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<document id="3757F833499C7BBA17E86111A04716B4" ID-DOI="10.11646/zootaxa.4092.2.4" ID-GBIF-Dataset="0847a84a-637d-4368-8bd8-0eb7171e5f59" ID-ISSN="1175-5326" ID-Zenodo-Dep="267859" ID-ZooBank="56072014-E67D-4866-A131-6693C2836B5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458118340065" checkinUser="plazi" docAuthor="Lambkin, Kevin J." docDate="2016" docId="03CD8786FF8EFF88FF3FF8B03A9EFD92" docLanguage="en" docName="zootaxa.4092.2.4.pdf.imf" docOrigin="Zootaxa 4092 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eoscartoides Evans 1956" docType="treatment" docVersion="8" lastPageNumber="209" masterDocId="FFF4FFFEFF8FFF8AFFA8FC593E79FFAF" masterDocTitle="Revision of the Dysmorphoptilidae (Hemiptera: Cicadomorpha: Prosboloidea) of the Queensland Triassic — Part 2" masterLastPageNumber="218" masterPageNumber="207" pageNumber="208" updateTime="1698184854291" updateUser="plazi">
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<mods:title id="CEFB09B004925CF99A967B447CE1FAFC">Revision of the Dysmorphoptilidae (Hemiptera: Cicadomorpha: Prosboloidea) of the Queensland Triassic — Part 2</mods:title>
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<treatment id="03CD8786FF8EFF88FF3FF8B03A9EFD92" ID-DOI="http://doi.org/10.5281/zenodo.6074519" ID-GBIF-Taxon="117914068" ID-Zenodo-Dep="6074519" LSID="urn:lsid:plazi:treatment:03CD8786FF8EFF88FF3FF8B03A9EFD92" httpUri="http://treatment.plazi.org/id/03CD8786FF8EFF88FF3FF8B03A9EFD92" lastPageId="2" lastPageNumber="209" pageId="1" pageNumber="208">
<subSubSection id="C37E651BFF8EFF8BFF3FF8B03FA0FAE9" pageId="1" pageNumber="208" type="nomenclature">
<paragraph id="8BDB3690FF8EFF8BFF3FF8B03FB6FAAC" blockId="1.[151,463,1257,1283]" box="[151,463,1257,1283]" pageId="1" pageNumber="208">
<heading id="D09381FCFF8EFF8BFF3FF8B03FB6FAAC" bold="true" box="[151,463,1257,1283]" fontSize="11" level="1" pageId="1" pageNumber="208" reason="1">
<taxonomicName id="4C644D13FF8EFF8BFF3FF8B03FB6FAAC" ID-CoL="8JL4M" authority="Evans, 1956" authorityName="Evans" authorityYear="1956" box="[151,463,1257,1283]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA82FF8EFF8BFF3FF8B03FB6FAAC" bold="true" box="[151,463,1257,1283]" pageId="1" pageNumber="208">
<emphasis id="B910EA82FF8EFF8BFF3FF8B03F4BFAAC" bold="true" box="[151,306,1257,1283]" italics="true" pageId="1" pageNumber="208">Eoscartoides</emphasis>
Evans, 1956
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDB3690FF8EFF8BFF3FF9693FA0FAE9" blockId="1.[151,887,1328,1380]" box="[151,473,1328,1350]" pageId="1" pageNumber="208">
<taxonomicName id="4C644D13FF8EFF8BFF3FF9693FACFAE9" authority="Evans, 1956: 220" authorityName="Evans" authorityPageNumber="220" authorityYear="1956" box="[151,469,1328,1350]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA82FF8EFF8BFF3FF9693F62FAE9" box="[151,283,1328,1350]" italics="true" pageId="1" pageNumber="208">Eoscartoides</emphasis>
Evans, 1956: 220
