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<treatment id="03C287E8FF87FFB6FF4BFE1AFAC9E5C6" ID-DOI="http://doi.org/10.5281/zenodo.7305358" ID-GBIF-Taxon="204305608" ID-Zenodo-Dep="7305358" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6" httpUri="http://treatment.plazi.org/id/03C287E8FF87FFB6FF4BFE1AFAC9E5C6" lastPageNumber="38" pageId="37" pageNumber="38" scope_family="Gekkonidae" scope_order="Squamata">
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<taxonomicName id="4C6B4D7DFF87FFB6FF4BFE1AFD4DE390" ID-CoL="6LPQG" authority="(Moreau de Jonnes, 1818)" baseAuthorityName="Moreau de Jonnes" baseAuthorityYear="1818" box="[151,760,391,418]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mabouia">
<emphasis id="B91FEAECFF87FFB6FF4BFE1AFD4DE390" bold="true" box="[151,760,391,418]" pageId="37" pageNumber="38">
<emphasis id="B91FEAECFF87FFB6FF4BFE1AFE1AE390" bold="true" box="[151,431,391,418]" italics="true" pageId="37" pageNumber="38">Hemidactylus mabouia</emphasis>
(
<bibRefCitation id="EFFA4B0FFF87FFB6FE62FE15FD45E390" author="Moreau de Jonnes, A." box="[446,752,391,418]" pageId="37" pageNumber="38" pagination="138 - 139" refId="ref85104" refString="Moreau de Jonnes, A. (1818) Monographie du Mabouia des murailles, ou Gecko Mabouia des Antilles. Bulletin des Sciences, Societe Philomatique, Paris, 3 (5), 138 - 139." type="journal article" year="1818">Moreau de Jonnès, 1818</bibRefCitation>
)
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<figureCitation id="13502A7BFF87FFB6FF4BFE31FEBCE3F4" box="[151,265,428,454]" captionStart="FIGURE 9" captionStartId="38.[151,250,1902,1926]" captionTargetBox="[184,1388,200,1863]" captionTargetId="figure-19@38.[157,1430,181,1878]" captionTargetPageId="38" captionText="FIGURE 9. From left to right: dorsal side of the trunk, ventral view of hind legs and tail, and ventral view of the head of Hemidactylus mabouia (A) (EBD 2746); and dorsal, lateral, and ventral views of the head of Lygodactylus thomensis (B) (MNCN 46132), and Lacertaspis gemmiventris (C) (EBD 31523) showing a marbled pattern. Photos: AS-V." figureDoi="http://doi.org/10.5281/zenodo.7285637" httpUri="https://zenodo.org/record/7285637/files/figure.png" pageId="37" pageNumber="38">Figure 9A</figureCitation>
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<treatmentCitation id="0ACA10EFFF87FFB6FF4BFE76FDC4E031" author="Moreau de Jonnes, A." box="[151,625,491,515]" page="138" pageId="37" pageNumber="38" year="1818">
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<emphasis id="B91FEAECFF87FFB6FF4BFE76FE8DE031" box="[151,312,491,515]" italics="true" pageId="37" pageNumber="38">Gekko mabouia</emphasis>
<bibRefCitation id="EFFA4B0FFF87FFB6FEE3FE71FDC4E031" author="Moreau de Jonnes, A." box="[319,625,491,515]" pageId="37" pageNumber="38" pagination="138 - 139" refId="ref85104" refString="Moreau de Jonnes, A. (1818) Monographie du Mabouia des murailles, ou Gecko Mabouia des Antilles. Bulletin des Sciences, Societe Philomatique, Paris, 3 (5), 138 - 139." type="journal article" year="1818">Moreau de Jonnès, 1818: 138</bibRefCitation>
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.
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locality: “en Amérique, dans les contrées continentales qui avoisinent au midi larchipel des Antilles, et quil est également répandu dans les îles même de larchipel, depuis la Trinité jusquà la
</treatmentCitationGroup>
<collectingCountry id="F37C766EFF87FFB6FF1BFDB4FE99E073" box="[199,300,553,577]" name="Jamaica" pageId="37" pageNumber="38">Jamaïque</collectingCountry>
”, restricted to
<collectingCountry id="F37C766EFF87FFB6FE61FDB4FD9BE072" box="[445,558,552,577]" name="Saint Vincent and the Grenadines" pageId="37" pageNumber="38">St. Vincent</collectingCountry>
in the Lesser Antilles by
<bibRefCitation id="EFFA4B0FFF87FFB6FCECFDB5FC68E073" author="Stejneger, L." box="[816,989,552,577]" pageId="37" pageNumber="38" pagination="549 - 724" refId="ref89945" refString="Stejneger, L. (1904) The herpetology of Porto Rico. Report of the United States National Museum, 1902, 549 - 724. https: // doi. org / 10.5962 / bhl. title. 11835" type="journal article" year="1904">Stejneger (1904)</bibRefCitation>
.
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<taxonomicName id="4C6B4D7DFF87FFB6FF4BFDDAFDDBE06D" authority="Wied-Neuwied, 1824: 101" authorityName="Wied-Neuwied" authorityPageNumber="101" authorityYear="1824" box="[151,622,583,607]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="incanescens">
<emphasis id="B91FEAECFF87FFB6FF4BFDDAFEEDE06D" box="[151,344,583,607]" italics="true" pageId="37" pageNumber="38">Gekko incanescens</emphasis>
<bibRefCitation id="EFFA4B0FFF87FFB6FE82FDDAFDDBE06D" author="Wied-Neuwied, M." box="[350,622,583,607]" pageId="37" pageNumber="38" pagination="661 - 673" refId="ref92366" refString="Wied-Neuwied, M. (1824) Verzeichniss der Amphibien, welche im zweyten Bande der Naturgeschichte Brasiliens vom Prinz Max von Neuwied werden beschrieben werden. Isis von Oken, 14, columns 661 - 673." type="journal article" year="1824">Wied-Neuwied, 1824: 101</bibRefCitation>
</taxonomicName>
.
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locality:
<collectingCountry id="F37C766EFF87FFB6FCD5FDDAFCFFE06D" box="[777,842,583,607]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
.
</paragraph>
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<emphasis id="B91FEAECFF87FFB6FF4BFDFBFEEAE04C" box="[151,351,614,638]" italics="true" pageId="37" pageNumber="38">Gecko tuberculosus</emphasis>
<bibRefCitation id="EFFA4B0FFF87FFB6FEB9FDFBFDBFE04C" author="Raddi, G." box="[357,522,614,638]" pageId="37" pageNumber="38" pagination="58 - 73" refId="ref87423" refString="Raddi, G. (1823) Continuazione della descrizione dei Rettili Brasiliani. Accademia Nazionale dei Quaranta, Memorie delle Societa Italiana Scienze, Modena, Italy, 19, 58 - 73." type="journal article" year="1823">Raddi, 1823: 63</bibRefCitation>
.
<typeStatus id="54D0885CFF87FFB6FDC8FDFBFDF3E04C" box="[532,582,614,638]" pageId="37" pageNumber="38">Type</typeStatus>
locality: warmer climates throughout the world, even on remote oceanic islands.
