCF3" bold="true" box="[199,328,763,789]" pageId="10" pageNumber="49">Etymology</emphasis>
|
||
. Named for
|
||
<typeStatus id="54D08847A418FFE0FE3AFD2D1F5DFCF3" box="[470,518,763,789]" pageId="10" pageNumber="49">type</typeStatus>
|
||
area, the province
|
||
<collectingRegion id="49AFF807A418FFE0FD30FD2D1E3AFCF3" box="[732,865,763,789]" country="Costa Rica" name="Guanacaste" pageId="10" pageNumber="49">Guanacaste</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371656EA418FFE0FF2BFCC81CA7FBBF" pageId="10" pageNumber="49" type="materials_examined">
|
||
<paragraph id="8BD436E5A418FFE0FF2BFCC8181DFC2F" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="49">
|
||
<emphasis id="B91FEAF7A418FFE0FF2BFCC81C74FCDE" bold="true" box="[199,303,798,824]" pageId="10" pageNumber="49">Material</emphasis>
|
||
.
|
||
<materialsCitation id="3B033CB8A418FFE0FEAFFCC91F80FCBB" ID-GBIF-Occurrence="4100370373" collectingDate="1985-12-07" collectingDateMax="1985-12-28" collectingDateMin="1985-12-07" collectionCode="NHMUK" collectorName="D. H. Janzen & I. D. Gauld" country="Costa Rica" elevation="300" location="Parque Nacional Santa Rosa" pageId="10" pageNumber="49" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="holotype">
|
||
<typeStatus id="54D08847A418FFE0FEAFFCC91CF0FCDF" box="[323,427,799,825]" pageId="10" pageNumber="49" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D6DFD6CA418FFE0FE54FCC91F52FCDF" box="[440,521,799,825]" pageId="10" pageNumber="49" type="female">female</specimenCount>
|
||
:
|
||
<collectingCountry id="F37C7675A418FFE0FDF4FCC91F9BFCDF" box="[536,704,799,825]" name="Costa Rica" pageId="10" pageNumber="49">COSTA RICA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49AFF807A418FFE0FD3FFCC91E0CFCDF" box="[723,855,799,825]" country="Costa Rica" name="Guanacaste" pageId="10" pageNumber="49">Guanacaste</collectingRegion>
|
||
,
|
||
<location id="8EB4603EA418FFE0FC84FCC919E2FCDF" LSID="urn:lsid:plazi:treatment:03C287F3A41BFFE0FF7BFAED1CA7FBBF:8EB4603EA418FFE0FC84FCC919E2FCDF" box="[872,1209,799,825]" country="Costa Rica" name="Parque Nacional Santa Rosa" pageId="10" pageNumber="49" stateProvince="Guanacaste">Parque Nacional Santa Rosa</location>
|
||
,
|
||
<quantity id="4C939B00A418FFE0FB26FCC91856FCDF" box="[1226,1293,799,825]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="49" unit="m" value="300.0">
|
||
<elevation id="0046D1D6A418FFE0FB26FCC91856FCDF" box="[1226,1293,799,825]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="49" unit="m" value="300.0">300m</elevation>
|
||
</quantity>
|
||
, SE-6-C,
|
||
<date id="FFD51025A418FFE0FA69FCC91C42FCBB" pageId="10" pageNumber="49" value="1985-12-07" valueMax="1985-12-28" valueMin="1985-12-07">
|
||
<collectingDate id="EF91E9CDA418FFE0FA69FCC91C42FCBB" pageId="10" pageNumber="49" value="1985-12-07" valueMax="1985-12-28" valueMin="1985-12-07">7- 28.xii.1985</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269E5333A418FFE0FECAFC951CE9FCBB" box="[294,434,835,861]" pageId="10" pageNumber="49">D.H. Janzen</collectorName>
|
||
&
|
||
<collectorName id="269E5333A418FFE0FE35FC951F0AFCBB" box="[473,593,835,861]" pageId="10" pageNumber="49">I.D. Gauld</collectorName>
|
||
(
|
||