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E651BFF8EFF8BFF3FF9163F8DF809" pageId="1" pageNumber="208" type="materials_examined">
<paragraph id="8BDB3690FF8EFF8BFF3FF9163D0EFACB" blockId="1.[151,887,1328,1380]" box="[151,887,1358,1380]" pageId="1" pageNumber="208">
<emphasis id="B910EA82FF8EFF8BFF3FF9163F5BFACB" bold="true" box="[151,290,1358,1380]" pageId="1" pageNumber="208">
<typeStatus id="54DF8832FF8EFF8BFF3FF9163EB5FACB" box="[151,204,1359,1380]" pageId="1" pageNumber="208">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C644D13FF8EFF8BFE81F9173C05FACB" authority="Evans, 1956" authorityName="Evans" authorityYear="1956" box="[297,636,1358,1380]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="species" species="bryani">
<emphasis id="B910EA82FF8EFF8BFE81F9173F8FFACB" box="[297,502,1358,1380]" italics="true" pageId="1" pageNumber="208">Eoscartoides bryani</emphasis>
Evans, 1956
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="8BDB3690FF8EFF8BFF3FF9D43C0DFA0D" blockId="1.[151,1035,1420,1473]" box="[151,628,1420,1442]" pageId="1" pageNumber="208">
<emphasis id="B910EA82FF8EFF8BFF3FF9D43F64FA0D" box="[151,285,1421,1442]" italics="true" pageId="1" pageNumber="208">Mesonirvana</emphasis>
Evans, 1956: 191192.
<emphasis id="B910EA82FF8EFF8BFDBAF9D53C0DFA0D" bold="true" box="[530,628,1420,1442]" pageId="1" pageNumber="208">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8BDB3690FF8EFF8BFF3FF9F23A72FA6E" blockId="1.[151,1035,1420,1473]" box="[151,1035,1451,1473]" pageId="1" pageNumber="208">
<emphasis id="B910EA82FF8EFF8BFF3FF9F23F5BFA6E" bold="true" box="[151,290,1451,1473]" pageId="1" pageNumber="208">
<typeStatus id="54DF8832FF8EFF8BFF3FF9F23EB5FA6F" box="[151,204,1451,1472]" pageId="1" pageNumber="208">Type</typeStatus>
species.
</emphasis>
<emphasis id="B910EA82FF8EFF8BFE81F9F53C7CFA6E" box="[297,517,1451,1473]" italics="true" pageId="1" pageNumber="208">Mesonirvana abrupta</emphasis>
Evans, 1956: 192, fig. 5E, by original designation.
</paragraph>
<paragraph id="8BDB3690FF8EFF8BFF3FF9B13C32FA51" blockId="1.[151,1317,1512,1564]" box="[151,587,1512,1534]" pageId="1" pageNumber="208">
<emphasis id="B910EA82FF8EFF8BFF3FF9B13F58FA51" box="[151,289,1512,1534]" italics="true" pageId="1" pageNumber="208">Mesocixiodes</emphasis>
Tillyard, 1922: 462 (
<emphasis id="B910EA82FF8EFF8BFE51F9B03C44FA51" box="[505,573,1513,1534]" italics="true" pageId="1" pageNumber="208">partim</emphasis>
).
</paragraph>
<paragraph id="8BDB3690FF8EFF8BFF3FFA5E3B5CF9B3" blockId="1.[151,1317,1512,1564]" box="[151,1317,1542,1564]" pageId="1" pageNumber="208">
<emphasis id="B910EA82FF8EFF8BFF3FFA5E3F5BF9B3" bold="true" box="[151,290,1542,1564]" pageId="1" pageNumber="208">
<typeStatus id="54DF8832FF8EFF8BFF3FFA5E3EB5F9B3" box="[151,204,1543,1564]" pageId="1" pageNumber="208">Type</typeStatus>
species.
</emphasis>
<emphasis id="B910EA82FF8EFF8BFE81FA5F3C4FF9B3" box="[297,566,1542,1564]" italics="true" pageId="1" pageNumber="208">Mesocixiodes termioneura</emphasis>
Tillyard, 1922: 462, text fig. 82, plate 53, fig. 34, by original designation.
