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<document id="06067776606D62F76C8C893479CB8704" ID-CLB-Dataset="182444" ID-DOI="10.11646/zootaxa.5202.1.1" ID-GBIF-Dataset="45ba25c1-47a9-4001-93d9-294815bf37e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7285600" ID-ZooBank="BF4831D2-D98B-4265-9138-03DB8607B826" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1667554680289" checkinUser="plazi" docAuthor="Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta &amp; Riva, Ignacio De La" docDate="2022" docId="03C287E8FFEEFFDEFF4BF910FEF7E39F" docLanguage="en" docName="zootaxa.5202.1.1.pdf" docOrigin="Zootaxa 5202 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5202.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Philothamnus girardi Bocage 1893" docType="treatment" docVersion="10" lastPageNumber="78" masterDocId="FFFBFF90FFA2FF93FFDCFF9DFFB5E232" masterDocTitle="Synopsis of the terrestrial Reptiles of Equatorial Guinea" masterLastPageNumber="197" masterPageNumber="1" pageNumber="77" updateTime="1732576365292" updateUser="ExternalLinkService">
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<mods:title id="527F30D0E29506D8BA1DF7B41BA0831E">Synopsis of the terrestrial Reptiles of Equatorial Guinea</mods:title>
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<mods:namePart id="3BC3600052925DF3DB8E9208759E7B56">Sánchez-Vialas, Alberto</mods:namePart>
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<mods:namePart id="71FE807F3F879B61942025A1AFD45153">Riva, Ignacio De La</mods:namePart>
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<mods:title id="651F74D52905315736D5585279351233">Zootaxa</mods:title>
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<treatment id="03C287E8FFEEFFDEFF4BF910FEF7E39F" ID-DOI="http://doi.org/10.5281/zenodo.7293465" ID-GBIF-Taxon="204305353" ID-Zenodo-Dep="7293465" LSID="urn:lsid:plazi:treatment:03C287E8FFEEFFDEFF4BF910FEF7E39F" httpUri="http://treatment.plazi.org/id/03C287E8FFEEFFDEFF4BF910FEF7E39F" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" scope_family="Colubridae" scope_order="Squamata">
<subSubSection id="C3716575FFEEFFDFFF4BF910FDF2E495" box="[151,583,1677,1704]" pageId="76" pageNumber="77" type="nomenclature">
<paragraph id="8BD436FEFFEEFFDFFF4BF910FDF2E495" blockId="76.[151,583,1677,1739]" box="[151,583,1677,1704]" pageId="76" pageNumber="77">
<heading id="D09C8192FFEEFFDFFF4BF910FDF2E495" bold="true" box="[151,583,1677,1704]" fontSize="11" level="2" pageId="76" pageNumber="77" reason="4">
<taxonomicName id="4C6B4D7DFFEEFFDFFF4BF910FDF2E495" ID-CoL="4GDBM" authority="Bocage, 1893" authorityName="Bocage" authorityYear="1893" box="[151,583,1677,1704]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="girardi">
<emphasis id="B91FEAECFFEEFFDFFF4BF910FDF2E495" bold="true" box="[151,583,1677,1704]" pageId="76" pageNumber="77">
<emphasis id="B91FEAECFFEEFFDFFF4BF910FE28E49A" bold="true" box="[151,413,1677,1704]" italics="true" pageId="76" pageNumber="77">Philothamnus girardi</emphasis>
Bocage, 1893
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEEFFDFFF4BF92CFEADE4F9" box="[151,280,1713,1739]" pageId="76" pageNumber="77" type="description">
<paragraph id="8BD436FEFFEEFFDFFF4BF92CFEADE4F9" blockId="76.[151,583,1677,1739]" box="[151,280,1713,1739]" pageId="76" pageNumber="77">