<collectionCode id="ED7AAE20A418FFE0FD8EFC951F88FCBB" box="[610,723,835,861]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="49" type="Museum">NHMUK</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B033CB8A418FFE0FD0AFC951E70FC67" ID-GBIF-Occurrence="4100370408" collectingDate="1986-02-08" collectingDateMax="1986-03-02" collectingDateMin="1986-02-08" collectionCode="MZLU, NHMUK" collectorName="D. H. Janzen & I. D. Gauld" country="Costa Rica" elevation="300" location="Parque Nacional Santa Rosa" pageId="10" pageNumber="49" specimenCount="4" specimenCount-female="4" stateProvince="Guanacaste" typeStatus="paratype">
|
||
<typeStatus id="54D08847A418FFE0FD0AFC951E08FCBB" box="[742,851,835,861]" pageId="10" pageNumber="49" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<specimenCount id="9D6DFD6CA418FFE0FC8BFC951ED2FCBB" box="[871,905,835,861]" pageId="10" pageNumber="49" type="female">3♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED7AAE20A418FFE0FC74FC951EB0FCBB" box="[920,1003,835,861]" collectionName="Sweden, Lund, Lund University" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="10" pageNumber="49">MZLU</collectionCode>
|
||
;
|
||
<collectionCode id="ED7AAE20A418FFE0FC14FC951933FCBB" box="[1016,1128,835,861]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="49" type="Museum">NHMUK</collectionCode>
|
||
):
|
||
<specimenCount id="9D6DFD6CA418FFE0FB97FC9519C5FCBB" box="[1147,1182,835,861]" pageId="10" pageNumber="49" type="female">1♀</specimenCount>
|
||
from same locality as holotype but collected
|
||
<date id="FFD51025A418FFE0FE76FCB11F6CFC67" box="[410,567,871,897]" pageId="10" pageNumber="49" value="1986-02-08" valueMax="1986-03-02" valueMin="1986-02-08">
|
||
<collectingDate id="EF91E9CDA418FFE0FE76FCB11F6CFC67" box="[410,567,871,897]" pageId="10" pageNumber="49" value="1986-02-08" valueMax="1986-03-02" valueMin="1986-02-08">8.ii-2.iii.1986</collectingDate>
|
||
</date>
|
||
; barcoded specimens
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="3B033CB8A418FFE0FCDAFCB11CDEFC2F" ID-GBIF-Occurrence="4100370342" collectingDate="2017-03-16" collectingDateMax="2017-03-23" collectingDateMin="2017-03-16" collectionCode="PL" collectorName="D. Janzen & W. Hallwachs" country="Costa Rica" elevation="831" latitude="10.7632" location="de Conservacion Guanacaste" longLatPrecision="7" longitude="-85.3338" pageId="10" pageNumber="49" specimenCode="BIOUG57030" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
|
||
<specimenCount id="9D6DFD6CA418FFE0FCDAFCB11E02FC67" box="[822,857,871,897]" pageId="10" pageNumber="49" type="female">1♀</specimenCount>
|
||
<collectingCountry id="F37C7675A418FFE0FC8DFCB1195CFC67" box="[865,1031,871,897]" name="Costa Rica" pageId="10" pageNumber="49">COSTA RICA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49AFF807A418FFE0FBFFFCB119CCFC67" box="[1043,1175,871,897]" country="Costa Rica" name="Guanacaste" pageId="10" pageNumber="49">Guanacaste</collectingRegion>
|
||
, Area
|
||
<location id="8EB4603EA418FFE0FB0CFCB11C40FC43" LSID="urn:lsid:plazi:treatment:03C287F3A41BFFE0FF7BFAED1CA7FBBF:8EB4603EA418FFE0FB0CFCB11C40FC43" country="Costa Rica" latitude="10.7632" longLatPrecision="7" longitude="-85.3338" name="de Conservacion Guanacaste" pageId="10" pageNumber="49" stateProvince="Guanacaste">de Conservacion Guanacaste</location>
|
||
,
|
||
<location id="8EB4603EA418FFE0FEC8FC5D1CEEFC43" LSID="urn:lsid:plazi:treatment:03C287F3A41BFFE0FF7BFAED1CA7FBBF:8EB4603EA418FFE0FEC8FC5D1CEEFC43" box="[292,437,907,933]" country="Costa Rica" latitude="10.7632" longLatPrecision="7" longitude="-85.3338" name="Sector Pailas" pageId="10" pageNumber="49" stateProvince="Guanacaste">Sector Pailas</location>
