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<document id="7C567D22259BCB96B7E52261AC8CFBED" ID-CLB-Dataset="4098" ID-DOI="10.11646/zootaxa.3861.6.3" ID-GBIF-Dataset="38808f1d-0656-4659-bc31-0e5eee066ed6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5247997" ID-ZooBank="8169FF7C-74C0-4385-8B67-09306D815CD2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623443819181" checkinUser="felipe" docAuthor="Angulo, Arturo, López, Myrna I., Bussing, William A. &amp; Murase, Atsunobu" docDate="2014" docId="03C2878AFFF7FF9D44D6FF1B848FDED2" docLanguage="en" docName="zt03861p574.pdf" docOrigin="Zootaxa 3861 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chimaeriformes" docType="key" docVersion="7" lastPageNumber="571" masterDocId="FFFBFFF2FFE6FF8C4441FF8C8113DB25" masterDocTitle="Records of chimaeroid fishes (Holocephali: Chimaeriformes) from the Pacific coast of Costa Rica, with the description of a new species of Chimera (Chimaeridae) from the eastern Pacific Ocean" masterLastPageNumber="574" masterPageNumber="554" pageNumber="571" updateTime="1699013252087" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C9DC10153BE178D420EFF31B97172E91">Records of chimaeroid fishes (Holocephali: Chimaeriformes) from the Pacific coast of Costa Rica, with the description of a new species of Chimera (Chimaeridae) from the eastern Pacific Ocean</mods:title>
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<mods:namePart id="6B64A51567774E4B8168042447917A0D">Angulo, Arturo</mods:namePart>
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<mods:namePart id="09632D18C95F69ACD4862C65A341A01F">López, Myrna I.</mods:namePart>
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<mods:name id="CB60A06210A858D9B0F5805174243D85" type="personal">
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<mods:namePart id="8C52ABF4A2FAFE3E6F41D87F45D80B0A">Bussing, William A.</mods:namePart>
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<mods:namePart id="846176AEE1806024CAEAA022EDCD5607">Murase, Atsunobu</mods:namePart>
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<mods:title id="F9951972279FE6566BB8FCA93894B3D5">Zootaxa</mods:title>
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<mods:date id="F43CC187FDEFB1FFAA5A413184A95271">2014</mods:date>
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<mods:number id="2CF5C6E54EB50C68C9A6BA97E791867C">2014-09-18</mods:number>
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<mods:number id="DA49EC2688BC4EDFFA65B5282E18EC15">3861</mods:number>
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<mods:identifier id="49BFD07396969D90F2D3B5090453E210" type="DOI">10.11646/zootaxa.3861.6.3</mods:identifier>
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<treatment id="03C2878AFFF7FF9D44D6FF1B848FDED2" ID-DOI="http://doi.org/10.5281/zenodo.5559872" ID-GBIF-Taxon="188606357" ID-Zenodo-Dep="5559872" LSID="urn:lsid:plazi:treatment:03C2878AFFF7FF9D44D6FF1B848FDED2" httpUri="http://treatment.plazi.org/id/03C2878AFFF7FF9D44D6FF1B848FDED2" lastPageNumber="571" pageId="17" pageNumber="571">
<subSubSection id="C3716517FFF7FF9D44D6FF1B8503DA65" pageId="17" pageNumber="571" type="nomenclature">
<paragraph id="8BD4369CFFF7FF9D44D6FF1B8240DB95" blockId="17.[151,851,151,176]" box="[151,851,151,176]" pageId="17" pageNumber="571">