<figureCitation id="13502A7BFFEEFFDFFF4BF92CFEADE4F9" box="[151,280,1713,1739]" captionStart="FIGURE 19" captionStartId="80.[151,250,1910,1934]" captionTargetBox="[169,1433,196,1867]" captionTargetId="figure-19@80.[154,1433,181,1886]" captionTargetPageId="80" captionText="FIGURE 19. From left to right: lateral, dorsal and ventral views of the head of: Philothamnus girardi (A) (EBD 27758), Philothamnus heterodermus (B) (MNCN 22864), and Philothamnus nitidus (C) (EBD 27534). Photos: AS-V." figureDoi="http://doi.org/10.5281/zenodo.7285679" httpUri="https://zenodo.org/record/7285679/files/figure.png" pageId="76" pageNumber="77">Figure 19A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEEFFDFFF4BF96BFA9FE53C" box="[151,1322,1782,1806]" pageId="76" pageNumber="77" type="reference_group">
<paragraph id="8BD436FEFFEEFFDFFF4BF96BFA9FE53C" blockId="76.[151,1322,1782,1806]" box="[151,1322,1782,1806]" pageId="76" pageNumber="77">
<treatmentCitationGroup id="AB7B11D0FFEEFFDFFF4BF96BFA9FE53C" box="[151,1322,1782,1806]" pageId="76" pageNumber="77">
<treatmentCitation id="0ACA10EFFFEEFFDFFF4BF96BFD82E53C" author="Bocage, J. V. B." box="[151,567,1782,1806]" page="46" pageId="76" pageNumber="77" year="1893">
<taxonomicName id="4C6B4D7DFFEEFFDFFF4BF96BFD82E53C" ID-CoL="4GDBM" authority="Bocage, 1893 a: 46" authorityName="Bocage" authorityPageNumber="46" authorityYear="1893" box="[151,567,1782,1806]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="girardi">
<emphasis id="B91FEAECFFEEFFDFFF4BF96BFEC4E53C" box="[151,369,1782,1806]" italics="true" pageId="76" pageNumber="77">Philothamnus girardi</emphasis>
<bibRefCitation id="EFFA4B0FFFEEFFDFFEABF96BFD82E53C" author="Bocage, J. V. B." box="[375,567,1782,1806]" pageId="76" pageNumber="77" pagination="45 - 46" refId="ref70843" refString="Bocage, J. V. B. (1893 a) Mammiferos, Aves e Reptis da Ilha de Anno-bom. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 3 (9), 45 - 46." type="journal article" year="1893">Bocage, 1893a: 46</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54D0885CFFEEFFDFFD9DF96BFDC6E53C" box="[577,627,1782,1806]" pageId="76" pageNumber="77">Type</typeStatus>
locality: “Anno Bom Island,
<collectingCountry id="F37C766EFFEEFFDFFC43F96BFC58E53C" box="[927,1005,1782,1806]" name="Guinea-Bissau" pageId="76" pageNumber="77">Guinea</collectingCountry>
”.
<collectingCountry id="F37C766EFFEEFFDFFC23F96BFBD5E53C" box="[1023,1120,1782,1806]" name="Equatorial Guinea" pageId="76" pageNumber="77">Annobon</collectingCountry>
,
<collectingCountry id="F37C766EFFEEFFDFFBB6F96BFA93E53C" box="[1130,1318,1782,1806]" name="Equatorial Guinea" pageId="76" pageNumber="77">Equatorial Guinea</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEEFFDFFF4BF8A1FAA1E564" box="[151,1300,1852,1879]" pageId="76" pageNumber="77" type="distribution">
<paragraph id="8BD436FEFFEEFFDFFF4BF8A1FAA1E564" blockId="76.[151,1437,1852,2023]" box="[151,1300,1852,1879]" pageId="76" pageNumber="77">
<emphasis id="B91FEAECFFEEFFDFFF4BF8A1FE84E564" bold="true" box="[151,305,1852,1878]" pageId="76" pageNumber="77">Distribution.</emphasis>
Endemic to
<collectingCountry id="F37C766EFFEEFFDFFE61F8A1FD90E564" box="[445,549,1852,1878]" name="Equatorial Guinea" pageId="76" pageNumber="77">Annobon</collectingCountry>
(
<bibRefCitation id="EFFA4B0FFFEEFFDFFDE8F8A1FD53E564" author="Bocage, J. V. B." box="[564,742,1852,1878]" pageId="76" pageNumber="77" pagination="45 - 46" refId="ref70843" refString="Bocage, J. V. B. (1893 a) Mammiferos, Aves e Reptis da Ilha de Anno-bom. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 3 (9), 45 - 46." type="journal article" year="1893">Bocage, 1893 a</bibRefCitation>
,b;