</paragraph>
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<taxonomicName id="4C6B4D7DFF87FFB6FF4BFD19FDF2E0AF" authority="Wied-Neuwied, 1824: 104" authorityName="Wied-Neuwied" authorityPageNumber="104" authorityYear="1824" box="[151,583,644,669]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="armatus">
<emphasis id="B91FEAECFF87FFB6FF4BFD19FE84E0AE" box="[151,305,644,668]" italics="true" pageId="37" pageNumber="38">Gekko armatus</emphasis>
<bibRefCitation id="EFFA4B0FFF87FFB6FEEBFD19FDF2E0AF" author="Wied-Neuwied, M." box="[311,583,644,669]" pageId="37" pageNumber="38" pagination="661 - 673" refId="ref92366" refString="Wied-Neuwied, M. (1824) Verzeichniss der Amphibien, welche im zweyten Bande der Naturgeschichte Brasiliens vom Prinz Max von Neuwied werden beschrieben werden. Isis von Oken, 14, columns 661 - 673." type="journal article" year="1824">Wied-Neuwied, 1824: 104</bibRefCitation>
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.
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locality:
<collectingCountry id="F37C766EFF87FFB6FD3EFD19FC96E0AE" box="[738,803,644,668]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8BD436FEFF87FFB6FF4BFD3EFCE8E089" blockId="37.[151,1436,491,792]" box="[151,861,675,700]" pageId="37" pageNumber="38">
<emphasis id="B91FEAECFF87FFB6FF4BFD3EFE8AE089" box="[151,319,675,699]" italics="true" pageId="37" pageNumber="38">Gecko aculeatus</emphasis>
<bibRefCitation id="EFFA4B0FFF87FFB6FE9AFD3EFE68E089" author="Spix, J. B." box="[326,477,675,699]" pageId="37" pageNumber="38" refId="ref89841" refString="Spix, J. B. (1825) Animalia Nova Sive Species Nova Lacertarum Quas in Itinere per Brasiliam Annis MDCCCXVII-MDCCCXX Jussu et Auspicius Maximiliani Josephi I. Bavariae Regis Suscepto Collegit et Descripsit Dr. J. B. de Spix. Hubschmanni, Monachii [Munchen], 26 pp., 28 pls. https: // doi. org / 10.5962 / bhl. title. 5117" type="book" year="1825">Spix, 1825: 16</bibRefCitation>
.
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locality:
<collectingRegion id="49AFF81CFF87FFB6FDA4FD3EFCBBE089" box="[632,782,675,699]" country="Brazil" name="Rio de Janeiro" pageId="37" pageNumber="38">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="F37C766EFF87FFB6FCCBFD3EFCEDE089" box="[791,856,675,699]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8BD436FEFF87FFB6FF4BFD5FFD4FE0E8" blockId="37.[151,1436,491,792]" box="[151,762,706,730]" pageId="37" pageNumber="38">
<emphasis id="B91FEAECFF87FFB6FF4BFD5FFE81E0EB" box="[151,308,706,730]" italics="true" pageId="37" pageNumber="38">Gecko cruciger</emphasis>
<bibRefCitation id="EFFA4B0FFF87FFB6FEE6FD5FFE67E0E8" author="Spix, J. B." box="[314,466,706,730]" pageId="37" pageNumber="38" refId="ref89841" refString="Spix, J. B. (1825) Animalia Nova Sive Species Nova Lacertarum Quas in Itinere per Brasiliam Annis MDCCCXVII-MDCCCXX Jussu et Auspicius Maximiliani Josephi I. Bavariae Regis Suscepto Collegit et Descripsit Dr. J. B. de Spix. Hubschmanni, Monachii [Munchen], 26 pp., 28 pls. https: // doi. org / 10.5962 / bhl. title. 5117" type="book" year="1825">Spix, 1825: 16</bibRefCitation>
.
<typeStatus id="54D0885CFF87FFB6FE07FD5FFDB8E0E8" box="[475,525,706,730]" pageId="37" pageNumber="38">Type</typeStatus>
locality:
<collectingRegion id="49AFF81CFF87FFB6FDB1FD5FFD1EE0E8" box="[621,683,706,730]" country="Brazil" name="Bahia" pageId="37" pageNumber="38">Bahia</collectingRegion>
,
<collectingCountry id="F37C766EFF87FFB6FD69FD5FFD43E0E8" box="[693,758,706,730]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8BD436FEFF87FFB6FF4BFD7DFC79E0CA" blockId="37.[151,1436,491,792]" box="[151,972,736,761]" pageId="37" pageNumber="38">
<taxonomicName id="4C6B4D7DFF87FFB6FF4BFD7DFD21E0CB" authority="(Moreau de Jonnes, 1818)" baseAuthorityName="Moreau de Jonnes" baseAuthorityYear="1818" box="[151,660,736,761]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mabouia">
<emphasis id="B91FEAECFF87FFB6FF4BFD7DFE37E0CA" box="[151,386,736,760]" italics="true" pageId="37" pageNumber="38">Hemidactylus mabouia</emphasis>
(
<bibRefCitation id="EFFA4B0FFF87FFB6FE53FD7CFD39E0CB" author="Moreau de Jonnes, A." box="[399,652,736,761]" pageId="37" pageNumber="38" pagination="138 - 139" refId="ref85104" refString="Moreau de Jonnes, A. (1818) Monographie du Mabouia des murailles, ou Gecko Mabouia des Antilles. Bulletin des Sciences, Societe Philomatique, Paris, 3 (5), 138 - 139." type="journal article" year="1818">Moreau de Jonnès, 1818</bibRefCitation>
)
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:
<bibRefCitation id="EFFA4B0FFF87FFB6FD42FD7DFC7DE0CA" author="Dumeril, A. M. C. &amp; Bibron, G." box="[670,968,736,761]" pageId="37" pageNumber="38" refId="ref75664" refString="Dumeril, A. M. C. &amp; Bibron, G. (1836) Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Vol. 3. Encyclopedique Roret, Paris, iv + 518 pp." type="book" year="1836">Duméril &amp; Bibron 1836: 362</bibRefCitation>
.
</paragraph>
<paragraph id="8BD436FEFF87FFB6FF4BFD62FBE6E125" blockId="37.[151,1436,491,792]" box="[151,1107,767,792]" pageId="37" pageNumber="38">
<taxonomicName id="4C6B4D7DFF87FFB6FF4BFD62FD69E125" authority="Boettger, 1878: 1" authorityName="Boettger" authorityPageNumber="1" authorityYear="1878" box="[151,732,767,792]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="variety" species="frenatus" variety="calabaricus">
<emphasis id="B91FEAECFF87FFB6FF4BFD62FECEE125" box="[151,379,767,791]" italics="true" pageId="37" pageNumber="38">Hemidactylus frenatus</emphasis>
var.
<emphasis id="B91FEAECFF87FFB6FE71FD62FD90E125" box="[429,549,767,791]" italics="true" pageId="37" pageNumber="38">calabaricus</emphasis>
Boettger, 1878: 1
</taxonomicName>
.