<heading id="D09C81F0FFF7FF9D44D6FF1B8240DB95" bold="true" box="[151,851,151,176]" fontSize="10" level="2" pageId="17" pageNumber="571" reason="2">
<emphasis id="B91FEA8EFFF7FF9D44D6FF1B8240DB95" bold="true" box="[151,851,151,176]" pageId="17" pageNumber="571">
Key to eastern Pacific species of the order
<taxonomicName id="4C6B4D1FFFF7FF9D46CEFF1B8240DB95" ID-CoL="X9" ID-ENA="7864" box="[655,851,151,176]" class="Holocephali" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="order">Chimaeriformes</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FF538503DA65" blockId="17.[151,1436,223,320]" pageId="17" pageNumber="571">
The following key is based on our research and data available in the literature (Didier &amp; Nakaya 1999, Didier &amp; Rosenberger 2002,
<bibRefCitation id="EFFA4B6DFFF7FF9D4531FE888351DA38" author="Barnett, L. K. &amp; Didier, D. A. &amp; Long, D. J. &amp; Ebert, D. A." box="[368,578,260,285]" pageId="17" pageNumber="571" pagination="27 - 38" refId="ref12350" refString="Barnett, L. K., Didier, D. A., Long, D. J. &amp; Ebert, D. A. (2006) Hydrolagus mccoskeri sp. nov., a new species of chimaeroid fish from the Galapagos Islands (Holocephali: Chimaeriformes: Chimaeridae). Zootaxa, 1328, 27 - 38." type="journal article" year="2006">
Barnett
<emphasis id="B91FEA8EFFF7FF9D4588FE898312DA39" box="[457,513,260,284]" italics="true" pageId="17" pageNumber="571">et al.</emphasis>
2006
</bibRefCitation>
,
<bibRefCitation id="EFFA4B6DFFF7FF9D460DFE8883CDDA38" author="Nelson, J. S." box="[588,734,260,285]" pageId="17" pageNumber="571" refId="ref14557" refString="Nelson, J. S. (2006) Fishes of the world. John Wiley &amp; Sons, New Jersey, 601 pp." type="book" year="2006">
<collectingRegion id="49AFF87EFFF7FF9D460DFE888388DA38" box="[588,667,260,285]" country="New Zealand" name="Nelson" pageId="17" pageNumber="571">Nelson</collectingRegion>
2006
</bibRefCitation>
,
<bibRefCitation id="EFFA4B6DFFF7FF9D46A9FE8882DDDA38" author="Quaranta, K. L. &amp; Didier, D. A. &amp; Long, D. J. &amp; Ebert, D. A." box="[744,974,260,285]" pageId="17" pageNumber="571" pagination="33 - 45" refId="ref14582" refString="Quaranta, K. L., Didier, D. A., Long, D. J. &amp; Ebert, D. A. (2006) A new species of chimaeroid, Hydrolagus alphus sp. nov. (Chimaeriformes: Chimaeridae) from the Galapagos Islands. Zootaxa, 1377, 33 - 45." type="journal article" year="2006">
Quaranta
<emphasis id="B91FEA8EFFF7FF9D4714FE89829EDA39" box="[853,909,260,284]" italics="true" pageId="17" pageNumber="571">et al.</emphasis>
2006
</bibRefCitation>
,
<bibRefCitation id="EFFA4B6DFFF7FF9D4798FE8885E0DA38" author="Didier, D. A. &amp; Meckley, T." box="[985,1267,260,285]" pageId="17" pageNumber="571" pagination="49 - 50" refId="ref13035" refString="Didier, D. A. &amp; Meckley, T. (2009 a) Rhinochimaeridae. In: Nakaya, K., Yabe, M., Imamura, H., Romero, M. C. &amp; Yoshida, M. (Eds.), Deep-sea fishes of Peru. Japan Deep Sea Trawlers Association, Tokyo, pp. 49 - 50." type="book chapter" year="2009">Didier &amp; Meckley 2009a</bibRefCitation>
, b,
<bibRefCitation id="EFFA4B6DFFF7FF9D4158FE8881C1DA65" author="James, K. &amp; Ebert, D. A. &amp; Long, D. J. &amp; Didier, D. A." pageId="17" pageNumber="571" pagination="59 - 68" refId="ref13776" refString="James, K. C, Ebert, D. A., Long, D. J. &amp; Didier, D. A. (2009) A new species of chimaera, Hydrolagus melanophasma sp. nov. (Chondrichthyes: Chimaeriformes: Chimaeridae), from the eastern North Pacific. Zootaxa, 2218, 59 - 68." type="journal article" year="2009">