<bibRefCitation id="EFFA4B0FFFEEFFDFFCDBF8A1FC09E565" author="Loveridge, A." box="[775,956,1852,1879]" pageId="76" pageNumber="77" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridge 1958</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFFEEFFDFFC14F8A0FB36E564" author="Jesus, J. &amp; Nagy, Z. T. &amp; Branch, W. R. &amp; Wink, M. &amp; Brehm, A. &amp; Harris, J. D." box="[968,1155,1852,1879]" pageId="76" pageNumber="77" pagination="41 - 48" refId="ref81022" refString="Jesus, J., Nagy, Z. T., Branch, W. R., Wink, M., Brehm, A. &amp; Harris, J. D. (2009) Phylogenetic relationships of African green snakes (genera Philothamnus and Hapsidophrys) from Sao Tome, Principe and Annobon islands based on mtDNA sequences, and comments on their colonization and taxonomy. Herpetological Journal, 19, 41 - 48." type="journal article" year="2009">
Jesus
<emphasis id="B91FEAECFFEEFFDFFBD6F8A0FBF7E564" box="[1034,1090,1852,1878]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
2009
</bibRefCitation>
) (
<figureCitation id="13502A7BFFEEFFDFFB45F8A1FAB2E564" box="[1177,1287,1852,1878]" captionStart="MAPS 21" captionStartId="79.[151,223,1943,1967]" captionTargetBox="[202,1385,192,1908]" captionTargetId="figure-18@79.[159,1429,181,1919]" captionTargetPageId="79" captionText="MAPS 21 A–C. Distribution maps for Equatorial Guinean records of (A) Hapsidophrys smaragdina; (B) Philothamnus carinatus; (C) Philothamnus girardi." figureDoi="http://doi.org/10.5281/zenodo.7304727" httpUri="https://zenodo.org/record/7304727/files/figure.png" pageId="76" pageNumber="77">Map 21C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEEFFDEFF1BF8FDFEF7E39F" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="discussion">
<paragraph id="8BD436FEFFEEFFDEFF1BF8FDFCFBE2CB" blockId="76.[151,1437,1852,2023]" lastBlockId="77.[151,1437,150,429]" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
<emphasis id="B91FEAECFFEEFFDFFF1BF8FDFEFAE548" bold="true" box="[199,335,1888,1914]" pageId="76" pageNumber="77">Comments.</emphasis>
<taxonomicName id="4C6B4D7DFFEEFFDFFE81F8FDFDE0E548" authorityName="Bocage" authorityYear="1893" box="[349,597,1888,1914]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="girardi">
<emphasis id="B91FEAECFFEEFFDFFE81F8FDFDE0E548" box="[349,597,1888,1914]" italics="true" pageId="76" pageNumber="77">Philothamnus girardi</emphasis>
</taxonomicName>
could probably be a junior synonym of
<taxonomicName id="4C6B4D7DFFEEFFDFFB8DF8FCFE09E5AC" authority="(Bocage, 1866 a) (Engelbrecht et al. 2019)" authorityName="Engelbrecht et al." authorityYear="2019" baseAuthorityName="Bocage" baseAuthorityYear="1866" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="B91FEAECFFEEFFDFFB8DF8FCFB7BE548" box="[1105,1230,1888,1914]" italics="true" pageId="76" pageNumber="77">P. dorsalis</emphasis>
(
<bibRefCitation id="EFFA4B0FFFEEFFDFFB38F8FDFA20E548" author="Bocage, J. V. B." box="[1252,1429,1888,1914]" pageId="76" pageNumber="77" pagination="37 - 56" refId="ref70648" refString="Bocage, J. V. B. (1866 a) Lista dos reptis das possessoes portuguezas d'Africa occidental que existem no Museu Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Primeira Serie, 1 (1), 37 - 56." type="journal article" year="1866">Bocage, 1866a</bibRefCitation>
) (