<typeStatus id="54D0885CFF87FFB6FD3BFC9DFCACE12A" box="[743,793,768,792]" pageId="37" pageNumber="38">Type</typeStatus>
locality: Old Calabar,
<collectingCountry id="F37C766EFF87FFB6FC23FD62FBFBE125" box="[1023,1102,767,791]" name="Nigeria" pageId="37" pageNumber="38">Nigeria</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716575FF87FFB6FF4BFCDDFCD9E1F6" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8BD436FEFF87FFB6FF4BFCDDFCD9E1F6" blockId="37.[151,1437,832,2036]" pageId="37" pageNumber="38">
<emphasis id="B91FEAECFF87FFB6FF4BFCDDFE9CE168" bold="true" box="[151,297,832,858]" pageId="37" pageNumber="38">Distribution</emphasis>
. Widely distributed in tropical areas both in the Old and New Worlds, including some Atlantic islands. In
<collectingCountry id="F37C766EFF87FFB6FF65FCFEFE3FE14F" box="[185,394,867,893]" name="Equatorial Guinea" pageId="37" pageNumber="38">Equatorial Guinea</collectingCountry>
it has been recorded in the islands of Bioko (at Malabo) (
<bibRefCitation id="EFFA4B0FFF87FFB6FBE1FCF9FAD7E14F" author="Carranza, S. &amp; Arnold, E. N." box="[1085,1378,867,894]" pageId="37" pageNumber="38" pagination="531 - 545" refId="ref73406" refString="Carranza, S. &amp; Arnold, E. N. (2006) Systematics, biogeography, and evolution of Hemidactylus geckos (Reptilia: Gekkonidae) elucidated using mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 38 (2), 531 - 545. https: // doi. org / 10.1016 / j. ympev. 2005.07.012" type="journal article" year="2006">Carranza &amp; Arnold 2006</bibRefCitation>
) and
<collectingCountry id="F37C766EFF87FFB6FF4BFC1AFF4AE193" box="[151,255,903,929]" name="Equatorial Guinea" pageId="37" pageNumber="38">Annobon</collectingCountry>
(
<bibRefCitation id="EFFA4B0FFF87FFB6FED2FC1AFE7FE193" author="Jesus, J. &amp; Brehm, A. &amp; Harris, D. J." box="[270,458,903,929]" pageId="37" pageNumber="38" pagination="20 - 22" refId="ref80654" refString="Jesus, J., Brehm, A. &amp; Harris, D. J. (2003) The herpetofauna of Annobon island, Gulf of Guinea, West Africa. Herpetological Bulletin, 86, 20 - 22." type="journal article" year="2003">
Jesus
<emphasis id="B91FEAECFF87FFB6FE8DFC15FE3CE193" box="[337,393,903,929]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2003
</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFF87FFB6FE09FC1AFD28E190" author="Rocha, S. &amp; Carretero, M. A. &amp; Harris, D. J." box="[469,669,903,930]" pageId="37" pageNumber="38" pagination="292 - 299" refId="ref87895" refString="Rocha, S., Carretero, M. A. &amp; Harris, D. J. (2005) Diversity and phylogenetic relationships of Hemidactylus geckos from the Comoro islands. Molecular Phylogenetics and Evolution, 35, 292 - 299. https: // doi. org / 10.1016 / j. ympev. 2004.11.023" type="journal article" year="2005">
Rocha
<emphasis id="B91FEAECFF87FFB6FDFFFC15FDEEE193" box="[547,603,903,929]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2005
</bibRefCitation>
), and in
<collectingCountry id="F37C766EFF87FFB6FD27FC1AFCD2E193" box="[763,871,903,929]" name="Equatorial Guinea" pageId="37" pageNumber="38">Río Muni</collectingCountry>
at the village of Sendje in Monte Mitra, within the Monte Alén National Park (
<bibRefCitation id="EFFA4B0FFF87FFB6FE10FC37FD5CE1F7" author="Gonwouo, N. L. &amp; Nsang, A." box="[460,745,938,965]" pageId="37" pageNumber="38" pagination="32 - 49" refId="ref78060" refString="Gonwouo, N. L. &amp; Nsang, A. (2005) An assessment of the reptiles and amphibians of the Monte Mitra forest, Monte Alen National Park. In: Sunderland, T. C. H. (Ed.), A Biodiversity Assessment of the Monte Mitra Forest, Monte Alen National Park, Equatorial Guinea. Smithsonian Institution, Washington, D. C., pp. 32 - 49." type="book chapter" year="2005">Gonwouo &amp; Nsang 2005</bibRefCitation>
) (
<figureCitation id="13502A7BFF87FFB6FD23FC37FCEBE1F6" box="[767,862,938,964]" captionStart="MAPS 8" captionStartId="39.[151,223,1943,1967]" captionTargetBox="[202,1383,192,1906]" captionTargetId="figure-18@39.[159,1427,181,1919]" captionTargetPageId="39" captionText="MAPS 8 AC. Distribution maps for Equatorial Guinean records of (A) Hemidactlyus aff. longicephalus (question mark represents a dubious record; see species account); (B) Hemidactylus mabouia; (C) Hemidactylus muriceus." figureDoi="http://doi.org/10.5281/zenodo.7304647" httpUri="https://zenodo.org/record/7304647/files/figure.png" pageId="37" pageNumber="38">Map 8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716575FF87FFB6FF1BFC53FAC9E5C6" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BD436FEFF87FFB6FF1BFC53FE6AE529" blockId="37.[151,1437,832,2036]" pageId="37" pageNumber="38">
<emphasis id="B91FEAECFF87FFB6FF1BFC53FEFAE1DA" bold="true" box="[199,335,974,1000]" pageId="37" pageNumber="38">Comments.</emphasis>
One of the most widespread species of the genus, as it has been introduced in several tropical regions. Likely originated in East Africa. Based on mtDNA (12S and CytB gene fragments) this species is very uniform across most of its distribution, which encompass tropical Africa, including Bioko (Malabo), South America, North America (Florida) and some Caribbean areas (
<bibRefCitation id="EFFA4B0FFF87FFB6FD3AFBA5FBB7E660" author="Carranza, S. &amp; Arnold, E. N." box="[742,1026,1080,1106]" pageId="37" pageNumber="38" pagination="531 - 545" refId="ref73406" refString="Carranza, S. &amp; Arnold, E. N. (2006) Systematics, biogeography, and evolution of Hemidactylus geckos (Reptilia: Gekkonidae) elucidated using mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 38 (2), 531 - 545. https: // doi. org / 10.1016 / j. ympev. 2005.07.012" type="journal article" year="2006">Carranza &amp; Arnold 2006</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFF87FFB6FBD1FBA4FB70E660" author="Rato, C. &amp; Martins, B. &amp; Rocha, R. &amp; Silva-Rocha, I." box="[1037,1221,1080,1107]" pageId="37" pageNumber="38" pagination="369 - 375" refId="ref87603" refString="Rato, C., Martins, B., Rocha, R. &amp; Silva-Rocha, I. (2021) Uncovered genetic diversity in Hemidactylus mabouia (Reptilia: Gekkonidae) from Madeira Island reveals uncertain sources of introduction. Amphibia-Reptilia, 42 (3), 369 - 375. https: // doi. org / 10.1163 / 15685381 - bja 10052" type="journal article" year="2021">
Rato
<emphasis id="B91FEAECFF87FFB6FB96FBA4FB37E660" box="[1098,1154,1080,1106]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2021
</bibRefCitation>
). However, despite the widespread genetic uniformity of
<taxonomicName id="4C6B4D7DFF87FFB6FD98FBC1FD7FE644" baseAuthorityName="Moreau de Jonnes" baseAuthorityYear="1818" box="[580,714,1116,1142]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mabouia">
<emphasis id="B91FEAECFF87FFB6FD98FBC1FD7FE644" box="[580,714,1116,1142]" italics="true" pageId="37" pageNumber="38">H. mabouia</emphasis>
</taxonomicName>
in most of its range, deep lineages and cryptic diversity across the eastern coast of Africa have been documented (
