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<document id="BD269D4CCDB8E5A88B2A52947C68A999" ID-DOI="10.5281/zenodo.5342544" ID-ISSN="0374-1036" ID-Zenodo-Dep="5342544" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630194263007" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2008" docId="03C287A7EF58FFA3FE06FD3FFD51B48C" docLanguage="en" docName="ActaEntMusNatPra.2008.48.1.37-50.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 48 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus tottenhami Last 1953" docType="treatment" docVersion="4" lastPageNumber="47" masterDocId="FFFBFFDFEF50FFA9FFC3FFCCFFB6B738" masterDocTitle="Revision of Afrotropical species of the Philonthus abyssinus species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="50" masterPageNumber="37" pageNumber="45" updateTime="1699182624215" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8397C0767860D62D0460B98F49015CF3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E4349FFCBA94904CFE05C354F8664AEE">
<mods:title id="A1CA1BE19EA352D470933428F5463511">Revision of Afrotropical species of the Philonthus abyssinus species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
</mods:titleInfo>
<mods:name id="C9C91BF5CE0B7168EAE676945444D05B" type="personal">
<mods:role id="6826DD458DD42A3A7412AF4A1B9E5CAE">
<mods:roleTerm id="1B550E9CACD0339DD2A1C3477E8E4D1D">Author</mods:roleTerm>
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<mods:namePart id="7356649388548C3899E1CDCDAFA9361B">Hromádka, Lubomír</mods:namePart>
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<mods:typeOfResource id="AC1829CF932FEED11711E40F6893309C">text</mods:typeOfResource>
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<mods:title id="F78E0E3EA5BF8F23B86885D46D898962">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="BDE649D39546A2B1D93BB0E7D5F52EC2">
<mods:date id="B0A8B138551DCE2E7CA4E43817E57ABE">2008</mods:date>
<mods:detail id="AD9D23B6C95095CA7E5CF66C607F5316" type="pubDate">
<mods:number id="E521C580DBCB4EFD8F363D7D3547D2BC">2008-08-15</mods:number>
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<mods:detail id="3D1B28BB33AC1393AA12DF334396937B" type="volume">
<mods:number id="358D3C45073212A669970EE4696C3A06">48</mods:number>
</mods:detail>
<mods:detail id="AB749764549037106D1ED0D17E85F217" type="issue">
<mods:number id="F3FBADD84D29C909AC53D3B497D3DA7A">1</mods:number>
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<mods:start id="9EB37ED94F7C70083B6F6F09793EC1B7">37</mods:start>
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<mods:classification id="B87BD55789216C617D9EE1E0C2173387">journal article</mods:classification>
<mods:identifier id="49A591A6BA3D87DB61B76EEF5BD41E1D" type="DOI">10.5281/zenodo.5342544</mods:identifier>
<mods:identifier id="8B7A6702FA28CD2D1B54E66CFED9AD2B" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="4717D00123B87DED37FC2E7C8CC566DA" type="Zenodo-Dep">5342544</mods:identifier>
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<treatment id="03C287A7EF58FFA3FE06FD3FFD51B48C" ID-DOI="http://doi.org/10.5281/zenodo.5448350" ID-Zenodo-Dep="5448350" LSID="urn:lsid:plazi:treatment:03C287A7EF58FFA3FE06FD3FFD51B48C" httpUri="http://treatment.plazi.org/id/03C287A7EF58FFA3FE06FD3FFD51B48C" lastPageId="10" lastPageNumber="47" pageId="8" pageNumber="45">
<subSubSection id="C371653AEF58FFA1FE06FD3FFC8EB434" box="[453,824,755,780]" pageId="8" pageNumber="45" type="nomenclature">
<paragraph id="8BD436B1EF58FFA1FE06FD3FFC8EB434" blockId="8.[453,824,755,780]" box="[453,824,755,780]" pageId="8" pageNumber="45">
<heading id="D09C81DDEF58FFA1FE06FD3FFC8EB434" bold="true" box="[453,824,755,780]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="45" reason="3">