<bibRefCitation id="EFFA4B0FFFEEFFDFFF43F819FE01E5AC" author="Engelbrecht, H. M. &amp; Branch, W. R. &amp; Greenbaum, E. &amp; Alexander, G. J. &amp; Jackson, K. &amp; Burger, M. &amp; Conradie, W. &amp; Kusamba, C. &amp; ZassiBoulou, A-G. &amp; Tolley, K. A." box="[159,436,1924,1950]" pageId="76" pageNumber="77" pagination="357 - 365" refId="ref76556" refString="Engelbrecht, H. M., Branch, W. R., Greenbaum, E., Alexander, G. J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., ZassiBoulou, A-G. &amp; Tolley, K. A. (2019) Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution, 130, 357 - 365. https: // doi. org / 10.1016 / j. ympev. 2018.10.023" type="journal article" year="2019">
Engelbrecht
<emphasis id="B91FEAECFFEEFFDFFEEEF818FEDBE5AC" box="[306,366,1924,1950]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
. However, as previously reported (
<bibRefCitation id="EFFA4B0FFFEEFFDFFC8EF819FBB8E5AD" author="Loveridge, A." box="[850,1037,1924,1951]" pageId="76" pageNumber="77" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridge 1958</bibRefCitation>
), all the specimens from
<collectingCountry id="F37C766EFFEEFFDFFAE8F819FA29E5AC" box="[1332,1436,1924,1950]" name="Equatorial Guinea" pageId="76" pageNumber="77">Annobon</collectingCountry>
examined by us have 13 scale rows at mid-body, instead of the 15 rows shown by
<taxonomicName id="4C6B4D7DFFEEFFDFFBF1F834FF69E5D4" authority="(Chippaux &amp; Jackson 2019)" baseAuthorityName="Chippaux &amp; Jackson" baseAuthorityYear="2019" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="B91FEAECFFEEFFDFFBF1F834FB16E5F0" box="[1069,1187,1960,1986]" italics="true" pageId="76" pageNumber="77">P. dorsalis</emphasis>
(
<bibRefCitation id="EFFA4B0FFFEEFFDFFB6DF835FF61E5D4" author="Chippaux, J. P. &amp; Jackson, K." pageId="76" pageNumber="77" refId="ref74358" refString="Chippaux, J. P. &amp; Jackson, K. (2019) Snakes of Central and Western Africa. Johns Hopkins University Press, Baltimore, Maryland, 429 pp." type="book" year="2019">Chippaux &amp; Jackson 2019</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C6B4D7DFFEEFFDFFF3AF851FE62E5D4" authorityName="Bocage" authorityYear="1893" box="[230,471,1996,2022]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="girardi">
<emphasis id="B91FEAECFFEEFFDFFF3AF851FE62E5D4" box="[230,471,1996,2022]" italics="true" pageId="76" pageNumber="77">Philothamnus girardi</emphasis>
</taxonomicName>
has been treated by some authors as a subspecies of
<taxonomicName id="4C6B4D7DFFEEFFDFFBF9F850FB5FE5D4" baseAuthorityName="Smith" baseAuthorityYear="1847" box="[1061,1258,1997,2022]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="semivariegatus">
<emphasis id="B91FEAECFFEEFFDFFBF9F850FB5FE5D4" box="[1061,1258,1997,2022]" italics="true" pageId="76" pageNumber="77">P. semivariegatus</emphasis>
</taxonomicName>
(Mertens 1934;
<bibRefCitation id="EFFA4B0FFFEFFFDEFF4BFF0AFEF8E283" author="Loveridge, A." box="[151,333,150,177]" pageId="77" pageNumber="78" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridge 1958</bibRefCitation>
), although this last taxon does not form a monophyletic unit and is not closely related to
<taxonomicName id="4C6B4D7DFFEFFFDEFAEEFF0AFA29E283" authorityName="Bocage" authorityYear="1893" box="[1330,1436,151,177]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="girardi">