<bibRefCitation id="EFFA4B0FFF87FFB6FD01FBE2FC78E6AB" author="Agarwal, I. &amp; Ceriaco, L. M. P. &amp; Metallinou, M. &amp; Jackman, T. R. &amp; Bauer, A. M." box="[733,973,1151,1178]" pageId="37" pageNumber="38" pagination="1 - 14" refId="ref69579" refString="Agarwal, I., Ceriaco, L. M. P., Metallinou, M., Jackman, T. R. &amp; Bauer, A. M. (2021) How the African house gecko (Hemidactylus mabouia) conquered the world. Royal Society Open Science, 8 (210749), 1 - 14. https: // doi. org / 10.1098 / rsos. 210749" type="journal article" year="2021">
Agarwal
<emphasis id="B91FEAECFF87FFB6FC94FB1DFC37E6AB" box="[840,898,1151,1177]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
, 2021
</bibRefCitation>
). The taxonomic status and relationships between some populations formerly recorded as
<taxonomicName id="4C6B4D7DFF87FFB6FD12FB3EFCE3E68F" baseAuthorityName="Moreau de Jonnes" baseAuthorityYear="1818" box="[718,854,1187,1213]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mabouia">
<emphasis id="B91FEAECFF87FFB6FD12FB3EFCE3E68F" box="[718,854,1187,1213]" italics="true" pageId="37" pageNumber="38">H. mabouia</emphasis>
</taxonomicName>
and the Malagasy
<taxonomicName id="4C6B4D7DFF87FFB6FBE4FB3EFADAE68F" authority="Gray, 1842" authorityName="Gray" authorityYear="1842" box="[1080,1391,1187,1213]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mercatorius">
<emphasis id="B91FEAECFF87FFB6FBE4FB3EFB51E68E" box="[1080,1252,1187,1212]" italics="true" pageId="37" pageNumber="38">H. mercatorius</emphasis>
Gray, 1842
</taxonomicName>
are in need of revision (
<bibRefCitation id="EFFA4B0FFF87FFB6FEA2FB5BFDFBE6D3" author="Rocha, S. &amp; Carretero, M. A. &amp; Harris, D. J." box="[382,590,1222,1249]" pageId="37" pageNumber="38" pagination="292 - 299" refId="ref87895" refString="Rocha, S., Carretero, M. A. &amp; Harris, D. J. (2005) Diversity and phylogenetic relationships of Hemidactylus geckos from the Comoro islands. Molecular Phylogenetics and Evolution, 35, 292 - 299. https: // doi. org / 10.1016 / j. ympev. 2004.11.023" type="journal article" year="2005">
Rocha
<emphasis id="B91FEAECFF87FFB6FE0CFB5AFDBEE6D2" box="[464,523,1222,1248]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2005
</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFF87FFB6FD80FB5BFC83E6D2" author="Rocha, S. &amp; Carretero, M. A. &amp; Harris, D. J." box="[604,822,1222,1248]" pageId="37" pageNumber="38" pagination="83 - 89" refId="ref87953" refString="Rocha, S., Carretero, M. A. &amp; Harris, D. J. (2010 a) On the diversity, colonization patterns and status of Hemidactylus spp. (Reptilia: Gekkonidae) from the Western Indian Ocean islands. The Herpetological Journal, 20, 83 - 89." type="journal article" year="2010">
Rocha
<emphasis id="B91FEAECFF87FFB6FD70FB5AFD53E6D2" box="[684,742,1222,1248]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2010a
</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFF87FFB6FC9EFB5BFB9FE6D2" author="Agarwal, I. &amp; Ceriaco, L. M. P. &amp; Metallinou, M. &amp; Jackman, T. R. &amp; Bauer, A. M." box="[834,1066,1222,1248]" pageId="37" pageNumber="38" pagination="1 - 14" refId="ref69579" refString="Agarwal, I., Ceriaco, L. M. P., Metallinou, M., Jackman, T. R. &amp; Bauer, A. M. (2021) How the African house gecko (Hemidactylus mabouia) conquered the world. Royal Society Open Science, 8 (210749), 1 - 14. https: // doi. org / 10.1098 / rsos. 210749" type="journal article" year="2021">
Agarwal
<emphasis id="B91FEAECFF87FFB6FC70FB5AFC53E6D2" box="[940,998,1222,1248]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2021
</bibRefCitation>
). Within this context, the results obtained by
<bibRefCitation id="EFFA4B0FFF87FFB6FEF9FB74FDB7E736" author="Rocha, S. &amp; Carretero, M. A. &amp; Harris, D. J." box="[293,514,1257,1284]" pageId="37" pageNumber="38" pagination="292 - 299" refId="ref87895" refString="Rocha, S., Carretero, M. A. &amp; Harris, D. J. (2005) Diversity and phylogenetic relationships of Hemidactylus geckos from the Comoro islands. Molecular Phylogenetics and Evolution, 35, 292 - 299. https: // doi. org / 10.1016 / j. ympev. 2004.11.023" type="journal article" year="2005">
Rocha
<emphasis id="B91FEAECFF87FFB6FEA9FB77FE1AE731" box="[373,431,1257,1283]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
(2005)
</bibRefCitation>
and
<bibRefCitation id="EFFA4B0FFF87FFB6FDE7FB74FC91E736" author="Rocha, S. &amp; Carretero, M. A. &amp; Harris, D. J." box="[571,804,1257,1284]" pageId="37" pageNumber="38" pagination="83 - 89" refId="ref87953" refString="Rocha, S., Carretero, M. A. &amp; Harris, D. J. (2010 a) On the diversity, colonization patterns and status of Hemidactylus spp. (Reptilia: Gekkonidae) from the Western Indian Ocean islands. The Herpetological Journal, 20, 83 - 89." type="journal article" year="2010">
Rocha
<emphasis id="B91FEAECFF87FFB6FD57FB77FD70E731" box="[651,709,1257,1283]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
(2010a)
</bibRefCitation>
based on mtDNA (16S gene fragment) and nuDNA (cmos gene fragment), showed that populations from
<collectingCountry id="F37C766EFF87FFB6FD3CFA90FCFDE715" box="[736,840,1293,1319]" name="Equatorial Guinea" pageId="37" pageNumber="38">Annobon</collectingCountry>
and some other Gulf of
<collectingCountry id="F37C766EFF87FFB6FBBDFA90FB07E715" box="[1121,1202,1293,1319]" name="Guinea-Bissau" pageId="37" pageNumber="38">Guinea</collectingCountry>
islands, such as
<collectingCountry id="F37C766EFF87FFB6FAAEFA90FED9E778" name="Sao Tome and Principe" pageId="37" pageNumber="38">São Tomé and Príncipe</collectingCountry>
, are nested within a clade formed by populations of
<taxonomicName id="4C6B4D7DFF87FFB6FC6CFAACFBE2E778" authorityName="Gray" authorityYear="1842" box="[944,1111,1329,1354]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mercatorius">
<emphasis id="B91FEAECFF87FFB6FC6CFAACFBE2E778" box="[944,1111,1329,1354]" italics="true" pageId="37" pageNumber="38">H. mercatorius</emphasis>
</taxonomicName>
from
<collectingCountry id="F37C766EFF87FFB6FB47FAADFA91E778" box="[1179,1316,1328,1354]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
,
<collectingCountry id="F37C766EFF87FFB6FAF3FAACFA2CE779" box="[1327,1433,1329,1355]" name="Comoros" pageId="37" pageNumber="38">Comoros</collectingCountry>
,
<collectingCountry id="F37C766EFF87FFB6FF4BFAC9FF44E75C" box="[151,241,1364,1390]" name="Uganda" pageId="37" pageNumber="38">Uganda</collectingCountry>
, and the Islands of Mahé (
<collectingCountry id="F37C766EFF87FFB6FDC0FAC9FD2CE75C" box="[540,665,1364,1390]" name="Seychelles" pageId="37" pageNumber="38">Seychelles</collectingCountry>
),
<collectingCountry id="F37C766EFF87FFB6FD71FAC9FCA4E75C" box="[685,785,1364,1390]" name="Tanzania" pageId="37" pageNumber="38">Zanzibar</collectingCountry>
and
<collectingRegion id="49AFF81CFF87FFB6FC94FAC9FC21E75C" box="[840,916,1364,1390]" country="Tanzania" name="Kusini Pemba" pageId="37" pageNumber="38">Pemba</collectingRegion>
(
<collectingCountry id="F37C766EFF87FFB6FC78FAC9FBA5E75C" box="[932,1040,1364,1390]" name="Tanzania" pageId="37" pageNumber="38">Tanzania</collectingCountry>
). As