James
<emphasis id="B91FEA8EFFF7FF9D4125FE89848FDA39" box="[1380,1436,260,284]" italics="true" pageId="17" pageNumber="571">et al.</emphasis>
2009
</bibRefCitation>
,
<bibRefCitation id="EFFA4B6DFFF7FF9D449CFEAB830CDA65" author="Gonzalez-Acosta, A. F. &amp; Castro-Aguirre, J. &amp; Didier, D. A. &amp; Velez-Marin, R. &amp; Burnes-Romo, L. A." box="[221,543,295,320]" pageId="17" pageNumber="571" pagination="197 - 201" refId="ref13626" refString="Gonzalez-Acosta, A. F., Castro-Aguirre, J. L, Didier, D. A., Velez-Marin, R. &amp; Burnes-Romo, L. A. (2010) Occurrence of Hydrolagus macrophthalmus (Chondrichthyes: Holocephali: Chimaeridae) in the northeastern Pacific. Revista Mexicana de Biodiversidad, 81 (1), 197 - 201. http: // dx. doi. org / 10.7550 / rmb. 17717" type="journal article" year="2010">
González-Acosta
<emphasis id="B91FEA8EFFF7FF9D45E5FEA580CEDA65" box="[420,477,296,320]" italics="true" pageId="17" pageNumber="571">et al.</emphasis>
2010
</bibRefCitation>
,
<bibRefCitation id="EFFA4B6DFFF7FF9D466BFEA4823CDA65" author="Bustamante, C. &amp; Flores, H. &amp; Concha-Perez, Y. &amp; Vargas-Caro, C. &amp; Lamilla, J. &amp; Bennett, M." box="[554,815,295,320]" pageId="17" pageNumber="571" pagination="236 - 242" refId="ref12453" refString="Bustamante, C., Flores, H., Concha-Perez, Y., Vargas-Caro, C., Lamilla, J. &amp; Bennett, M. (2012) First record of Hydrolagus melanophasma James, Ebert, Long &amp; Didier, 2009 (Chondrichthyes, Chimaeriformes, Holocephali) from the southeastern Pacific Ocean. Latin American Journal of Aquatic Research, 40 (1), 236 - 242. http: // dx. doi. org / 10.3856 / vol 40 - issue 1 - fulltext- 23" type="journal article" year="2012">
Bustamante
<emphasis id="B91FEA8EFFF7FF9D46F4FEA583FEDA65" box="[693,749,296,320]" italics="true" pageId="17" pageNumber="571">et al.</emphasis>
2012
</bibRefCitation>
,
<bibRefCitation id="EFFA4B6DFFF7FF9D477BFEAB8517DA65" author="Didier, D. A. &amp; Kemper, J. M. &amp; Ebert, D. A." box="[826,1028,295,320]" pageId="17" pageNumber="571" pagination="97 - 124" refId="ref13283" refString="Didier, D. A., Kemper, J. M. &amp; Ebert, D. A. (2012) Phylogeny and classification of extant Holocephalans. In: Carrier, J. C., Musick, J. A. &amp; Heithaus, M. R. (Eds.), Biology of sharks and their relatives. CRC Press, Boca Raton, pp. 97 - 124." type="book chapter" year="2012">
Didier
<emphasis id="B91FEA8EFFF7FF9D47C9FEA582D2DA65" box="[904,961,296,320]" italics="true" pageId="17" pageNumber="571">et al.</emphasis>
2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716517FFF7FF9D44D6FEE6848FDED2" pageId="17" pageNumber="571" type="key">
<keyStep id="309F2E39FFF7FF9D44D6FEE68488DAF2" pageId="17" pageNumber="571">
<paragraph id="8BD4369CFFF7FF9D44D6FEE68485DAB9" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FEE68485DAB9" pageId="17" pageNumber="571">
1 Snout with elongate, flexible, hooklike process; lateral line canals closed; second dorsal fin not elongate; tail heterocercal (
<collectingCountry id="F37C760CFFF7FF9D44A6FE0B802ADAB9" box="[231,313,391,412]" name="Ecuador" pageId="17" pageNumber="571">Ecuador</collectingCountry>
to
<collectingCountry id="F37C760CFFF7FF9D4516FE0B80ABDAB9" box="[343,440,391,412]" name="Argentina" pageId="17" pageNumber="571">Argentina</collectingCountry>
).............................................