|
||
, Pailas Dos PL12-5,
|
||
<geoCoordinate id="EE5F5022A418FFE0FD7AFC5D1E4BFC43" box="[662,784,907,933]" degrees="10.7632" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="5" value="10.7632">10.7632°N</geoCoordinate>
|
||
<geoCoordinate id="EE5F5022A418FFE0FCF9FC5C1ECEFC43" box="[789,917,906,933]" degrees="85.3338" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="5" value="-85.3338">85.3338°W</geoCoordinate>
|
||
,
|
||
<quantity id="4C939B00A418FFE0FC72FC5D1E85FC43" box="[926,990,907,933]" metricMagnitude="2" metricUnit="m" metricValue="8.309999999999999" pageId="10" pageNumber="49" unit="m" value="831.0">
|
||
<elevation id="0046D1D6A418FFE0FC72FC5D1E85FC43" box="[926,990,907,933]" metricMagnitude="2" metricUnit="m" metricValue="8.309999999999999" pageId="10" pageNumber="49" unit="m" value="831.0">831m</elevation>
|
||
</quantity>
|
||
<date id="FFD51025A418FFE0FC0EFC5D19DFFC43" box="[994,1156,907,933]" pageId="10" pageNumber="49" value="2017-03-16" valueMax="2017-03-23" valueMin="2017-03-16">
|
||
<collectingDate id="EF91E9CDA418FFE0FC0EFC5D19DFFC43" box="[994,1156,907,933]" pageId="10" pageNumber="49" value="2017-03-16" valueMax="2017-03-23" valueMin="2017-03-16">16-23.iii.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269E5333A418FFE0FB60FC5D19A0FC43" box="[1164,1275,907,933]" pageId="10" pageNumber="49">D. Janzen</collectorName>
|
||
,
|
||
<collectorName id="269E5333A418FFE0FAEFFC5D18C7FC43" box="[1283,1436,907,933]" pageId="10" pageNumber="49">W. Hallwachs</collectorName>
|
||
(
|
||
<specimenCode id="DBCD9E9EA418FFE0FF73FC781C12FC2F" box="[159,329,942,969]" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="10" pageNumber="49" type="University or college">BIOUG57030</specimenCode>
|
||
-G11)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B033CB8A418FFE0FE7CFC791819FC2F" ID-GBIF-Occurrence="4100370351" box="[400,1346,943,969]" collectingDate="2017-02-02" collectingDateMax="2017-02-23" collectingDateMin="2017-02-02" collectionCode="BIOUG" collectorName="D. Janzen & Hallwachs" country="Costa Rica" pageId="10" pageNumber="49" specimenCode="BIOUG60816" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
|
||
<specimenCount id="9D6DFD6CA418FFE0FE7CFC791CE8FC2F" box="[400,435,943,969]" pageId="10" pageNumber="49" type="female">1♀</specimenCount>
|
||
from same locality as previous but collected
|
||
<date id="FFD51025A418FFE0FC43FC791914FC2F" box="[943,1103,943,969]" pageId="10" pageNumber="49" value="2017-02-23" valueMax="2017-02-02" valueMin="2017-02-23">
|
||
<collectingDate id="EF91E9CDA418FFE0FC43FC791914FC2F" box="[943,1103,943,969]" pageId="10" pageNumber="49" value="2017-02-23" valueMax="2017-02-02" valueMin="2017-02-23">23.ii-2.ii.2017</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="DBCD9E9EA418FFE0FBB2FC791853FC2F" box="[1118,1288,943,969]" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="10" pageNumber="49" type="University or college">BIOUG60816</specimenCode>
|
||
-C11)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD436E5A418FFE0FF7BFC051CA7FBBF" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="49">
|
||