<taxonomicName id="4C6B4D32EF58FFA1FE06FD3FFC8EB434" ID-CoL="4GBZG" authority="Last, 1953" authorityName="Last" authorityYear="1953" box="[453,824,755,780]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="tottenhami">
<emphasis id="B91FEAA3EF58FFA1FE06FD3FFC8EB434" bold="true" box="[453,824,755,780]" pageId="8" pageNumber="45">
<emphasis id="B91FEAA3EF58FFA1FE06FD3FFD0BB434" bold="true" box="[453,701,755,780]" italics="true" pageId="8" pageNumber="45">Philonthus tottenhami</emphasis>
<bibRefCitation id="EFFA4B40EF58FFA1FD00FD38FC8EB434" author="LAST H. R." box="[707,824,756,780]" pageId="8" pageNumber="45" pagination="251 - 252" refId="ref5952" refString="LAST H. R. 1953: Two new species of African Philonthus (Coleoptera, Staphylinidae). Entomologist's Monthly Magazine 91: 251 - 252." type="journal article" year="1953">Last, 1953</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C371653AEF58FFA1FD84FCDEFD00B41D" box="[583,694,786,805]" pageId="8" pageNumber="45" type="description">
<paragraph id="8BD436B1EF58FFA1FD84FCDEFD00B41D" blockId="8.[151,694,786,840]" box="[583,694,786,805]" pageId="8" pageNumber="45">
(
<figureCitation id="13502A34EF58FFA1FD8EFCDEFD07B41D" box="[589,689,786,805]" captionStart="Figs" captionStartId="9.[113,152,1547,1566]" captionTargetBox="[142,1055,217,1514]" captionTargetId="figure-11@9.[105,1097,198,1568]" captionTargetPageId="9" captionText="Figs. 17-22. Philonthus tottenhami Last, 1953. 17 – aedeagus, ventral view; 18 – aedeagus, lateral view; 19 – apical portion of sternite VIII, ventral view; 20 – sternite IX, ventral view; 21 – female tergite X, dorsal view; 22 – gonocoxites of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5342552" httpUri="https://zenodo.org/record/5342552/files/figure.png" pageId="8" pageNumber="45">Figs. 17-22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C371653AEF58FFA1FF54FCFAFE53B470" box="[151,485,820,840]" pageId="8" pageNumber="45" type="reference_group">
<paragraph id="8BD436B1EF58FFA1FF54FCFAFE53B470" blockId="8.[151,694,786,840]" box="[151,485,820,840]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7B119FEF58FFA1FF54FCFAFE53B470" box="[151,485,820,840]" pageId="8" pageNumber="45">
<treatmentCitation id="0ACA10A0EF58FFA1FF54FCFAFE57B470" author="LAST H. R." box="[151,481,820,840]" page="114" pageId="8" pageNumber="45" year="1953">
<taxonomicName id="4C6B4D32EF58FFA1FF54FCFAFE57B470" ID-CoL="4GBZG" authority="Last, 1953: 114" authorityName="Last" authorityPageNumber="114" authorityYear="1953" box="[151,481,820,840]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="tottenhami">
<emphasis id="B91FEAA3EF58FFA1FF54FCFAFEE2B470" box="[151,340,822,840]" italics="true" pageId="8" pageNumber="45">Philonthus tottenhami</emphasis>
<bibRefCitation id="EFFA4B40EF58FFA1FE9AFCF9FE57B470" author="LAST H. R." box="[345,481,820,840]" pageId="8" pageNumber="45" pagination="251 - 252" refId="ref5952" refString="LAST H. R. 1953: Two new species of African Philonthus (Coleoptera, Staphylinidae). Entomologist's Monthly Magazine 91: 251 - 252." type="journal article" year="1953">Last, 1953: 114</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C371653AEF58FFA1FF54FC92FB8FB4F9" pageId="8" pageNumber="45" type="materials_examined">
<paragraph id="8BD436B1EF58FFA1FF54FC92FDF3B44A" blockId="8.[151,1128,861,962]" box="[151,581,861,882]" pageId="8" pageNumber="45">
<emphasis id="B91FEAA3EF58FFA1FF54FC92FEA6B449" bold="true" box="[151,272,861,881]" pageId="8" pageNumber="45">
<typeStatus id="54D08813EF58FFA1FF54FC92FF72B449" box="[151,196,862,881]" pageId="8" pageNumber="45">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F37C7621EF58FFA1FED6FC92FE32B449" box="[277,388,862,881]" name="South Africa" pageId="8" pageNumber="45">South Africa</collectingCountry>
, Natal, Qudeni Forest.