<taxonomicName id="4C6B4D1FFFF7FF9D47A6FE0B8485DAB9" ID-CoL="PVZB" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[999,1430,391,412]" class="Holocephali" family="Callorhinchidae" genus="Callorhinchus" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="callorynchus">
<emphasis id="B91FEA8EFFF7FF9D47A6FE0B85E1DAB9" box="[999,1266,391,412]" italics="true" pageId="17" pageNumber="571">Callorhinchus callorynchus</emphasis>
(Linnaeus 1758)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FE298488DAF2" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FE298488DAF2" pageId="17" pageNumber="571">- Snout short and rounded or long and pointed, not hooklike; lateral line canals are open grooves; second dorsal fin elongate; tail diphycercal.......................................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E39FFF7FF9D44D6FE538488D969" pageId="17" pageNumber="571">
<paragraph id="8BD4369CFFF7FF9D44D6FE538488D937" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FE538488D937" pageId="17" pageNumber="571">2 Snout short and bluntly rounded; caudal-fin axis horizontal with the fin nearly symmetrical, epaxial and hypaxial lobes equal sized............................................................................................... 3</keyLead>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FD968488D969" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FD968488D969" pageId="17" pageNumber="571">- Snout elongated and pointed; caudal-fin axis weakly raised with the fin asymmetrical, epaxial caudal-fin lobe narrower than hypaxial lobe........................................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E39FFF7FF9D44D6FDD98488D9A2" pageId="17" pageNumber="571">
<paragraph id="8BD4369CFFF7FF9D44D6FDD9848FD94F" blockId="17.[151,1436,362,1527]" box="[151,1436,596,618]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FDD9848FD94F" box="[151,1436,596,618]" pageId="17" pageNumber="571">
3 Anal fin present (
<collectingCountry id="F37C760CFFF7FF9D45C9FDD980E1D94F" box="[392,498,597,618]" name="Costa Rica" pageId="17" pageNumber="571">Costa Rica</collectingCountry>
to
<collectingCountry id="F37C760CFFF7FF9D4651FDD9832DD94F" box="[528,574,597,618]" name="Peru" pageId="17" pageNumber="571">Peru</collectingCountry>
, probably more widespread in the southeastern Pacific)....
<taxonomicName id="4C6B4D1FFFF7FF9D40C6FDD9845AD94F" ID-CoL="5Y3XB" authorityName="Angulo, Lopez, Bussing &amp; Murase" authorityYear="2014" box="[1159,1353,597,618]" class="Holocephali" family="Chimaeridae" genus="Chimaera" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="B91FEA8EFFF7FF9D40C6FDD9845AD94F" box="[1159,1353,597,618]" italics="true" pageId="17" pageNumber="571">Chimaera orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57F5FFF7FF9D410EFDD8848FD94F" box="[1359,1436,596,618]" pageId="17" pageNumber="571" rank="species">
<emphasis id="B91FEA8EFFF7FF9D410EFDD88485D94F" bold="true" box="[1359,1430,596,618]" pageId="17" pageNumber="571">sp. nov</emphasis>
.