<emphasis id="B91FEAECFFEFFFDEFAEEFF0AFA29E283" box="[1330,1436,151,177]" italics="true" pageId="77" pageNumber="78">P. girardi</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFFA4B0FFFEFFFDEFF13FF26FE51E2E7" author="Engelbrecht, H. M. &amp; Branch, W. R. &amp; Greenbaum, E. &amp; Alexander, G. J. &amp; Jackson, K. &amp; Burger, M. &amp; Conradie, W. &amp; Kusamba, C. &amp; ZassiBoulou, A-G. &amp; Tolley, K. A." box="[207,484,187,213]" pageId="77" pageNumber="78" pagination="357 - 365" refId="ref76556" refString="Engelbrecht, H. M., Branch, W. R., Greenbaum, E., Alexander, G. J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., ZassiBoulou, A-G. &amp; Tolley, K. A. (2019) Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution, 130, 357 - 365. https: // doi. org / 10.1016 / j. ympev. 2018.10.023" type="journal article" year="2019">
Engelbrecht
<emphasis id="B91FEAECFFEFFFDEFEBDFF26FE28E2E7" box="[353,413,187,213]" italics="true" pageId="77" pageNumber="78">et al.</emphasis>
2019
</bibRefCitation>
).
<taxonomicName id="4C6B4D7DFFEFFFDEFE26FF26FCA5E2E7" authorityName="Bocage" authorityYear="1882" box="[506,784,187,213]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="thomensis">
<emphasis id="B91FEAECFFEFFFDEFE26FF26FCA5E2E7" box="[506,784,187,213]" italics="true" pageId="77" pageNumber="78">Philothamnus thomensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F37C766EFFEFFFDEFC80FF26FBABE2E7" box="[860,1054,187,213]" name="Sao Tome Island" pageId="77" pageNumber="78">São Tomé Island</collectingCountry>
is the sister species to the clade formed by
<taxonomicName id="4C6B4D7DFFEFFFDEFECEFF42FEC9E2CB" authorityName="Bocage" authorityYear="1893" box="[274,380,223,249]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="girardi">
<emphasis id="B91FEAECFFEFFFDEFECEFF42FEC9E2CB" box="[274,380,223,249]" italics="true" pageId="77" pageNumber="78">P. girardi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D7DFFEFFFDEFE6EFF42FCFFE2CB" authority="(Engelbrecht et al. 2019)" baseAuthorityName="Engelbrecht" baseAuthorityYear="2019" box="[434,842,223,249]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="B91FEAECFFEFFFDEFE6EFF42FD9DE2CB" box="[434,552,223,249]" italics="true" pageId="77" pageNumber="78">P. dorsalis</emphasis>
(
<bibRefCitation id="EFFA4B0FFFEFFFDEFDEAFF42FCF6E2CB" author="Engelbrecht, H. M. &amp; Branch, W. R. &amp; Greenbaum, E. &amp; Alexander, G. J. &amp; Jackson, K. &amp; Burger, M. &amp; Conradie, W. &amp; Kusamba, C. &amp; ZassiBoulou, A-G. &amp; Tolley, K. A." box="[566,835,223,249]" pageId="77" pageNumber="78" pagination="357 - 365" refId="ref76556" refString="Engelbrecht, H. M., Branch, W. R., Greenbaum, E., Alexander, G. J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., ZassiBoulou, A-G. &amp; Tolley, K. A. (2019) Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution, 130, 357 - 365. https: // doi. org / 10.1016 / j. ympev. 2018.10.023" type="journal article" year="2019">
Engelbrecht
<emphasis id="B91FEAECFFEFFFDEFD1BFF42FD4AE2CB" box="[711,767,223,249]" italics="true" pageId="77" pageNumber="78">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BD436FEFFEFFFDEFF1BFE9FFEF7E39F" blockId="77.[151,1437,150,429]" pageId="77" pageNumber="78">
<emphasis id="B91FEAECFFEFFFDEFF1BFE9FFE7EE32E" bold="true" box="[199,459,258,284]" pageId="77" pageNumber="78">Specimens examined.</emphasis>
<materialsCitation id="3B033CA3FFEFFFDEFE0AFE9EFE86E373" ID-GBIF-Occurrence="3962324711" accessionNumber="EBD 27758" collectingDate="1987-11" collectionCode="EBD" country="Equatorial Guinea" location="Fourteen" pageId="77" pageNumber="78" specimenCount="1">
<location id="8EB46025FFEFFFDEFE0AFE9EFD8FE32F" LSID="urn:lsid:plazi:treatment:03C287E8FFEEFFDEFF4BF910FEF7E39F:8EB46025FFEFFFDEFE0AFE9EFD8FE32F" box="[470,570,259,285]" country="Equatorial Guinea" name="Fourteen" pageId="77" pageNumber="78">Fourteen</location>
specimens.