</paragraph>
<paragraph id="8BD436B1EF58FFA1FF54FCB5FE16B4B4" blockId="8.[151,1128,861,962]" box="[151,416,888,908]" pageId="8" pageNumber="45">
<emphasis id="B91FEAA3EF58FFA1FF54FCB5FEAAB4B4" bold="true" box="[151,284,888,908]" pageId="8" pageNumber="45">
<typeStatus id="54D08813EF58FFA1FF54FCB5FF72B4B4" box="[151,196,889,908]" pageId="8" pageNumber="45">Type</typeStatus>
material.
</emphasis>
Not examined.
</paragraph>
<paragraph id="8BD436B1EF58FFA1FF54FC5FFB8FB4F9" blockId="8.[151,1128,861,962]" pageId="8" pageNumber="45">
<emphasis id="B91FEAA3EF58FFA1FF54FC5FFDEAB49F" bold="true" box="[151,604,915,935]" pageId="8" pageNumber="45">
Additional material examined.
<collectingCountry id="F37C7621EF58FFA1FE70FC5FFDEEB49F" box="[435,600,915,935]" name="South Africa" pageId="8" pageNumber="45">SOUTH AFRICA</collectingCountry>
:
</emphasis>
NATAL, 1 J
<specimenCount id="9D6DFD38EF58FFA1FD06FC58FD53B49F" box="[709,741,910,935]" pageId="8" pageNumber="45" type="female">1 ♀</specimenCount>
, ‘
<quantity id="4C939B54EF58FFA1FD30FC58FC91B49F" box="[755,807,916,935]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="45" unit="km" value="75.0">75km</quantity>
WSW Estcourt, Cathedral Parks For. Sta, Forestry Sta.
<date id="FFD51071EF58FFA1FEECFC62FE1CB4F9" box="[303,426,942,961]" pageId="8" pageNumber="45" value="1979-12-09" valueMax="1979-12-12" valueMin="1979-12-09">9.-12.xii.1979</date>
, veld pasture,
<quantity id="4C939B54EF58FFA1FDE6FC62FDD0B4FA" box="[549,614,942,962]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="45" unit="m" value="1400.0">1400 m</quantity>
a.s.l., S. + J. Peck, mega-dung traps’ (
<collectionCode id="ED7AAE74EF58FFA1FC6FFC62FC5BB4F9" box="[940,1005,942,961]" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="45" type="Museum">FMNH</collectionCode>
,
<collectionCode id="ED7AAE74EF58FFA1FC36FC62FB99B4F9" box="[1013,1071,942,961]" pageId="8" pageNumber="45">LHPC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C371653AEF58FFA3FF54FC16FC58B40C" lastPageId="10" lastPageNumber="47" pageId="8" pageNumber="45" type="description">
<paragraph id="8BD436B1EF58FFA1FF54FC16FBF1B4C9" blockId="8.[151,1127,986,1586]" box="[151,1095,986,1010]" pageId="8" pageNumber="45">
<emphasis id="B91FEAA3EF58FFA1FF54FC16FE80B4CA" bold="true" box="[151,310,986,1010]" pageId="8" pageNumber="45">Redescription</emphasis>
. Body length
<quantity id="4C939B54EF58FFA1FE0EFC16FD9BB4C9" box="[461,557,986,1010]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="8" pageNumber="45" unit="mm" value="11.8">11.8 mm</quantity>
, length of fore body (to the end of elytra)
<quantity id="4C939B54EF58FFA1FC2DFC16FBF5B4C9" box="[1006,1091,986,1010]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="8" pageNumber="45" unit="mm" value="5.2">5.2 mm</quantity>
.