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FDFE8488D9A2" blockId="17.[151,1436,362,1527]" box="[151,1435,626,647]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FDFE8488D9A2" box="[151,1435,626,647]" pageId="17" pageNumber="571">- Anal fin absent...................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E39FFF7FF9D44D6FD038488D9E7" pageId="17" pageNumber="571">
<paragraph id="8BD4369CFFF7FF9D44D6FD038488D981" blockId="17.[151,1436,362,1527]" box="[151,1435,655,676]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FD038488D981" box="[151,1435,655,676]" pageId="17" pageNumber="571">4 Anterior and posterior regions of second dorsal-fin considerably taller than middle region............................ 5</keyLead>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FD218488D9E7" blockId="17.[151,1436,362,1527]" box="[151,1435,685,706]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FD218488D9E7" box="[151,1435,685,706]" pageId="17" pageNumber="571">- Second dorsal-fin uniform in height throughout............................................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E39FFF7FF9D44D6FD468488D812" pageId="17" pageNumber="571">
<paragraph id="8BD4369CFFF7FF9D44D6FD46848FD9D9" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FD46848FD9D9" pageId="17" pageNumber="571">
5 Anterior edge of dorsal-fin spine not serrated; uniform dark brown to reddish-brown across entire body with numerous white spots on the head and trunk (Gulf of Alaska to
<collectingCountry id="F37C760CFFF7FF9D46E2FD6B821FD9D9" box="[675,780,743,764]" name="Costa Rica" pageId="17" pageNumber="571">Costa Rica</collectingCountry>
)...................
<taxonomicName id="4C6B4D1FFFF7FF9D404EFD6B848FD9D9" ID-CoL="6MK28" authority="(Lay &amp; Bennett 1839)" baseAuthorityName="Lay &amp; Bennett" baseAuthorityYear="1839" box="[1039,1436,743,764]" class="Holocephali" family="Chimaeridae" genus="Hydrolagus" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="colliei">
<emphasis id="B91FEA8EFFF7FF9D404EFD6B85D2D9D9" box="[1039,1217,743,764]" italics="true" pageId="17" pageNumber="571">Hydrolagus colliei</emphasis>
(Lay &amp; Bennett 1839)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FC898488D812" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FC898488D812" pageId="17" pageNumber="571">- Anterior edge of dorsal-fin spine serrated; uniform brown across entire body with or without a distinct white spot on the lateral side above the pectoral fins............................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E39FFF7FF9D44D6FCB3848FD8EF" pageId="17" pageNumber="571">
<paragraph id="8BD4369CFFF7FF9D44D6FCB3848FD8AA" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FCB3848FD8AA" pageId="17" pageNumber="571">
6 Snout blunt; EYL more than 40% HDL; tail region short and stout, PCA less than 57% BDL; uniform brown across entire body with a distinct white spot on the lateral side above the pectoral fins (
<collectingRegion id="49AFF87EFFF7FF9D47E2FCD18546D857" box="[931,1109,861,882]" country="Ecuador" name="Galapagos" pageId="17" pageNumber="571">Galapagos Islands</collectingRegion>
).................................................................................
<taxonomicName id="4C6B4D1FFFF7FF9D4739FCF6848FD8AA" ID-CoL="6MK29" authority="Quaranta, Didier, Long &amp; Ebert 2006" authorityName="Quaranta, Didier, Long &amp; Ebert" authorityYear="2006" box="[888,1436,890,911]" class="Holocephali" family="Chimaeridae" genus="Hydrolagus" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="alphus">
<emphasis id="B91FEA8EFFF7FF9D4739FCF6853CD8AA" box="[888,1071,890,911]" italics="true" pageId="17" pageNumber="571">Hydrolagus alphus</emphasis>
<bibRefCitation id="EFFA4B6DFFF7FF9D4075FCF6848FD8AA" author="Quaranta, K. L. &amp; Didier, D. A. &amp; Long, D. J. &amp; Ebert, D. A." box="[1076,1436,890,911]" pageId="17" pageNumber="571" pagination="33 - 45" refId="ref14582" refString="Quaranta, K. L., Didier, D. A., Long, D. J. &amp; Ebert, D. A. (2006) A new species of chimaeroid, Hydrolagus alphus sp. nov. (Chimaeriformes: Chimaeridae) from the Galapagos Islands. Zootaxa, 1377, 33 - 45." type="journal article" year="2006">Quaranta, Didier, Long &amp; Ebert 2006</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FC1B848FD8EF" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FC1B848FD8EF" pageId="17" pageNumber="571">
- Snout pointed at tip; EYL less than 40% HDL; tail region elongate and slender, PCA more than 58% BDL; uniform brown across entire body with no white markings (
<collectingCountry id="F37C760CFFF7FF9D46C5FC3983DCD8EF" box="[644,719,949,970]" name="Mexico" pageId="17" pageNumber="571">Mexico</collectingCountry>
to
<collectingCountry id="F37C760CFFF7FF9D46AFFC398230D8EF" box="[750,803,949,970]" name="Chile" pageId="17" pageNumber="571">Chile</collectingCountry>
)................