</paragraph>
<paragraph id="8BDB3690FF8EFF8BFF3FFA103F8DF809" blockId="1.[151,1437,1609,2030]" pageId="1" pageNumber="208">
<emphasis id="B910EA82FF8EFF8BFF3FFA103FDEF9CE" bold="true" box="[151,423,1609,1633]" pageId="1" pageNumber="208">Diagnostic characters.</emphasis>
Tegmen over
<quantity id="4C9C9B75FF8EFF8BFDF0FA133CD5F9CE" box="[600,684,1610,1634]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="208" unit="mm" value="11.0">11 mm</quantity>
long, lobate; precostal carina and marginal membrane distinct; hypocostal carina short; punctation very fine, somewhat coarser basally and posterior to Pcu; hyposubcostal carina long, only very slightly concave; strigil in basal costal space large and well developed, quite coarsely rugose, extending in an arc from just above R to costal margin; no swellings detected on hind margin of clavus, its proximal corner strongly bent downwards; stem of RA very short, its primary fork just beyond clavus ½ length, proximal to level of CuA fork; stem of RA1 directed anteriorly in line with stem of RA, entering costal margin well before tegmen apex, with at least 2 anterior branches; RA2 extensive, occupying a large area antero-apically, with at least 3 anterior branches; RA2 and
<collectingCountry id="F3737600FF8EFF8BFDFBFB1F3C0CF8F2" box="[595,629,1862,1885]" name="Poland" pageId="1" pageNumber="208">RP</collectingCountry>
joined by a crossvein; M1+2 deeply forked; M3+4 simple, or with a terminal fork only; M and CuA joined basally by a short crossvein (the “arculus”); apical m-cu inclined; CuA1 gently concave; CuA2 straight.
</paragraph>
</subSubSection>
<subSubSection id="C37E651BFF8EFF88FF6FFBEB3A9EFD92" lastPageId="2" lastPageNumber="209" pageId="1" pageNumber="208" type="description">
<paragraph id="8BDB3690FF8EFF88FF6FFBEB3F07FEEF" blockId="1.[151,1437,1609,2030]" lastBlockId="2.[151,1436,151,573]" lastPageId="2" lastPageNumber="209" pageId="1" pageNumber="208">
<emphasis id="B910EA82FF8EFF8BFF6FFBEB3F68F866" bold="true" box="[199,273,1970,1993]" pageId="1" pageNumber="208">Notes.</emphasis>
The
<typeStatus id="54DF8832FF8EFF8BFEFFFBEB3FFEF866" box="[343,391,1970,1993]" pageId="1" pageNumber="208">type</typeStatus>
species of
<taxonomicName id="4C644D13FF8EFF8BFDB0FBE83CD3F865" box="[536,682,1969,1994]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA82FF8EFF8BFDB0FBE83CD3F865" box="[536,682,1969,1994]" italics="true" pageId="1" pageNumber="208">Eoscartoides</emphasis>
</taxonomicName>
and
<emphasis id="B910EA82FF8EFF8BFD46FBEB3DFBF866" box="[750,898,1970,1993]" italics="true" pageId="1" pageNumber="208">Mesonirvana</emphasis>
are synonyms.