</paragraph>
<paragraph id="8BD436B1EF58FFA1FF76FC36FE48B36A" blockId="8.[151,1127,986,1586]" pageId="8" pageNumber="45">Colouration. Black, clypeus along anterior margin and antennal sockets narrowly brown; mandibles, maxillary and labial palpi black-brown; antennae and legs black; terminal tarsomeres of all tarsi somewhat paler.</paragraph>
<paragraph id="8BD436B1EF58FFA1FF76FB96FDC0B20A" blockId="8.[151,1127,986,1586]" pageId="8" pageNumber="45">Head transverse (ratio 44: 30), parallel-sided. Eyes small, shorter than temples (ratio 12: 17). Clypeus with wide and deep depression medially. Posterior angles of head capsule obtusely rounded, with one long black bristle. Medial interocular punctures widely separated, distance between medial interocular punctures almost three times as long as distance between medial and lateral interocular punctures, two coarse punctures at posterior margin of eyes. Temporal area impunctate. Surface with very fine microsculpture of transverse waves; sides with several black bristles of unequal length.</paragraph>
<paragraph id="8BD436B1EF58FFA1FF76FAF6FC56B28A" blockId="8.[151,1127,986,1586]" pageId="8" pageNumber="45">Antennae reaching posterior half of pronotum when reclined, antennomeres 1-3 and 11 longer than wide, antennomeres 4-6 as long as wide, antennomeres 7-10 a little wider than long. Antennomere 1 distinctly shorter than antennomeres 2-3 combined, antennomere 2 shorter than antennomere 3, antennomere 5 a little shorter than antennomere 2.</paragraph>
<paragraph id="8BD436B1EF58FFA1FF76FA76FC68B10A" blockId="8.[151,1127,986,1586]" pageId="8" pageNumber="45">Pronotum parallel-sided, longer than wide (ratio 47: 43); anterior angles rectangular, with many dark bristles of unequal length; posterior angles strongly rounded. Each dorsal row with 4 coarse punctures; each sublateral row with 2 punctures. Pronotal margins with several black bristles of unequal length. Surface with microsculpture similar to that on head.</paragraph>
<caption id="DF146639EF59FFA0FFB2F9C7FE21B16C" ID-DOI="http://doi.org/10.5281/zenodo.5342552" ID-Zenodo-Dep="5342552" httpUri="https://zenodo.org/record/5342552/files/figure.png" pageId="9" pageNumber="46" startId="9.[113,152,1547,1566]" targetBox="[142,1055,217,1514]" targetPageId="9">
<paragraph id="8BD436B1EF59FFA0FFB2F9C7FE21B16C" blockId="9.[113,1089,1547,1620]" pageId="9" pageNumber="46">
Figs. 17-22.
<taxonomicName id="4C6B4D32EF59FFA0FF1FF9C0FE4EB126" authority="Last, 1953" authorityName="Last" authorityYear="1953" box="[220,504,1547,1567]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="tottenhami">
<emphasis id="B91FEAA3EF59FFA0FF1FF9C0FE2EB126" box="[220,408,1548,1566]" italics="true" pageId="9" pageNumber="46">Philonthus tottenhami</emphasis>
<bibRefCitation id="EFFA4B40EF59FFA0FE5EF9C0FE4EB126" author="LAST H. R." box="[413,504,1547,1567]" pageId="9" pageNumber="46" pagination="251 - 252" refId="ref5952" refString="LAST H. R. 1953: Two new species of African Philonthus (Coleoptera, Staphylinidae). Entomologist's Monthly Magazine 91: 251 - 252." type="journal article" year="1953">Last, 1953</bibRefCitation>
</taxonomicName>
. 17 – aedeagus, ventral view; 18 – aedeagus, lateral view; 19 – apical portion of sternite VIII, ventral view; 20 – sternite IX, ventral view; 21 – female tergite X, dorsal view; 22 – gonocoxites of female genital segment.