<taxonomicName id="4C6B4D1FFFF7FF9D47BBFC39848FD8EF" ID-CoL="6MJQ5" authority="de Buen 1959" authorityName="de Buen" authorityYear="1959" box="[1018,1436,949,970]" class="Holocephali" family="Chimaeridae" genus="Hydrolagus" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="macrophthalmus">
<emphasis id="B91FEA8EFFF7FF9D47BBFC398403D8EF" box="[1018,1296,949,970]" italics="true" pageId="17" pageNumber="571">Hydrolagus macrophthalmus</emphasis>
de Buen 1959
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E39FFF7FF9D44D6FC5E8488DFB2" pageId="17" pageNumber="571">
<paragraph id="8BD4369CFFF7FF9D44D6FC5E848FDF1A" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FC5E848FDF1A" pageId="17" pageNumber="571">
7 EYL more than 10% BDL; two narrowly spaced columns of serrations on the posterolateral edges of the distal 6675% of spine (in adults); uniform overall medium grey on dorsal and lateral parts extending to near the ventrum, and dorsum with many irregular circular and elongate white blotches (
<collectingRegion id="49AFF87EFFF7FF9D469AFB81829EDF07" box="[731,909,1037,1058]" country="Ecuador" name="Galapagos" pageId="17" pageNumber="571">Galapagos Islands</collectingRegion>
).................................................................................................
<taxonomicName id="4C6B4D1FFFF7FF9D4726FBA6848FDF1A" ID-CoL="3N7F4" authority="Barnett, Didier, Long &amp; Ebert 2006" authorityName="Barnett, Didier, Long &amp; Ebert" authorityYear="2006" box="[871,1436,1066,1087]" class="Holocephali" family="Chimaeridae" genus="Hydrolagus" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="mccoskeri">
<emphasis id="B91FEA8EFFF7FF9D4726FBA6852CDF1A" box="[871,1087,1066,1087]" italics="true" pageId="17" pageNumber="571">Hydrolagus mccoskeri</emphasis>
<bibRefCitation id="EFFA4B6DFFF7FF9D4007FBA6848FDF1A" author="Barnett, L. K. &amp; Didier, D. A. &amp; Long, D. J. &amp; Ebert, D. A." box="[1094,1436,1066,1087]" pageId="17" pageNumber="571" pagination="27 - 38" refId="ref12350" refString="Barnett, L. K., Didier, D. A., Long, D. J. &amp; Ebert, D. A. (2006) Hydrolagus mccoskeri sp. nov., a new species of chimaeroid fish from the Galapagos Islands (Holocephali: Chimaeriformes: Chimaeridae). Zootaxa, 1328, 27 - 38." type="journal article" year="2006">Barnett, Didier, Long &amp; Ebert 2006</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FBCB8488DFB2" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FBCB8488DFB2" pageId="17" pageNumber="571">
- EYL less than 9%
<materialsCitation id="3B033CC1FFF7FF9D45E1FBCB8488DFB2" ID-GBIF-Occurrence="3382541307" collectionCode="BDL" country="Chile" location="Southern" pageId="17" pageNumber="571" specimenCount="1" stateProvince="California">
<collectionCode id="ED7AAE59FFF7FF9D45E1FBCB80C0DF79" box="[416,467,1095,1116]" pageId="17" pageNumber="571">BDL</collectionCode>
; spine serrations very worn, posterior edge of spine serrated for last 6.513% of spine length (in adults); uniform dark brown to black across entire body with a lighter band over the snout (
<location id="8EB46047FFF7FF9D402EFBE985D4DF5F" LSID="urn:lsid:plazi:treatment:03C2878AFFF7FF9D44D6FF1B848FDED2:8EB46047FFF7FF9D402EFBE985D4DF5F" box="[1135,1223,1125,1146]" country="Chile" name="Southern" pageId="17" pageNumber="571" stateProvince="California">Southern</location>
<collectingRegion id="49AFF87EFFF7FF9D408EFBE98420DF5F" box="[1231,1331,1125,1146]" country="United States of America" name="California" pageId="17" pageNumber="571">California</collectingRegion>
, U.S.A. to
<collectingCountry id="F37C760CFFF7FF9D44A0FB0E8004DFB2" box="[225,279,1154,1175]" name="Chile" pageId="17" pageNumber="571">Chile</collectingCountry>
).............................................