<taxonomicName id="4C6B4D7DFF87FFB6FB9BFAC9FADAE75C" authorityName="Gray" authorityYear="1842" box="[1095,1391,1364,1390]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mercatorius">
<emphasis id="B91FEAECFF87FFB6FB9BFAC9FADAE75C" box="[1095,1391,1364,1390]" italics="true" pageId="37" pageNumber="38">Hemidactylus mercatorius</emphasis>
</taxonomicName>
has been described from northern
<collectingCountry id="F37C766EFF87FFB6FE2AFAEAFDC8E7A3" box="[502,637,1399,1425]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
[restricted to
<collectingCountry id="F37C766EFF87FFB6FCFFFAEAFC2EE7A3" box="[803,923,1399,1425]" name="Seychelles" pageId="37" pageNumber="38">Seychelles</collectingCountry>
by
<bibRefCitation id="EFFA4B0FFF87FFB6FC17FAEAFB2DE7A0" author="Loveridge, A." box="[971,1176,1399,1426]" pageId="37" pageNumber="38" pagination="1 - 469" refId="ref83041" refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469." type="journal article" year="1947">Loveridge (1947)</bibRefCitation>
],
<bibRefCitation id="EFFA4B0FFF87FFB6FB72FAEAFA29E7A0" author="Rocha, S. &amp; Carretero, M. A. &amp; Harris, D. J." box="[1198,1436,1399,1426]" pageId="37" pageNumber="38" pagination="83 - 89" refId="ref87953" refString="Rocha, S., Carretero, M. A. &amp; Harris, D. J. (2010 a) On the diversity, colonization patterns and status of Hemidactylus spp. (Reptilia: Gekkonidae) from the Western Indian Ocean islands. The Herpetological Journal, 20, 83 - 89." type="journal article" year="2010">
Rocha
<emphasis id="B91FEAECFF87FFB6FADCFAE5FA8EE7A3" box="[1280,1339,1399,1425]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
(2010a)
</bibRefCitation>
suggested the use of
<taxonomicName id="4C6B4D7DFF87FFB6FE5FFA01FD99E787" authorityName="Gray" authorityYear="1842" box="[387,556,1436,1461]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mercatorius">
<emphasis id="B91FEAECFF87FFB6FE5FFA01FD99E787" box="[387,556,1436,1461]" italics="true" pageId="37" pageNumber="38">H. mercatorius</emphasis>
</taxonomicName>
for the populations of the islands of the Gulf of
<collectingCountry id="F37C766EFF87FFB6FB8FFA06FB11E787" box="[1107,1188,1435,1461]" name="Guinea-Bissau" pageId="37" pageNumber="38">Guinea</collectingCountry>
(
<collectingCountry id="F37C766EFF87FFB6FB6FFA06FA95E787" box="[1203,1312,1435,1461]" name="Equatorial Guinea" pageId="37" pageNumber="38">Annobon</collectingCountry>
,
<collectingCountry id="F37C766EFF87FFB6FAF0FA06FE99E7EA" name="Sao Tome and Principe" pageId="37" pageNumber="38">São Tomé and Príncipe</collectingCountry>
), leaving the status of continental populations pending further studies. By contrast, the specimens from Bioko (Malabo) and
<collectingCountry id="F37C766EFF87FFB6FE15FA7FFD61E7CE" box="[457,724,1506,1532]" name="Sao Tome and Principe" pageId="37" pageNumber="38">São Tomé and Príncipe</collectingCountry>
sequenced by
<bibRefCitation id="EFFA4B0FFF87FFB6FC5EFA7FFB07E7CE" author="Carranza, S. &amp; Arnold, E. N." box="[898,1202,1506,1532]" pageId="37" pageNumber="38" pagination="531 - 545" refId="ref73406" refString="Carranza, S. &amp; Arnold, E. N. (2006) Systematics, biogeography, and evolution of Hemidactylus geckos (Reptilia: Gekkonidae) elucidated using mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 38 (2), 531 - 545. https: // doi. org / 10.1016 / j. ympev. 2005.07.012" type="journal article" year="2006">Carranza &amp; Arnold (2006)</bibRefCitation>
did not cluster with two Malagasy samples of
<taxonomicName id="4C6B4D7DFF87FFB6FE14F99BFDC6E42D" authorityName="Gray" authorityYear="1842" box="[456,627,1542,1567]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mercatorius">
<emphasis id="B91FEAECFF87FFB6FE14F99BFDC6E42D" box="[456,627,1542,1567]" italics="true" pageId="37" pageNumber="38">H. mercatorius</emphasis>
</taxonomicName>
, thus considering the Atlantic populations as
<taxonomicName id="4C6B4D7DFF87FFB6FB5BF99BFF6EE471" authority="(Rato et al. 2021)" baseAuthorityName="Rato" baseAuthorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mabouia">
<emphasis id="B91FEAECFF87FFB6FB5BF99BFABAE42D" box="[1159,1295,1541,1567]" italics="true" pageId="37" pageNumber="38">H. mabouia</emphasis>
(
<bibRefCitation id="EFFA4B0FFF87FFB6FAFCF998FF66E471" author="Rato, C. &amp; Martins, B. &amp; Rocha, R. &amp; Silva-Rocha, I." pageId="37" pageNumber="38" pagination="369 - 375" refId="ref87603" refString="Rato, C., Martins, B., Rocha, R. &amp; Silva-Rocha, I. (2021) Uncovered genetic diversity in Hemidactylus mabouia (Reptilia: Gekkonidae) from Madeira Island reveals uncertain sources of introduction. Amphibia-Reptilia, 42 (3), 369 - 375. https: // doi. org / 10.1163 / 15685381 - bja 10052" type="journal article" year="2021">
Rato
<emphasis id="B91FEAECFF87FFB6FABDF99BFA28E42D" box="[1377,1437,1541,1567]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2021
</bibRefCitation>
)
</taxonomicName>
, which is congruent with the results obtained by
<bibRefCitation id="EFFA4B0FFF87FFB6FCC5F9B4FBA3E471" author="Agarwal, I. &amp; Ceriaco, L. M. P. &amp; Metallinou, M. &amp; Jackman, T. R. &amp; Bauer, A. M." box="[793,1046,1577,1603]" pageId="37" pageNumber="38" pagination="1 - 14" refId="ref69579" refString="Agarwal, I., Ceriaco, L. M. P., Metallinou, M., Jackman, T. R. &amp; Bauer, A. M. (2021) How the African house gecko (Hemidactylus mabouia) conquered the world. Royal Society Open Science, 8 (210749), 1 - 14. https: // doi. org / 10.1098 / rsos. 210749" type="journal article" year="2021">
Agarwal
<emphasis id="B91FEAECFF87FFB6FC5FF9B7FC0AE471" box="[899,959,1577,1603]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
(2021)
</bibRefCitation>
. Until the taxonomic identities of these populations alongside with those from mainland Africa are properly investigated, we tentatively follow the application of the most conservative designation,
<taxonomicName id="4C6B4D7DFF87FFB6FD05F9EFFCD4E4B9" box="[729,865,1649,1675]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mabouia">
<emphasis id="B91FEAECFF87FFB6FD05F9EFFCD4E4B9" box="[729,865,1649,1675]" italics="true" pageId="37" pageNumber="38">H. mabouia</emphasis>
</taxonomicName>
, for the
<collectingCountry id="F37C766EFF87FFB6FC16F9ECFB2EE4B9" box="[970,1179,1649,1675]" name="Equatorial Guinea" pageId="37" pageNumber="38">Equatorial Guinea</collectingCountry>
populations (
<bibRefCitation id="EFFA4B0FFF87FFB6FAE6F9ECFEADE49D" author="Agarwal, I. &amp; Ceriaco, L. M. P. &amp; Metallinou, M. &amp; Jackman, T. R. &amp; Bauer, A. M." pageId="37" pageNumber="38" pagination="1 - 14" refId="ref69579" refString="Agarwal, I., Ceriaco, L. M. P., Metallinou, M., Jackman, T. R. &amp; Bauer, A. M. (2021) How the African house gecko (Hemidactylus mabouia) conquered the world. Royal Society Open Science, 8 (210749), 1 - 14. https: // doi. org / 10.1098 / rsos. 210749" type="journal article" year="2021">
Agarwal
<emphasis id="B91FEAECFF87FFB6FF4BF90BFF67E49D" box="[151,210,1685,1711]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2021
</bibRefCitation>
).