</paragraph>
</caption>
<paragraph id="8BD436B1EF5AFFA3FF76FF17FDEBB60B" blockId="10.[151,1127,219,948]" pageId="10" pageNumber="47">Scutellum very coarsely and densely punctate, diameters of punctures almost twice as large as eye-facets. Distance between punctures much smaller than diameters of punctures. Surface with traces of fine microsculpture.</paragraph>
<paragraph id="8BD436B1EF5AFFA3FF76FEF7FE85B6EB" blockId="10.[151,1127,219,948]" pageId="10" pageNumber="47">Elytra wider than long (ratio 57: 53), slightly widened posteriad. Punctation dense and coarse, punctures a little smaller than those on scutellum, separated in transverse direction mostly by one puncture diameter. Anterior pronotal angles with one long bristle; pronotal margins with many long bristles of unequal length. Surface without microsculpture; setation brown-yellow.</paragraph>
<paragraph id="8BD436B1EF5AFFA3FF76FE10FCA1B50C" blockId="10.[151,1127,219,948]" pageId="10" pageNumber="47">Legs. Metatarsus longer than metatibia (ratio 31: 27), all tibiae with many bristles of different length. Metatarsomere 1 distinctly longer than metatarsomeres 2-3 combined, metatarsomere 5 approximately as long as metatarsomere 1.</paragraph>
<paragraph id="8BD436B1EF5AFFA3FF76FDF7FC19B58B" blockId="10.[151,1127,219,948]" pageId="10" pageNumber="47">Abdomen from fifth visible tergite slightly narrowed towards both base and apex. First four visible tergites with two basal lines, elevated area between basal lines on first three tergites almost impunctate, that on fourth tergite sparsely punctate. Punctation of visible tergites very sparse and fine. Surface without microsculpture, very shiny; setation dark.</paragraph>
<paragraph id="8BD436B1EF5AFFA3FF76FD77FE24B42C" blockId="10.[151,1127,219,948]" pageId="10" pageNumber="47">
Male. Protarsomeres 1-3 moderately dilated, each densely covered with modified pale setae ventrally, protarsomere 4 narrow and small; sternite VIII (
<figureCitation id="13502A34EF5AFFA3FCFBFD17FCC9B5CB" box="[824,895,731,755]" captionStart="Figs" captionStartId="9.[113,152,1547,1566]" captionTargetBox="[142,1055,217,1514]" captionTargetId="figure-11@9.[105,1097,198,1568]" captionTargetPageId="9" captionText="Figs. 17-22. Philonthus tottenhami Last, 1953. 17 – aedeagus, ventral view; 18 – aedeagus, lateral view; 19 – apical portion of sternite VIII, ventral view; 20 – sternite IX, ventral view; 21 – female tergite X, dorsal view; 22 – gonocoxites of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5342552" httpUri="https://zenodo.org/record/5342552/files/figure.png" pageId="10" pageNumber="47">Fig.19</figureCitation>
), sternite IX (
<figureCitation id="13502A34EF5AFFA3FBD2FD17FBEDB5CB" box="[1041,1115,731,755]" captionStart="Figs" captionStartId="9.[113,152,1547,1566]" captionTargetBox="[142,1055,217,1514]" captionTargetId="figure-11@9.[105,1097,198,1568]" captionTargetPageId="9" captionText="Figs. 17-22. Philonthus tottenhami Last, 1953. 17 – aedeagus, ventral view; 18 – aedeagus, lateral view; 19 – apical portion of sternite VIII, ventral view; 20 – sternite IX, ventral view; 21 – female tergite X, dorsal view; 22 – gonocoxites of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5342552" httpUri="https://zenodo.org/record/5342552/files/figure.png" pageId="10" pageNumber="47">Fig.20</figureCitation>
), aedeagus (
<figureCitation id="13502A34EF5AFFA3FECBFD37FE33B42C" box="[264,389,763,788]" captionStart="Figs" captionStartId="9.[113,152,1547,1566]" captionTargetBox="[142,1055,217,1514]" captionTargetId="figure-11@9.[105,1097,198,1568]" captionTargetPageId="9" captionText="Figs. 17-22. Philonthus tottenhami Last, 1953. 17 – aedeagus, ventral view; 18 – aedeagus, lateral view; 19 – apical portion of sternite VIII, ventral view; 20 – sternite IX, ventral view; 21 – female tergite X, dorsal view; 22 – gonocoxites of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5342552" httpUri="https://zenodo.org/record/5342552/files/figure.png" pageId="10" pageNumber="47">Figs. 17-18</figureCitation>
).