<taxonomicName id="4C6B4D1FFFF7FF9D4705FB0E8488DFB2" ID-CoL="3N7F6" authority="James, Ebert, Long &amp; Didier 2009" authorityName="James, Ebert, Long &amp; Didier" authorityYear="2009" box="[836,1435,1154,1175]" class="Holocephali" family="Chimaeridae" genus="Hydrolagus" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="melanophasma">
<emphasis id="B91FEA8EFFF7FF9D4705FB0E8558DFB2" box="[836,1099,1154,1175]" italics="true" pageId="17" pageNumber="571">Hydrolagus melanophasma</emphasis>
<bibRefCitation id="EFFA4B6DFFF7FF9D4010FB0E8488DFB2" author="James, K. &amp; Ebert, D. A. &amp; Long, D. J. &amp; Didier, D. A." box="[1105,1435,1154,1175]" pageId="17" pageNumber="571" pagination="59 - 68" refId="ref13776" refString="James, K. C, Ebert, D. A., Long, D. J. &amp; Didier, D. A. (2009) A new species of chimaera, Hydrolagus melanophasma sp. nov. (Chondrichthyes: Chimaeriformes: Chimaeridae), from the eastern North Pacific. Zootaxa, 2218, 59 - 68." type="journal article" year="2009">James, Ebert, Long &amp; Didier 2009</bibRefCitation>
</taxonomicName>
</materialsCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E39FFF7FF9D44D6FB138488DE29" pageId="17" pageNumber="571">
<paragraph id="8BD4369CFFF7FF9D44D6FB138488DFF7" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FB138488DFF7" pageId="17" pageNumber="571">
8 Tooth-plates with ridges and knobs; eyes virtually above mouth; upper edge of caudal fin without denticles or tubercles (Southern
<collectingRegion id="49AFF87EFFF7FF9D4544FB31807ADFF7" box="[261,361,1213,1234]" country="United States of America" name="California" pageId="17" pageNumber="571">California</collectingRegion>
,
<collectingCountry id="F37C760CFFF7FF9D4533FB3180A4DFF7" box="[370,439,1213,1234]" name="United States of America" pageId="17" pageNumber="571">U.S.A.</collectingCountry>
, to
<collectingCountry id="F37C760CFFF7FF9D4598FB318318DFF7" box="[473,523,1213,1234]" name="Peru" pageId="17" pageNumber="571">Peru</collectingCountry>
).........................................
<taxonomicName id="4C6B4D1FFFF7FF9D4044FB318488DFF7" ID-CoL="6LFS7" authority="Goode &amp; Bean 1895" authorityName="Goode &amp; Bean" authorityYear="1895" box="[1029,1435,1213,1234]" class="Holocephali" family="Rhinochimaeridae" genus="Harriotta" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="raleighana">
<emphasis id="B91FEA8EFFF7FF9D4044FB3185C3DFF7" box="[1029,1232,1213,1234]" italics="true" pageId="17" pageNumber="571">Harriotta raleighana</emphasis>
Goode &amp; Bean 1895
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FB568488DE29" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FB568488DE29" pageId="17" pageNumber="571">- Tooth-plates nearly smooth; eyes distinctly behind level of mouth; caudal tubercles present along the upper edge of caudal fin .................................................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="309F2E39FFF7FF9D44D6FA99848FDED2" pageId="17" pageNumber="571">
<paragraph id="8BD4369CFFF7FF9D44D6FA998485DEA7" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FA998485DEA7" pageId="17" pageNumber="571">
9 Snout broad and paddle-shaped with fleshy tip; junction of supraorbital and infraorbital canals on ventral side of snout closer to the tip of the snout than to the nasal canal; ONC/TIO greater than 1.4, TIO/SWF less than 1.5, TIO/LNC less than 3.0; 2547 dorsal caudal tubercles in mature specimens; even dark brown color over entire body (
<collectingCountry id="F37C760CFFF7FF9D401FFAC385D4DE41" box="[1118,1223,1359,1380]" name="Costa Rica" pageId="17" pageNumber="571">Costa Rica</collectingCountry>
to
<collectingCountry id="F37C760CFFF7FF9D40A7FAC38407DE41" box="[1254,1300,1359,1380]" name="Peru" pageId="17" pageNumber="571">Peru</collectingCountry>
).............................................................