<bibRefCitation id="EFFA4B0FFF87FFB6FEF2F908FDADE49D" author="Ceriaco, L. M. &amp; Marques, M. P. &amp; Bauer, A. M." box="[302,536,1685,1711]" pageId="37" pageNumber="38" pagination="91 - 108" refId="ref73685" refString="Ceriaco, L. M., Marques, M. P. &amp; Bauer, A. M. (2018) Miscellanea Herpetologica Sanctithomae, with a provisional checklist of the terrestrial herpetofauna of Sao Tome, Principe and Annobon islands. Zootaxa, 4387 (1), 91 - 108. https: // doi. org / 10.11646 / zootaxa. 4387.1.4" type="journal article" year="2018">
Ceríaco
<emphasis id="B91FEAECFF87FFB6FE53F90BFE7FE49D" box="[399,458,1685,1711]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
(2018
</bibRefCitation>
,
<bibRefCitation id="EFFA4B0FFF87FFB6FDFAF908FDD5E49D" author="Ceriaco, L. M. &amp; Marques, M. P. &amp; Bell, R. B. &amp; Bauer, A." box="[550,608,1685,1711]" pageId="37" pageNumber="38" pagination="1 - 534" refId="ref74096" refString="Ceriaco, L. M., Marques, M. P., Bell, R. B. &amp; Bauer, A. (2022) The Terrestrial Reptiles of the Gulf of Guinea Oceanic Islands. In: Ceriaco, L. M., Lima, R. F., Melo, M., Bell, R. C. (Eds.), Biodiversity of the Gulf of Guinea Oceanic Islands: science and conservation. Springer, Cham, pp. 1 - 534." type="book chapter" year="2022">2022</bibRefCitation>
) used the name
<taxonomicName id="4C6B4D7DFF87FFB6FCFDF90BFC1DE49D" box="[801,936,1685,1711]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mabouia">
<emphasis id="B91FEAECFF87FFB6FCFDF90BFC1DE49D" box="[801,936,1685,1711]" italics="true" pageId="37" pageNumber="38">H. mabouia</emphasis>
</taxonomicName>
for the populations from the islands of the Gulf of
<collectingCountry id="F37C766EFF87FFB6FF28F924FEFCE4E1" box="[244,329,1721,1747]" name="Guinea-Bissau" pageId="37" pageNumber="38">Guinea</collectingCountry>
. In
<collectingCountry id="F37C766EFF87FFB6FEA9F924FE50E4E1" box="[373,485,1721,1747]" name="Equatorial Guinea" pageId="37" pageNumber="38">Río Muni</collectingCountry>
, the species frequents human settlements (
<bibRefCitation id="EFFA4B0FFF87FFB6FC10F924FB44E4E6" author="Gonwouo, N. L. &amp; Nsang, A." box="[972,1265,1721,1748]" pageId="37" pageNumber="38" pagination="32 - 49" refId="ref78060" refString="Gonwouo, N. L. &amp; Nsang, A. (2005) An assessment of the reptiles and amphibians of the Monte Mitra forest, Monte Alen National Park. In: Sunderland, T. C. H. (Ed.), A Biodiversity Assessment of the Monte Mitra Forest, Monte Alen National Park, Equatorial Guinea. Smithsonian Institution, Washington, D. C., pp. 32 - 49." type="book chapter" year="2005">Gonwouo &amp; Nsang 2005</bibRefCitation>
). Pérez del
<bibRefCitation id="EFFA4B0FFF87FFB6FAABF924FF57E4C5" author="Val, J." pageId="37" pageNumber="38" refId="ref85942" refString="Perez del Val, J. (2001) Catalogo de las Colecciones Zoologicas de Guinea Ecuatorial del Museo Nacional de Ciencias Naturales, II. Vertebrados. Manuales Tecnicos de Museologia 11, MNCN-CSIC, Madrid, 90 pp." type="book" year="2001">Val (2001)</bibRefCitation>
listed the preserved specimens MNCN 15664 and MNCN 1566615668 as
<taxonomicName id="4C6B4D7DFF87FFB6FB95F943FB7AE4C5" box="[1097,1231,1757,1783]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="mabouia">
<emphasis id="B91FEAECFF87FFB6FB95F943FB7AE4C5" box="[1097,1231,1757,1783]" italics="true" pageId="37" pageNumber="38">H. mabouia</emphasis>
</taxonomicName>
, which are herein re-identified as
<taxonomicName id="4C6B4D7DFF87FFB6FE9BF89FFE62E529" authorityName="Hallowell" authorityYear="1854" box="[327,471,1793,1819]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="angulatus">
<emphasis id="B91FEAECFF87FFB6FE9BF89FFE62E529" box="[327,471,1793,1819]" italics="true" pageId="37" pageNumber="38">H. angulatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD436FEFF87FFB6FF1BF8B9FAC9E5C6" blockId="37.[151,1437,832,2036]" pageId="37" pageNumber="38">
<emphasis id="B91FEAECFF87FFB6FF1BF8B9FE77E50C" bold="true" box="[199,450,1828,1854]" pageId="37" pageNumber="38">Specimens examined.</emphasis>
<materialsCitation id="3B033CA3FF87FFB6FE1AF8B8FB5DE50D" ID-GBIF-Occurrence="3962324797" accessionNumber="EBD 18384" box="[454,1256,1829,1855]" collectingDate="1984-09-19" collectionCode="EBD" country="Equatorial Guinea" location="Thirty-five" municipality="Bata" pageId="37" pageNumber="38" specimenCount="1">
<location id="8EB46025FF87FFB6FE1AF8B8FDF5E50D" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FE1AF8B8FDF5E50D" box="[454,576,1829,1855]" country="Equatorial Guinea" municipality="Bata" name="Thirty-five" pageId="37" pageNumber="38">Thirty-five</location>
specimens.