<collectingCountry id="F37C766EFFEFFFDEFD17FE9EFC29E32F" box="[715,924,259,285]" name="Equatorial Guinea" pageId="77" pageNumber="78">Equatorial Guinea</collectingCountry>
(without specific locality),
<date id="FFD5103EFFEFFFDEFB3FFE9EFA29E32F" box="[1251,1436,259,285]" pageId="77" pageNumber="78" value="1987-11">
<collectingDate id="EF91E9D6FFEFFFDEFB3FFE9EFA29E32F" box="[1251,1436,259,285]" pageId="77" pageNumber="78" value="1987-11">November 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEFFFDEFF43FEBAFF6CE373" box="[159,217,295,321]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD</collectionCode>
27758)
</materialsCitation>
.
<materialsCitation id="3B033CA3FFEFFFDEFEE0FEBAFF5FE357" ID-GBIF-Occurrence="3962324462" accessionNumber="EBD 27762" collectingDate="1987-11-22" collectionCode="EBD" country="Equatorial Guinea" location="Annobon" pageId="77" pageNumber="78" specimenCode="MNCN 15834, MNCN 23167" specimenCount="2">
<collectingCountry id="F37C766EFFEFFFDEFEE0FEBAFE1CE373" box="[316,425,295,321]" name="Equatorial Guinea" pageId="77" pageNumber="78">Annobon</collectingCountry>
:
<collectingCountry id="F37C766EFFEFFFDEFE6DFEBAFDA9E373" box="[433,540,295,321]" name="Equatorial Guinea" pageId="77" pageNumber="78">Annobon</collectingCountry>
, 1959 [
<specimenCode id="DBCD9E85FFEFFFDEFDB1FEBAFCA1E373" box="[621,788,294,321]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="77" pageNumber="78" type="Museum">MNCN 15834</specimenCode>
,
<specimenCode id="DBCD9E85FFEFFFDEFCC3FEBAFC75E373" box="[799,960,295,321]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="77" pageNumber="78" type="Museum">MNCN 23167</specimenCode>
(
<specimenCount id="9D6DFD77FFEFFFDEFC0CFEBAFBD7E373" box="[976,1122,295,321]" pageId="77" pageNumber="78" type="generic">2 specimens</specimenCount>
)],
<date id="FFD5103EFFEFFFDEFBA7FEBAFAE1E373" box="[1147,1364,295,321]" pageId="77" pageNumber="78" value="1987-11-22">
<collectingDate id="EF91E9D6FFEFFFDEFBA7FEBAFAE1E373" box="[1147,1364,295,321]" pageId="77" pageNumber="78" value="1987-11-22">22 November 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEFFFDEFABFFEBAFA28E373" box="[1379,1437,295,321]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD</collectionCode>
27762)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEFFFDEFF2BFED6FB9FE357" ID-GBIF-Occurrence="3962324613" accessionNumber="EBD 27766, EBD 27757" box="[247,1066,330,357]" country="Equatorial Guinea" location="San Antonio" municipality="November" pageId="77" pageNumber="78" specimenCount="1">
<location id="8EB46025FFEFFFDEFF2BFED6FE3DE357" LSID="urn:lsid:plazi:treatment:03C287E8FFEEFFDEFF4BF910FEF7E39F:8EB46025FFEFFFDEFF2BFED6FE3DE357" box="[247,392,331,357]" country="Equatorial Guinea" municipality="November" name="San Antonio" pageId="77" pageNumber="78">San Antonio</location>
,
<collectingCountry id="F37C766EFFEFFFDEFE4FFED6FE4BE357" box="[403,510,331,357]" name="Equatorial Guinea" pageId="77" pageNumber="78">Annobon</collectingCountry>