<taxonomicName id="4C6B4D1FFFF7FF9D4706FAE18485DEA7" ID-CoL="4S86C" authority="Compagno, Stehmann &amp; Ebert 1990" authorityName="Compagno, Stehmann, &amp; Ebert" authorityYear="1990" box="[839,1430,1389,1410]" class="Holocephali" family="Rhinochimaeridae" genus="Rhinochimaera" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="africana">
<emphasis id="B91FEA8EFFF7FF9D4706FAE18521DEA7" box="[839,1074,1389,1410]" italics="true" pageId="17" pageNumber="571">Rhinochimaera africana</emphasis>
<bibRefCitation id="EFFA4B6DFFF7FF9D4078FAE18485DEA7" author="Compagno, L. J. V. &amp; Stehmann, M. &amp; Ebert, D. A." box="[1081,1430,1389,1410]" pageId="17" pageNumber="571" pagination="201 - 222" refId="ref12743" refString="Compagno, L. J. V., Stehmann, M. &amp; Ebert, D. A. (1990) Rhinochimaera africana, a new longnose Chimaera from Southern Africa, with comments on the systematics and distribution of the genus Rhinochimaera Garman, 1901 (Chondrichthyes, Chimaeriformes, Rhinochimaeridae). African Journal of Marine Science, 9 (1), 201 - 222. http: // dx. doi. org / 10.2989 / 025776190784378646" type="journal article" year="1990">Compagno, Stehmann &amp; Ebert 1990</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD4369CFFF7FF9D44D6FA06848FDED2" blockId="17.[151,1436,362,1527]" pageId="17" pageNumber="571">
<keyLead id="309A95A9FFF7FF9D44D6FA06848FDED2" pageId="17" pageNumber="571">
- Snout narrow and conical shaped with the tip narrow and stiff; junction of supraorbital and infraorbital canals on ventral side of snout nearly equidistant between the tip of the snout and the nasal canal; ONC/TIO less than 1.4, TIO/SWF greater than 1.5, TIO/LNC greater than 3.0; 3662 dorsal caudal tubercles in mature specimens; pale brownish-gray body color with fins darker (
<collectingCountry id="F37C760CFFF7FF9D44A6FA6E8005DED2" box="[231,278,1506,1527]" name="Peru" pageId="17" pageNumber="571">Peru</collectingCountry>
)..............................................................
<taxonomicName id="4C6B4D1FFFF7FF9D404BFA6E848FDED2" ID-CoL="78TNC" authority="(Mitsukuri 1895)" baseAuthorityName="Mitsukuri" baseAuthorityYear="1895" box="[1034,1436,1506,1527]" class="Holocephali" family="Rhinochimaeridae" genus="Rhinochimaera" kingdom="Animalia" order="Chimaeriformes" pageId="17" pageNumber="571" phylum="Chordata" rank="species" species="pacifica">
<emphasis id="B91FEA8EFFF7FF9D404BFA6E85E2DED2" box="[1034,1265,1506,1527]" italics="true" pageId="17" pageNumber="571">Rhinochimaera pacifica</emphasis>
(Mitsukuri 1895)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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