<collectingCountry id="F37C766EFF87FFB6FD18F8B8FC84E50D" box="[708,817,1829,1855]" name="Equatorial Guinea" pageId="37" pageNumber="38">Río Muni</collectingCountry>
:
<collectingMunicipality id="6BB0AC84FF87FFB6FCE6F8B8FCC5E50D" box="[826,880,1829,1855]" pageId="37" pageNumber="38">Bata</collectingMunicipality>
,
<date id="FFD5103EFF87FFB6FCA4F8B8FBFEE50D" box="[888,1099,1829,1855]" pageId="37" pageNumber="38" value="1984-09-19">
<collectingDate id="EF91E9D6FF87FFB6FCA4F8B8FBFEE50D" box="[888,1099,1829,1855]" pageId="37" pageNumber="38" value="1984-09-19">19 September 1984</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFF87FFB6FB84F8B8FB24E50D" box="[1112,1169,1829,1855]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD</collectionCode>
18384)
</materialsCitation>
;
<materialsCitation id="3B033CA3FF87FFB6FB2CF8B8FDA2E551" ID-GBIF-Occurrence="3962324508" accessionNumber="EBD 18608" collectingDate="1984-07-08" collectionCode="EBD" country="Equatorial Guinea" location="Alosa" municipality="Niefang" pageId="37" pageNumber="38" specimenCode="EBD 18608-18609" specimenCount="1">
<location id="8EB46025FF87FFB6FB2CF8B8FA86E50D" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FB2CF8B8FA86E50D" box="[1264,1331,1829,1855]" country="Equatorial Guinea" municipality="Niefang" name="Alosa" pageId="37" pageNumber="38">Alosa</location>
,
<collectingMunicipality id="6BB0AC84FF87FFB6FAE0F8B8FA2DE50D" box="[1340,1432,1829,1855]" pageId="37" pageNumber="38">Niefang</collectingMunicipality>
,
<date id="FFD5103EFF87FFB6FF4BF8D4FEA8E551" box="[151,285,1865,1891]" pageId="37" pageNumber="38" value="1984-07-08">
<collectingDate id="EF91E9D6FF87FFB6FF4BF8D4FEA8E551" box="[151,285,1865,1891]" pageId="37" pageNumber="38" value="1984-07-08">8 July 1984</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FF87FFB6FEF3F8D4FDBBE551" box="[303,526,1865,1891]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD 1860818609</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CA3FF87FFB6FDF8F8D4FB1BE551" ID-GBIF-Occurrence="3962324559" accessionNumber="EBD 18664" box="[548,1198,1865,1891]" collectingDate="1984-06-13" collectionCode="EBD" country="Equatorial Guinea" county="Km" location="Ebian" municipality="Bata" pageId="37" pageNumber="38" specimenCount="1">
from
<location id="8EB46025FF87FFB6FDB8F8D4FD10E551" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FDB8F8D4FD10E551" box="[612,677,1865,1891]" country="Equatorial Guinea" county="Km" municipality="Bata" name="Ebian" pageId="37" pageNumber="38">Ebian</location>
to
<collectingMunicipality id="6BB0AC84FF87FFB6FD11F8D4FCB6E551" box="[717,771,1865,1891]" pageId="37" pageNumber="38">Bata</collectingMunicipality>
,
<collectingCounty id="62B54E72FF87FFB6FCCCF8D4FC8FE551" box="[784,826,1865,1891]" pageId="37" pageNumber="38">Km</collectingCounty>
19,
<date id="FFD5103EFF87FFB6FCB3F8D4FBBDE551" box="[879,1032,1865,1891]" pageId="37" pageNumber="38" value="1984-06-13">
<collectingDate id="EF91E9D6FF87FFB6FCB3F8D4FBBDE551" box="[879,1032,1865,1891]" pageId="37" pageNumber="38" value="1984-06-13">13 June 1984</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFF87FFB6FBC5F8D4FBE7E551" box="[1049,1106,1865,1891]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD</collectionCode>
18664)
</materialsCitation>
;
<materialsCitation id="3B033CA3FF87FFB6FB66F8D4FF5CE5B5" ID-GBIF-Occurrence="3962324474" accessionNumber="EBD 18669" collectionCode="EBD" country="Equatorial Guinea" location="Asonga" municipality="Bata" pageId="37" pageNumber="38" specimenCount="1">
<location id="8EB46025FF87FFB6FB66F8D4FAA4E551" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FB66F8D4FAA4E551" box="[1210,1297,1865,1891]" country="Equatorial Guinea" municipality="Bata" name="Asonga" pageId="37" pageNumber="38">Asonga</location>
,
<collectingMunicipality id="6BB0AC84FF87FFB6FAC3F8D4FAE7E551" box="[1311,1362,1865,1891]" pageId="37" pageNumber="38">Bata</collectingMunicipality>
(
<collectionCode id="ED7AAE3BFF87FFB6FABFF8D4FA29E551" box="[1379,1436,1865,1891]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD</collectionCode>
18669)
</materialsCitation>
;
<materialsCitation id="3B033CA3FF87FFB6FF2FF8F0FCBBE5B5" ID-GBIF-Occurrence="3962324343" accessionNumber="EBD 18696" box="[243,782,1901,1927]" collectingDate="1984-06-09" collectionCode="EBD" country="Equatorial Guinea" location="Hotel Panafrica" municipality="Bata" pageId="37" pageNumber="38" specimenCount="1">
<location id="8EB46025FF87FFB6FF2FF8F0FE16E5B5" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FF2FF8F0FE16E5B5" box="[243,419,1901,1927]" country="Equatorial Guinea" municipality="Bata" name="Hotel Panafrica" pageId="37" pageNumber="38">Hotel Panáfrica</location>
,
<collectingMunicipality id="6BB0AC84FF87FFB6FE70F8F0FE57E5B5" box="[428,482,1901,1927]" pageId="37" pageNumber="38">Bata</collectingMunicipality>
,
<date id="FFD5103EFF87FFB6FE30F8F0FDDAE5B5" box="[492,623,1901,1927]" pageId="37" pageNumber="38" value="1984-06-09">
<collectingDate id="EF91E9D6FF87FFB6FE30F8F0FDDAE5B5" box="[492,623,1901,1927]" pageId="37" pageNumber="38" value="1984-06-09">9 June 1984</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFF87FFB6FDA1F8F0FD03E5B5" box="[637,694,1901,1927]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD</collectionCode>
18696)
</materialsCitation>
;
<materialsCitation id="3B033CA3FF87FFB6FCC5F8F0FD25E599" ID-GBIF-Occurrence="3962324354" collectingDate="1963-04-15" collectingDateMax="1964-05-26" collectingDateMin="1963-04-15" collectionCode="EBD" country="Equatorial Guinea" county="Bata" location="Finca" municipality="Cocos" pageId="37" pageNumber="38" specimenCode="EBD 2735-2739, EBD 2742-2743, EBD 2747" specimenCount="1">
<location id="8EB46025FF87FFB6FCC5F8F0FCE2E5B5" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FCC5F8F0FCE2E5B5" box="[793,855,1901,1927]" country="Equatorial Guinea" county="Bata" municipality="Cocos" name="Finca" pageId="37" pageNumber="38">Finca</location>
de los
<collectingMunicipality id="6BB0AC84FF87FFB6FC7EF8F0FC5FE5B5" box="[930,1002,1901,1927]" pageId="37" pageNumber="38">Cocos</collectingMunicipality>
,
<collectingCounty id="62B54E72FF87FFB6FC28F8F0FB9FE5B5" box="[1012,1066,1901,1927]" pageId="37" pageNumber="38">Bata</collectingCounty>
,
<date id="FFD5103EFF87FFB6FBE8F8F0FB79E5B5" box="[1076,1228,1901,1928]" pageId="37" pageNumber="38" value="1963-04-15">