, 19
<collectingMunicipality id="6BB0AC84FFEFFFDEFDECFED6FD1DE357" box="[560,680,331,357]" pageId="77" pageNumber="78">November</collectingMunicipality>
, 1987 (EBD 27766, EBD 27757)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEFFFDEFBE9FED6FE78E3BB" ID-GBIF-Occurrence="3962324596" accessionNumber="EBD 27769" collectingDate="1987-11-25" collectionCode="EBD" country="Equatorial Guinea" location="Annobon" pageId="77" pageNumber="78" specimenCode="EBD 27769-27772" specimenCount="1">
<collectingCountry id="F37C766EFFEFFFDEFBE9FED6FB28E357" box="[1077,1181,331,357]" name="Equatorial Guinea" pageId="77" pageNumber="78">Annobon</collectingCountry>
airport,
<date id="FFD5103EFFEFFFDEFADDFED7FF7AE3BB" pageId="77" pageNumber="78" value="1987-11-25">
<collectingDate id="EF91E9D6FFEFFFDEFADDFED7FF7AE3BB" pageId="77" pageNumber="78" value="1987-11-25">25 November 1987</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FFEFFFDEFF3DFEF2FE70E3BB" box="[225,453,367,393]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD 27769–27772</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEFFFDEFE07FEF2FBD7E3BB" ID-GBIF-Occurrence="3962324660" accessionNumber="EBD 27759" box="[475,1122,366,393]" collectingDate="1987-11-11" collectionCode="EBD" country="San Pedro, Annobon" location="San Pedro, Annobon" pageId="77" pageNumber="78" specimenCount="1">
<collectingCountry id="F37C766EFFEFFFDEFE07FEF2FD79E3BB" box="[475,716,367,393]" isEnumeration="true" pageId="77" pageNumber="78">San Pedro, Annobon</collectingCountry>
,
<date id="FFD5103EFFEFFFDEFD06FEF2FC0CE3BB" box="[730,953,367,393]" pageId="77" pageNumber="78" value="1987-11-11">
<collectingDate id="EF91E9D6FFEFFFDEFD06FEF2FC0CE3BB" box="[730,953,367,393]" pageId="77" pageNumber="78" value="1987-11-11">11 November 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEFFFDEFC17FEF2FBB0E3BB" box="[971,1029,367,393]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD</collectionCode>
27759)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEFFFDEFBACFEF2FE8AE39F" ID-GBIF-Occurrence="3962324673" accessionNumber="EBD 27760" collectionCode="EBD" country="Equatorial Guinea" location="San Juan" pageId="77" pageNumber="78" specimenCode="EBD 27760-27761" specimenCount="1">
<location id="8EB46025FFEFFFDEFBACFEF2FB6EE3BB" LSID="urn:lsid:plazi:treatment:03C287E8FFEEFFDEFF4BF910FEF7E39F:8EB46025FFEFFFDEFBACFEF2FB6EE3BB" box="[1136,1243,367,393]" country="Equatorial Guinea" name="San Juan" pageId="77" pageNumber="78">San Juan</location>
,
<collectingCountry id="F37C766EFFEFFFDEFB34FEF2FAE5E3BB" box="[1256,1360,367,393]" name="Equatorial Guinea" pageId="77" pageNumber="78">Annobon</collectingCountry>
(
<specimenCode id="DBCD9E85FFEFFFDEFABFFEF2FE82E39F" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD 27760–27761</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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