</paragraph>
<paragraph id="8BD436B1EF5AFFA3FF76FCD7FC58B40C" blockId="10.[151,1127,219,948]" box="[181,1006,795,820]" pageId="10" pageNumber="47">
Female. Tergite X (
<figureCitation id="13502A34EF5AFFA3FE47FCD7FE65B40C" box="[388,467,795,820]" captionStart="Figs" captionStartId="9.[113,152,1547,1566]" captionTargetBox="[142,1055,217,1514]" captionTargetId="figure-11@9.[105,1097,198,1568]" captionTargetPageId="9" captionText="Figs. 17-22. Philonthus tottenhami Last, 1953. 17 – aedeagus, ventral view; 18 – aedeagus, lateral view; 19 – apical portion of sternite VIII, ventral view; 20 – sternite IX, ventral view; 21 – female tergite X, dorsal view; 22 – gonocoxites of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5342552" httpUri="https://zenodo.org/record/5342552/files/figure.png" pageId="10" pageNumber="47">Fig. 21</figureCitation>
), gonocoxites of female genital segment (
<figureCitation id="13502A34EF5AFFA3FC52FCD7FC57B40C" box="[913,993,795,820]" captionStart="Figs" captionStartId="9.[113,152,1547,1566]" captionTargetBox="[142,1055,217,1514]" captionTargetId="figure-11@9.[105,1097,198,1568]" captionTargetPageId="9" captionText="Figs. 17-22. Philonthus tottenhami Last, 1953. 17 – aedeagus, ventral view; 18 – aedeagus, lateral view; 19 – apical portion of sternite VIII, ventral view; 20 – sternite IX, ventral view; 21 – female tergite X, dorsal view; 22 – gonocoxites of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5342552" httpUri="https://zenodo.org/record/5342552/files/figure.png" pageId="10" pageNumber="47">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C371653AEF5AFFA3FF54FCF7FD4CB4AB" pageId="10" pageNumber="47" type="diagnosis">
<paragraph id="8BD436B1EF5AFFA3FF54FCF7FD94B44B" blockId="10.[151,1127,219,948]" pageId="10" pageNumber="47">
<emphasis id="B91FEAA3EF5AFFA3FF54FCF7FE3CB46B" bold="true" box="[151,394,827,851]" pageId="10" pageNumber="47">Differential diagnosis</emphasis>
.
<taxonomicName id="4C6B4D32EF5AFFA3FE5BFCF1FD32B46C" authorityName="Last" authorityYear="1953" box="[408,644,829,852]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="tottenhami">
<emphasis id="B91FEAA3EF5AFFA3FE5BFCF1FD32B46C" box="[408,644,829,852]" italics="true" pageId="10" pageNumber="47">Philonthus tottenhami</emphasis>
</taxonomicName>
may be distinguished from all species of this group by denser punctation of elytra.
</paragraph>
<paragraph id="8BD436B1EF5AFFA3FF54FCB7FD4CB4AB" blockId="10.[151,1127,219,948]" box="[151,762,891,916]" pageId="10" pageNumber="47">
<emphasis id="B91FEAA3EF5AFFA3FF54FCB7FEA3B4AB" bold="true" box="[151,277,891,915]" pageId="10" pageNumber="47">Bionomics.</emphasis>
Collected in mega dung trap on veld pasture.
</paragraph>
</subSubSection>
<subSubSection id="C371653AEF5AFFA3FF54FC57FD51B48C" box="[151,743,923,948]" pageId="10" pageNumber="47" type="distribution">
<paragraph id="8BD436B1EF5AFFA3FF54FC57FD51B48C" blockId="10.[151,1127,219,948]" box="[151,743,923,948]" pageId="10" pageNumber="47">
<emphasis id="B91FEAA3EF5AFFA3FF54FC57FE94B48B" bold="true" box="[151,290,923,947]" pageId="10" pageNumber="47">Distribution</emphasis>
.
<collectingCountry id="F37C7621EF5AFFA3FEF3FC57FE01B48B" box="[304,439,923,947]" name="South Africa" pageId="10" pageNumber="47">South Africa</collectingCountry>
(
<bibRefCitation id="EFFA4B40EF5AFFA3FE05FC50FD8FB48B" author="LAST H. R." box="[454,569,923,948]" pageId="10" pageNumber="47" pagination="251 - 252" refId="ref5952" refString="LAST H. R. 1953: Two new species of African Philonthus (Coleoptera, Staphylinidae). Entomologist's Monthly Magazine 91: 251 - 252." type="journal article" year="1953">LAST 1953</bibRefCitation>
,
<bibRefCitation id="EFFA4B40EF5AFFA3FD87FC50FD6DB48C" author="HERMAN L. H." box="[580,731,924,948]" pageId="10" pageNumber="47" pagination="2441 - 3020" refId="ref5882" refString="HERMAN L. H. 2001: Catalog of the Staphylinidae (Insecta, Coleoptera). 1758 to the end of the second millenium. V. Staphylinine group (Part 2). Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History 265: 2441 - 3020." type="journal article" year="2001">HERMAN 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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