<collectingDate id="EF91E9D6FF87FFB6FBE8F8F0FB79E5B5" box="[1076,1228,1901,1928]" pageId="37" pageNumber="38" value="1963-04-15">15 April 1963</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FF87FFB6FB06F8F0FA2DE5BA" box="[1242,1432,1901,1928]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD 27352739</specimenCode>
,
<specimenCode id="DBCD9E85FF87FFB6FF4BF80CFEE2E599" box="[151,343,1937,1963]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD 27422743</specimenCode>
),
<date id="FFD5103EFF87FFB6FEB5F80CFE48E599" box="[361,509,1937,1963]" pageId="37" pageNumber="38" value="1964-05-26">
<collectingDate id="EF91E9D6FF87FFB6FEB5F80CFE48E599" box="[361,509,1937,1963]" pageId="37" pageNumber="38" value="1964-05-26">26 May 1964</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FF87FFB6FDD0F80CFD3DE599" box="[524,648,1937,1963]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD 2747</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CA3FF87FFB6FD47F80CFAB4E599" ID-GBIF-Occurrence="3962324583" box="[667,1281,1937,1963]" collectingDate="1964-06-09" collectionCode="EBD" country="Equatorial Guinea" location="Boca del Rio Etembue" pageId="37" pageNumber="38" specimenCode="EBD 2740-2741" specimenCount="1">
<location id="8EB46025FF87FFB6FD47F80CFC22E599" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FD47F80CFC22E599" box="[667,919,1937,1963]" country="Equatorial Guinea" name="Boca del Rio Etembue" pageId="37" pageNumber="38">Boca del Río Etembue</location>
,
<date id="FFD5103EFF87FFB6FC7FF80CFB9DE599" box="[931,1064,1937,1963]" pageId="37" pageNumber="38" value="1964-06-09">
<collectingDate id="EF91E9D6FF87FFB6FC7FF80CFB9DE599" box="[931,1064,1937,1963]" pageId="37" pageNumber="38" value="1964-06-09">9 June 1964</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FF87FFB6FBEBF80CFB4CE599" box="[1079,1273,1937,1963]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD 27402741</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CA3FF87FFB6FAD0F80CFD95E5E2" ID-GBIF-Occurrence="3962324480" collectionCode="EBD" country="Equatorial Guinea" location="Boca del Rio Aye" pageId="37" pageNumber="38" specimenCode="EBD 2745-2746" specimenCount="1">
<location id="8EB46025FF87FFB6FAD0F80CFF73E5FD" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FAD0F80CFF73E5FD" country="Equatorial Guinea" name="Boca del Rio Aye" pageId="37" pageNumber="38">Boca del Río Aye</location>
,
<collectingCountry id="F37C766EFF87FFB6FF08F828FEF4E5FD" box="[212,321,1973,1999]" name="Equatorial Guinea" pageId="37" pageNumber="38">Río Muni</collectingCountry>
(
<specimenCode id="DBCD9E85FF87FFB6FE8FF828FDADE5E2" box="[339,536,1973,2000]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD 27452746</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CA3FF87FFB6FDF1F828FB8AE5E2" ID-GBIF-Occurrence="3962324415" accessionNumber="EBD 31500" box="[557,1087,1973,2000]" collectionCode="EBD" country="Equatorial Guinea" location="San Joaquin de Ndyacom" pageId="37" pageNumber="38" specimenCount="1">
<location id="8EB46025FF87FFB6FDF1F828FCE1E5FD" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FDF1F828FCE1E5FD" box="[557,852,1973,1999]" country="Equatorial Guinea" name="San Joaquin de Ndyacom" pageId="37" pageNumber="38">San Joaquín de Ndyacom</location>
, 1990 (
<collectionCode id="ED7AAE3BFF87FFB6FC76F828FC56E5FD" box="[938,995,1973,1999]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD</collectionCode>
31500)
</materialsCitation>
.
<materialsCitation id="3B033CA3FF87FFB6FB91F828FE30E5C6" ID-GBIF-Occurrence="3962324755" accessionNumber="EBD 21395" collectingDate="1986-07-20" collectionCode="EBD" country="Equatorial Guinea" location="Bioko" municipality="Malabo" pageId="37" pageNumber="38" specimenCode="EBD 21395-21396" specimenCount="1">
<location id="8EB46025FF87FFB6FB91F828FB20E5FD" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FB91F828FB20E5FD" box="[1101,1173,1973,1999]" country="Equatorial Guinea" municipality="Malabo" name="Bioko" pageId="37" pageNumber="38">Bioko</location>
:
<collectingMunicipality id="6BB0AC84FF87FFB6FB7EF828FB4CE5FD" box="[1186,1273,1973,1999]" pageId="37" pageNumber="38">Malabo</collectingMunicipality>
,
<date id="FFD5103EFF87FFB6FADBF828FA29E5FD" box="[1287,1436,1973,1999]" pageId="37" pageNumber="38" value="1986-07-20">
<collectingDate id="EF91E9D6FF87FFB6FADBF828FA29E5FD" box="[1287,1436,1973,1999]" pageId="37" pageNumber="38" value="1986-07-20">20 July 1986</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FF87FFB6FF43F844FEC8E5C6" box="[159,381,2009,2036]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD 2139521396</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CA3FF87FFB6FE53F844FACDE5C6" ID-GBIF-Occurrence="3962324309" accessionNumber="EBD 18683, EBD 22562" box="[399,1400,2009,2036]" collectingDate="1984-06-02" collectionCode="EBD" country="Equatorial Guinea" county="Malabo" location="Las Caracolas" municipality="Finca Carboneras" pageId="37" pageNumber="38" specimenCode="EBD 18683-18695" specimenCount="1">
<location id="8EB46025FF87FFB6FE53F844FD9BE5C1" LSID="urn:lsid:plazi:treatment:03C287E8FF87FFB6FF4BFE1AFAC9E5C6:8EB46025FF87FFB6FE53F844FD9BE5C1" box="[399,558,2009,2035]" country="Equatorial Guinea" county="Malabo" municipality="Finca Carboneras" name="Las Caracolas" pageId="37" pageNumber="38">Las Caracolas</location>
,
<collectingMunicipality id="6BB0AC84FF87FFB6FDE5F844FD4AE5C1" box="[569,767,2009,2035]" pageId="37" pageNumber="38">Finca Carboneras</collectingMunicipality>
,
<collectingCounty id="62B54E72FF87FFB6FCD6F844FCD4E5C1" box="[778,865,2009,2035]" pageId="37" pageNumber="38">Malabo</collectingCounty>
,
<date id="FFD5103EFF87FFB6FCB0F844FC47E5C1" box="[876,1010,2009,2035]" pageId="37" pageNumber="38" value="1984-06-02">
<collectingDate id="EF91E9D6FF87FFB6FCB0F844FC47E5C1" box="[876,1010,2009,2035]" pageId="37" pageNumber="38" value="1984-06-02">2 June 1984</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FF87FFB6FBDDF844FB69E5C6" box="[1025,1244,2009,2036]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD 1868318695</specimenCode>
,
<collectionCode id="ED7AAE3BFF87FFB6FB3BF844FAABE5C1" box="[1255,1310,2009,2035]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="37" pageNumber="38">EBD</collectionCode>
22562)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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