<collectingCountry id="F3737600FF8EFF8BFBEDFBEB3A1DF865" box="[1093,1124,1970,1994]" name="American Samoa" pageId="1" pageNumber="208">As</collectingCountry>
both generic names were established in the same paper, as first reviser (ICZN Article 24), I have chosen
<taxonomicName id="4C644D13FF8EFF8BFBA2FB8C3AE4F841" box="[1034,1181,2005,2030]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA82FF8EFF8BFBA2FB8C3AE4F841" box="[1034,1181,2005,2030]" italics="true" pageId="1" pageNumber="208">Eoscartoides</emphasis>
</taxonomicName>
as the senior synonym on the grounds that its
<typeStatus id="54DF8832FF8DFF88FE3EFCC13FBFFF00" box="[406,454,152,175]" pageId="2" pageNumber="209">type</typeStatus>
specimen is better preserved and that all subsequent published specimens (Evans 1961) have been identified as
<taxonomicName id="4C644D13FF8DFF88FE0BFCE53CFFFF7A" box="[419,646,188,213]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="species" species="bryani">
<emphasis id="B910EA82FF8DFF88FE0BFCE53CFFFF7A" box="[419,646,188,213]" italics="true" pageId="2" pageNumber="209">Eoscartoides bryani</emphasis>
</taxonomicName>
. The
<typeStatus id="54DF8832FF8DFF88FD60FCE43C81FF7B" box="[712,760,189,212]" pageId="2" pageNumber="209">type</typeStatus>
specimen of the
<typeStatus id="54DF8832FF8DFF88FC14FCE43D95FF7B" box="[956,1004,189,212]" pageId="2" pageNumber="209">type</typeStatus>
species,
<emphasis id="B910EA82FF8DFF88FBFCFCE43B30FF7A" box="[1108,1353,188,213]" italics="true" pageId="2" pageNumber="209">Mesonirvana abrupta</emphasis>
Evans, preserves only the apical region of the tegmen and is somewhat distorted, whereas that of
<taxonomicName id="4C644D13FF8DFF88FB15FCB93B50FF57" box="[1213,1321,223,248]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="species" species="bryani">
<emphasis id="B910EA82FF8DFF88FB15FCB93B50FF57" box="[1213,1321,223,248]" italics="true" pageId="2" pageNumber="209">E. bryani</emphasis>
</taxonomicName>
is almost complete, and in particular shows the characteristic strigil in the basal costal space (see photographs at pages 44 and 56 of Jell 2004).
</paragraph>
<paragraph id="8BDB3690FF8DFF88FF6FFD153A9EFD92" blockId="2.[151,1436,151,573]" pageId="2" pageNumber="209">
The large and distinctly shaped strigil, the very short stem of RA, and the basic pattern of forking of M (M1+2 deep, M3+4 simple) are considered as the apomorphies for
<taxonomicName id="4C644D13FF8DFF88FC94FD363DB7FE27" box="[828,974,367,392]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA82FF8DFF88FC94FD363DB7FE27" box="[828,974,367,392]" italics="true" pageId="2" pageNumber="209">Eoscartoides</emphasis>
</taxonomicName>
. The structure and shape of the strigil is apparently the same as that of the unnamed Triassic sspecies from the Madygen Formation of
<collectingCountry id="F3737600FF8DFF88FAB3FDCC3BE5FE03" box="[1307,1436,405,428]" name="Kyrgyzstan" pageId="2" pageNumber="209">Kyrgyzstan</collectingCountry>
illustrated by Shcherbakov and Popov (2002, fig.184) which also appears to have similar venation to
<taxonomicName id="4C644D13FF8DFF88FAABFDEE3BECFE7F" box="[1283,1429,439,464]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA82FF8DFF88FAABFDEE3BECFE7F" box="[1283,1429,439,464]" italics="true" pageId="2" pageNumber="209">Eoscartoides</emphasis>
</taxonomicName>
. Even though only one specimen of each of the three species of
<taxonomicName id="4C644D13FF8DFF88FCD4FD853A77FE5A" box="[892,1038,476,501]" class="Insecta" family="Dysmorphoptilidae" genus="Eoscartoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA82FF8DFF88FCD4FD853A77FE5A" box="[892,1038,476,501]" italics="true" pageId="2" pageNumber="209">Eoscartoides</emphasis>
</taxonomicName>
preserves a substantial part of the clavus (QM F3697a/b, GSQ 318a, QM
<accessionNumber id="9437AB73FF8DFF88FDF9FDA63CD2FDB7" box="[593,683,511,536]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F58592" pageId="2" pageNumber="209" type="EnaNcbi">F58592</accessionNumber>
), the peculiar bend at the proximal corner is present in all three and is probably the basal part of the commissural carina (Dmitry Shcherbakov, pers com.) (Figs 3,4,9).
</paragraph>
</subSubSection>
</treatment>
</document>