<materialsCitation id="3B033CB8A418FFE0FF7BFC051CA3FBBF" ID-GBIF-Occurrence="4100370396" collectingDate="2015-04-23" collectingDateMax="2015-04-30" collectingDateMin="2015-04-23" collectionCode="MZLU" collectorName="D. Janzen W. Hallwachs" country="Costa Rica" elevation="820" latitude="10.7631" location="Area de Conservacion Guanacaste" longLatPrecision="7" longitude="-85.3344" municipality="Sector Pailas" pageId="10" pageNumber="49" specimenCode="BIOUG 44761" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste">
|
||
Non-type material:
|
||
<specimenCount id="9D6DFD6CA418FFE0FE9BFC051CC1FC0B" box="[375,410,979,1005]" pageId="10" pageNumber="49" type="female">1♀</specimenCount>
|
||
<collectingCountry id="F37C7675A418FFE0FE4FFC051F11FC0B" box="[419,586,979,1005]" name="Costa Rica" pageId="10" pageNumber="49">COSTA RICA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49AFF807A418FFE0FDBBFC051F80FC0B" box="[599,731,979,1005]" country="Costa Rica" name="Guanacaste" pageId="10" pageNumber="49">Guanacaste</collectingRegion>
|
||
,
|
||
<location id="8EB4603EA418FFE0FD0BFC051929FC0B" LSID="urn:lsid:plazi:treatment:03C287F3A41BFFE0FF7BFAED1CA7FBBF:8EB4603EA418FFE0FD0BFC051929FC0B" box="[743,1138,979,1005]" country="Costa Rica" latitude="10.7631" longLatPrecision="7" longitude="-85.3344" municipality="Sector Pailas" name="Area de Conservacion Guanacaste" pageId="10" pageNumber="49" stateProvince="Guanacaste">Area de Conservacion Guanacaste</location>
|
||
;
|
||
<collectingMunicipality id="6BB0AC9FA418FFE0FB93FC05184DFC0B" box="[1151,1302,979,1005]" pageId="10" pageNumber="49">Sector Pailas</collectingMunicipality>
|
||
, Pailas Dos PL12-3,
|
||
<geoCoordinate id="EE5F5022A418FFE0FF10FC211C2DFBF7" box="[252,374,1015,1041]" degrees="10.7631" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="5" value="10.7631">10.7631°N</geoCoordinate>
|
||
<geoCoordinate id="EE5F5022A418FFE0FE93FC201F5BFBF7" box="[383,512,1014,1041]" degrees="85.3344" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="5" value="-85.3344">85.3344°W</geoCoordinate>
|
||
,
|
||
<quantity id="4C939B00A418FFE0FDE2FC211F0BFBF7" box="[526,592,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="10" pageNumber="49" unit="m" value="820.0">
|
||
<elevation id="0046D1D6A418FFE0FDE2FC211F0BFBF7" box="[526,592,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="10" pageNumber="49" unit="m" value="820.0">820m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFD51025A418FFE0FDB2FC201FA7FBF7" box="[606,764,1014,1041]" pageId="10" pageNumber="49" value="2015-04-23" valueMax="2015-04-30" valueMin="2015-04-23">
|
||
<collectingDate id="EF91E9CDA418FFE0FDB2FC201FA7FBF7" box="[606,764,1014,1041]" pageId="10" pageNumber="49" value="2015-04-23" valueMax="2015-04-30" valueMin="2015-04-23">23-30.iv.2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269E5333A418FFE0FCE5FC21197FFBF7" box="[777,1060,1015,1041]" pageId="10" pageNumber="49">D. Janzen W. Hallwachs</collectorName>
|
||
, (strongly fragmented specimen) (
|
||
<specimenCode id="DBCD9E9EA418FFE0FF73FBCD1C11FBD3" box="[159,330,1051,1077]" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="10" pageNumber="49" type="University or college">BIOUG 44761</specimenCode>
|
||
-D10) (
|
||
<collectionCode id="ED7AAE20A418FFE0FE75FBCD1CB4FBD3" box="[409,495,1051,1077]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="10" pageNumber="49">MZLU</collectionCode>
|
||
). This specimen is barcoded, but because of bad fragmentation with parts missing it is not included as type material
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |