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<document id="F5471D63BDE90C2A3E4B3D015934E2F4" ID-CLB-Dataset="55626" ID-DOI="10.11646/zootaxa.5133.4.2" ID-GBIF-Dataset="0cd92d63-eeac-48d6-9c9d-270f866de7d3" ID-ISSN="1175-5326" ID-Zenodo-Dep="6530956" ID-ZooBank="BD6A434A-21D7-49FC-A09F-FEF4BC740B14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652085552068" checkinUser="plazi" docAuthor="Joharchi, Omid, Döker, Ismail &amp; Khaustov, Vladimir A." docDate="2022" docId="D25AD024FFA8983201C7AB5372D3FCC4" docLanguage="en" docName="zootaxa.5133.4.2.pdf" docOrigin="Zootaxa 5133 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmolaelaps latisetis Joharchi 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="498" masterDocId="2E63A85CFFA0983E0150AC5D7325FFD1" masterDocTitle="New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from Russia, with a review of the Russian species of the genus" masterLastPageNumber="508" masterPageNumber="486" pageNumber="494" updateTime="1699329983419" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9CB1D51572B46A59EDE71101786EDEA1">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from Russia, with a review of the Russian species of the genus</mods:title>
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<mods:namePart id="E70C2B10F966266007C251ECAD63E66C">Joharchi, Omid</mods:namePart>
<mods:affiliation id="D17A0691F54FCB1CFE0726231186B7AE">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Semakova Str. 10, 625003 Tyumen, Russia.</mods:affiliation>
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<mods:namePart id="1DBEB93D55623F7F46EFD40FD62EF4DB">Döker, Ismail</mods:namePart>
<mods:nameIdentifier id="42514009E698684B3B4C942260519EBB" type="ORCID">0000-0002-2741-4946</mods:nameIdentifier>
<mods:affiliation id="E2973162971C9E9EFAE72C11133A20FF">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Semakova Str. 10, 625003 Tyumen, Russia. &amp; Cukurova University, Agricultural Faculty, Department of Plant Protection, Acarology Laboratory, 01330, Adana, Turkey. https: // orcid. org / 0000 - 0002 - 2741 - 4946 (Joharchi) https: // orcid. org / 0000 - 0002 - 1412 - 1554 (Döker) https: // orcid. org / 0000 - 0003 - 2831 - 7213 (Khaustov)</mods:affiliation>
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<mods:namePart id="8647B2C0DDA9FA55F89F35ABD1577482">Khaustov, Vladimir A.</mods:namePart>
<mods:affiliation id="704C671D7701FBD180AC01FA02881C7F">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Semakova Str. 10, 625003 Tyumen, Russia.</mods:affiliation>
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<mods:title id="07B053605FC3E536E6401410A8C19A63">Zootaxa</mods:title>
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<mods:date id="AC94B458DEBBF21BC5D66D072A8394DD">2022</mods:date>
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<mods:number id="7ED51EFE1CE21ADD95A1B26E4283BEEF">2022-05-06</mods:number>
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<treatment id="D25AD024FFA8983201C7AB5372D3FCC4" ID-DOI="http://doi.org/10.5281/zenodo.6530966" ID-GBIF-Taxon="195277957" ID-Zenodo-Dep="6530966" LSID="urn:lsid:plazi:treatment:D25AD024FFA8983201C7AB5372D3FCC4" httpUri="http://treatment.plazi.org/id/D25AD024FFA8983201C7AB5372D3FCC4" lastPageId="12" lastPageNumber="498" pageId="8" pageNumber="494">
<subSubSection id="12E932B9FFA8983601C7AB537159F8F8" box="[151,636,1806,1833]" pageId="8" pageNumber="494" type="nomenclature">
<paragraph id="5A4C6132FFA8983601C7AB537159F8F8" blockId="8.[151,636,1806,1869]" box="[151,636,1806,1833]" pageId="8" pageNumber="494">
<heading id="0104D65EFFA8983601C7AB537159F8F8" bold="true" box="[151,636,1806,1833]" fontSize="11" level="1" pageId="8" pageNumber="494" reason="1">
<emphasis id="6887BD20FFA8983601C7AB537159F8F8" bold="true" box="[151,636,1806,1833]" pageId="8" pageNumber="494">
<taxonomicName id="9DF31AB1FFA8983601C7AB537132F8F9" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[151,535,1806,1833]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="latisetis" status="sp. nov.">
<emphasis id="6887BD20FFA8983601C7AB537284F8F8" bold="true" box="[151,417,1806,1833]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps latisetis</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA89836034FAB527159F8F8" box="[543,636,1807,1833]" pageId="8" pageNumber="494" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA8983601C7AB6E726EF89C" box="[151,331,1842,1869]" pageId="8" pageNumber="494" type="description">
<paragraph id="5A4C6132FFA8983601C7AB6E726EF89C" blockId="8.[151,636,1806,1869]" box="[151,331,1842,1869]" pageId="8" pageNumber="494">
(
<figureCitation id="C2C87DB7FFA8983601CFAB6E7266F89C" box="[159,323,1842,1869]" captionStart-0="FIGURES 2526" captionStart-1="FIGURES 2729" captionStart-2="FIGURES 3040" captionStart-3="FIGURE 4144" captionStartId-0="9.[151,264,1943,1967]" captionStartId-1="10.[151,264,1468,1492]" captionStartId-2="13.[151,264,1874,1898]" captionStartId-3="14.[151,250,1943,1967]" captionTargetBox-0="[193,1391,190,1899]" captionTargetBox-1="[171,1417,193,1432]" captionTargetBox-2="[214,1374,184,1849]" captionTargetBox-3="[183,1406,196,1906]" captionTargetId-0="figure-19@9.[179,1408,181,1919]" captionTargetId-1="figure-18@10.[151,1436,181,1444]" captionTargetId-2="figure-19@13.[212,1375,181,1851]" captionTargetId-3="figure-18@14.[175,1413,181,1919]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." captionText-1="FIGURES 2729. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." captionText-2="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." captionText-3="FIGURE 4144. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.6530976" figureDoi-1="http://doi.org/10.5281/zenodo.6530978" figureDoi-2="http://doi.org/10.5281/zenodo.6531011" figureDoi-3="http://doi.org/10.5281/zenodo.6530984" httpUri-0="https://zenodo.org/record/6530976/files/figure.png" httpUri-1="https://zenodo.org/record/6530978/files/figure.png" httpUri-2="https://zenodo.org/record/6531011/files/figure.png" httpUri-3="https://zenodo.org/record/6530984/files/figure.png" pageId="8" pageNumber="494">Figures 2544</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA8983601C7AB26776DF80C" pageId="8" pageNumber="494" type="materials_examined">
<paragraph id="5A4C6132FFA8983601C7AB26776DF80C" blockId="8.[151,1437,1914,2014]" pageId="8" pageNumber="494">
<emphasis id="6887BD20FFA8983601C7AB267261F845" bold="true" box="[151,324,1914,1941]" pageId="8" pageNumber="494">Type material.</emphasis>
<materialsCitation id="EA9B6B6FFFA898360019AB26728CF80C" ID-GBIF-Occurrence="3763889303" collectedFrom="in the nest of Lasius flavus (Fabricius, 1782) (Hymenoptera: Formicidae)" collectingDate="2021-08-11" collectionCode="TUMZ" collectorName="O. Joharchi" country="Russia" latitude="46.71375" location="on the coast of the Sea of Japan (East Sea)" longLatPrecision="10" longitude="141.88722" pageId="8" pageNumber="494" specimenCount="1" specimenCount-female="1" stateProvince="Sakhalin Island" typeStatus="holotype">
<typeStatus id="8548DF90FFA898360019AB267291F844" box="[329,436,1915,1941]" pageId="8" pageNumber="494" type="holotype">Holotype</typeStatus>
,
<specimenCount id="4CF5AABBFFA8983600EEAB267128F844" box="[446,525,1915,1941]" pageId="8" pageNumber="494" type="female">female</specimenCount>
,
<location id="5F2C37E9FFA898360347AB2670CCF844" LSID="urn:lsid:plazi:treatment:D25AD024FFA8983201C7AB5372D3FCC4:5F2C37E9FFA898360347AB2670CCF844" box="[535,1001,1915,1941]" country="Russia" latitude="46.71375" longLatPrecision="10" longitude="141.88722" name="on the coast of the Sea of Japan (East Sea)" pageId="8" pageNumber="494" stateProvince="Sakhalin Island">on the coast of the Sea of Japan (East Sea)</location>
,
<collectingRegion id="9837AFD0FFA8983602A3AB267787F844" box="[1011,1186,1915,1941]" pageId="8" pageNumber="494">Sakhalin Island</collectingRegion>
,
<collectingCountry id="22E421A2FFA8983605FCAB2677DCF844" box="[1196,1273,1915,1941]" name="Russia" pageId="8" pageNumber="494">Russia</collectingCountry>
,
<geoCoordinate id="3FC707F5FFA898360453AB2776B8F844" box="[1283,1437,1914,1941]" degrees="46" direction="north" minutes="42" orientation="latitude" pageId="8" pageNumber="494" precision="1" seconds="49.5" value="46.71375">46°42'49.5&quot;N</geoCoordinate>
<geoCoordinate id="3FC707F5FFA8983601C7ABC3720DF868" box="[151,296,1950,1977]" degrees="141" direction="east" minutes="53" orientation="longitude" pageId="8" pageNumber="494" precision="15" seconds="14" value="141.88722">141°53'14&quot;E</geoCoordinate>
,
<date id="2E4D47F2FFA898360068ABC272D1F868" box="[312,500,1951,1977]" pageId="8" pageNumber="494" value="2021-08-11">
<collectingDate id="3E09BE1AFFA898360068ABC272D1F868" box="[312,500,1951,1977]" pageId="8" pageNumber="494" value="2021-08-11">11 August 2021</collectingDate>
</date>
,
<collectorName id="F70604E4FFA898360353ABC271ADF868" box="[515,648,1951,1977]" pageId="8" pageNumber="494">O. Joharchi</collectorName>
coll.,
<collectedFrom id="E9EF22CFFFA898360387ABC27205F80F" pageId="8" pageNumber="494">
in the nest of
<taxonomicName id="9DF31AB1FFA8983602D0ABC277CDF868" authority="(Fabricius, 1782)" baseAuthorityName="Fabricius" baseAuthorityYear="1782" box="[896,1256,1951,1977]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="flavus">
<emphasis id="6887BD20FFA8983602D0ABC27732F868" box="[896,1047,1951,1977]" italics="true" pageId="8" pageNumber="494">Lasius flavus</emphasis>
(Fabricius, 1782)
</taxonomicName>
(
<taxonomicName id="9DF31AB1FFA8983605AAABC276BCF868" box="[1274,1433,1951,1977]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="9DF31AB1FFA8983601C7AB9E723CF80F" box="[151,281,1987,2014]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
)
</collectedFrom>
(in
<collectionCode id="3CE2F9F7FFA89836001DAB9E7287F80C" box="[333,418,1987,2013]" pageId="8" pageNumber="494">TUMZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EA9B6B6FFFA8983600E3AB9E7760F80C" ID-GBIF-Occurrence="3763889302" box="[435,1093,1987,2013]" pageId="8" pageNumber="494" typeStatus="paratype">
<typeStatus id="8548DF90FFA8983600E3AB9E7101F80C" box="[435,548,1987,2013]" pageId="8" pageNumber="494" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="4CF5AABBFFA89836037FAB9E719FF80C" box="[559,698,1987,2013]" pageId="8" pageNumber="494" type="female">two females</specimenCount>
, same data as
<typeStatus id="8548DF90FFA89836020AAB9E7099F80C" box="[858,956,1987,2013]" pageId="8" pageNumber="494" type="holotype">holotype</typeStatus>
(in
<collectionCode id="3CE2F9F7FFA8983602B8AB9E7718F80C" box="[1000,1085,1987,2013]" pageId="8" pageNumber="494">TUMZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="0E8C31BAFFA9983701C7ABCA7237F805" ID-DOI="http://doi.org/10.5281/zenodo.6530976" ID-Zenodo-Dep="6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="9" pageNumber="495" startId="9.[151,264,1943,1967]" targetBox="[193,1391,190,1899]" targetPageId="9">
<paragraph id="5A4C6132FFA9983701C7ABCA7237F805" blockId="9.[151,1437,1943,2004]" pageId="9" pageNumber="495">
<emphasis id="6887BD20FFA9983701C7ABCA7276F87E" bold="true" box="[151,339,1943,1967]" pageId="9" pageNumber="495">FIGURES 2526.</emphasis>
<taxonomicName id="9DF31AB1FFA998370007ABC571B1F861" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[343,660,1944,1968]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="495" phylum="Arthropoda" rank="species" species="latisetis" status="sp. nov.">
<emphasis id="6887BD20FFA998370007ABC5711FF861" box="[343,570,1944,1968]" italics="true" pageId="9" pageNumber="495">Cosmolaelaps latisetis</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA9983703C9ABC571CCF87E" box="[665,745,1944,1968]" pageId="9" pageNumber="495" rank="species">
<emphasis id="6887BD20FFA9983703C9ABC571C6F861" bold="true" box="[665,739,1944,1968]" pageId="9" pageNumber="495">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale.
</paragraph>
</caption>
<caption id="0E8C31BAFFAA983401C7A9E17603FA05" ID-DOI="http://doi.org/10.5281/zenodo.6530978" ID-Zenodo-Dep="6530978" box="[151,1318,1468,1492]" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="10" pageNumber="496" startId="10.[151,264,1468,1492]" targetBox="[171,1417,193,1432]" targetPageId="10">
<paragraph id="5A4C6132FFAA983401C7A9E17603FA05" blockId="10.[151,1318,1468,1492]" box="[151,1318,1468,1492]" pageId="10" pageNumber="496">
<emphasis id="6887BD20FFAA983401C7A9E17271FA05" bold="true" box="[151,340,1468,1492]" pageId="10" pageNumber="496">FIGURES 2729.</emphasis>
<taxonomicName id="9DF31AB1FFAA9834000AA9E171BEFA05" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[346,667,1468,1492]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="latisetis" status="sp. nov.">
<emphasis id="6887BD20FFAA9834000AA9E1711AFA05" box="[346,575,1468,1492]" italics="true" pageId="10" pageNumber="496">Cosmolaelaps latisetis</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFAA983403F1A9E171D6FA05" box="[673,755,1468,1492]" pageId="10" pageNumber="496" rank="species">
<emphasis id="6887BD20FFAA983403F1A9E171C8FA05" bold="true" box="[673,749,1468,1492]" pageId="10" pageNumber="496">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 27, subcapitulum; 28, epistome; 29, chelicera.
</paragraph>
</caption>
<subSubSection id="12E932B9FFAA98340197AA5F71BBF855" pageId="10" pageNumber="496" type="diagnosis">
<paragraph id="5A4C6132FFAA98340197AA5F71BBF855" blockId="10.[151,1437,1538,2033]" pageId="10" pageNumber="496">
<emphasis id="6887BD20FFAA98340197AA5F721FF9CD" bold="true" box="[199,314,1538,1564]" pageId="10" pageNumber="496">Diagnosis</emphasis>
(adult female). Dorsal shield well reticulate, with 39 pairs of scimitar-like setae, reaching well past base of next posterior setae,
<emphasis id="6887BD20FFAA983400B3AA7A7124F990" box="[483,513,1575,1601]" italics="true" pageId="10" pageNumber="496">Z5</emphasis>
shorter than
<emphasis id="6887BD20FFAA983403CBAA7A7193F990" box="[667,694,1575,1601]" italics="true" pageId="10" pageNumber="496">J5</emphasis>
(ratio of J5/Z5 ≈ 1.5); presternal area poorly sclerotised, sternal shield with lineate-reticulate ornamentation on lateral surface, bearing three pairs of long setae, extending slightly beyond of next posterior setae, ratio of shield length/width (at broadest level) ≃ 0.65; genital shield ratio of length/ width (at broadest level) ≃ 1.9, shield ornamented posteromedially by seven cells flanked by a median Λ-shaped ornamentation, anal shield ratio of length/width (at broadest level) ≃ 1.2, post-anal seta thickened, sparsely barbed, apically bent (hook shaped); distance between genital and anal shields is shorter than length of anal shield, almost all opisthogastric setae well thickened (especially
<emphasis id="6887BD20FFAA9834038BAAA27026F8C8" box="[731,771,1791,1817]" italics="true" pageId="10" pageNumber="496">Jv5</emphasis>
, apically bent) and arising on minute sclerotised platelet, except
<emphasis id="6887BD20FFAA983401B7AB7F720FF8ED" box="[231,298,1826,1853]" italics="true" pageId="10" pageNumber="496">Jv12</emphasis>
and
<emphasis id="6887BD20FFAA9834000FAB7F72ACF8ED" box="[351,393,1826,1852]" italics="true" pageId="10" pageNumber="496">Zv1</emphasis>
. Fixed digit of chelicera with five teeth, including two large proximal teeth (posterior to pilus dentilis). Legs without spatulate setae but most setae conspicuously thickened and some inserted on small tubercles. Legs I and IV shorter than length of idiosoma.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFAA98320197ABD372ADFD1C" lastPageId="12" lastPageNumber="498" pageId="10" pageNumber="496" type="description">
<paragraph id="5A4C6132FFAA98340197ABD372D8F879" blockId="10.[151,1437,1538,2033]" box="[199,509,1934,1960]" pageId="10" pageNumber="496">
<emphasis id="6887BD20FFAA98340197ABD37292F879" bold="true" box="[199,439,1934,1960]" pageId="10" pageNumber="496">Description. Female</emphasis>
(n=3)
</paragraph>
<paragraph id="5A4C6132FFAA98350197ABEF7075FEB4" blockId="10.[151,1437,1538,2033]" lastBlockId="11.[151,1437,151,2014]" lastPageId="11" lastPageNumber="497" pageId="10" pageNumber="496">
<emphasis id="6887BD20FFAA98340197ABEF72A6F81D" box="[199,387,1970,1996]" italics="true" pageId="10" pageNumber="496">Dorsal idiosoma</emphasis>
(
<figureCitation id="C2C87DB7FFAA983400C2ABEF72CCF81C" box="[402,489,1970,1997]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="10" pageNumber="496">Figs 25</figureCitation>
,
<figureCitation id="C2C87DB7FFAA983400A6ABEF714CF81D" box="[502,617,1970,1996]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="10" pageNumber="496">30, 32, 33</figureCitation>
). Dorsal shield semi-circular, large, reticulate throughout, 498512 long, 404410 wide, covering entire dorsal idiosoma. Shield with 39 pairs of very long scimitar-like setae, 22 podonotal, 17 opisthonotal, including two pairs of
<emphasis id="6887BD20FFAB98350301ACCA7148FF61" box="[593,621,151,176]" italics="true" pageId="11" pageNumber="497">Zx</emphasis>
setae between
<emphasis id="6887BD20FFAB98350247ACCA7006FF61" box="[791,803,151,176]" italics="true" pageId="11" pageNumber="497">J</emphasis>
and
<emphasis id="6887BD20FFAB9835020AACCA704FFF61" box="[858,874,151,176]" italics="true" pageId="11" pageNumber="497">Z</emphasis>
setae, and three unpaired setae
<emphasis id="6887BD20FFAB9835059EACCA77C2FF61" box="[1230,1255,151,176]" italics="true" pageId="11" pageNumber="497">Jx</emphasis>
between
<emphasis id="6887BD20FFAB98350402ACCA7649FF60" box="[1362,1388,151,177]" italics="true" pageId="11" pageNumber="497">J2</emphasis>
and
<emphasis id="6887BD20FFAB983501C7ACE67397FF04" box="[151,178,187,213]" italics="true" pageId="11" pageNumber="497">J4</emphasis>
, setae reaching well past base of next posterior setae, mostly subequal in length (6073), all dorsal setae very thick and slightly swollen at base, except
<emphasis id="6887BD20FFAB98350329AC8271ABFF28" box="[633,654,223,249]" italics="true" pageId="11" pageNumber="497">j1</emphasis>
(2023) and
<emphasis id="6887BD20FFAB9835027BAC827061FF28" box="[811,836,223,249]" italics="true" pageId="11" pageNumber="497">z1</emphasis>
(1720) simple and shortest, ratio of
<emphasis id="6887BD20FFAB983505A9AC827631FF28" box="[1273,1300,223,249]" italics="true" pageId="11" pageNumber="497">J5</emphasis>
(3740)/Z5 (2225) ≈ 1.5,
<emphasis id="6887BD20FFAB9835006BAD5E727CFECC" box="[315,345,259,285]" italics="true" pageId="11" pageNumber="497">Z5</emphasis>
sparsely barbed (
<figureCitation id="C2C87DB7FFAB9835034BAD5E7154FECC" box="[539,625,258,285]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 25</figureCitation>
,
<figureCitation id="C2C87DB7FFAB9835032CAD5E71C8FECC" box="[636,749,259,285]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">30, 32, 33</figureCitation>
). Dorsal shield with 23 pairs of pore-like structures, including seven pairs of gland openings (
<emphasis id="6887BD20FFAB98350351AD7A710EFE90" box="[513,555,295,321]" italics="true" pageId="11" pageNumber="497">gd1</emphasis>
,
<emphasis id="6887BD20FFAB9835036CAD7A7143FE90" box="[572,614,295,321]" italics="true" pageId="11" pageNumber="497">gd2</emphasis>
,
<emphasis id="6887BD20FFAB98350326AD7A7184FE90" box="[630,673,295,321]" italics="true" pageId="11" pageNumber="497">gd4</emphasis>
,
<emphasis id="6887BD20FFAB983503E1AD7A71FEFE90" box="[689,731,295,321]" italics="true" pageId="11" pageNumber="497">gd5</emphasis>
,
<emphasis id="6887BD20FFAB983503BCAD7B7033FE91" box="[748,790,294,320]" italics="true" pageId="11" pageNumber="497">gd6</emphasis>
,
<emphasis id="6887BD20FFAB98350276AD7A7075FE90" box="[806,848,295,321]" italics="true" pageId="11" pageNumber="497">gd8</emphasis>
,
<emphasis id="6887BD20FFAB98350231AD7A70AFFE90" box="[865,906,295,321]" italics="true" pageId="11" pageNumber="497">gd9</emphasis>
) and 16 pairs of poroids. Shape, position and relative length and shape of setae shown in
<figureCitation id="C2C87DB7FFAB983503D0AD1671D2FEB4" box="[640,759,330,357]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figures 25</figureCitation>
and
<figureCitation id="C2C87DB7FFAB9835027DAD167069FEB4" box="[813,844,331,357]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">30</figureCitation>
.
</paragraph>
<paragraph id="5A4C6132FFAB98350197AD327089FA4C" blockId="11.[151,1437,151,2014]" pageId="11" pageNumber="497">
<emphasis id="6887BD20FFAB98350197AD3272A2FE58" box="[199,391,367,393]" italics="true" pageId="11" pageNumber="497">Ventral idiosoma</emphasis>
(
<figureCitation id="C2C87DB7FFAB983500C7AD3272CAFE58" box="[407,495,367,393]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB983500ADAD327156FE58" box="[509,627,367,393]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34, 40</figureCitation>
). Tritosternum with paired pilose laciniae (7780), fused basally (68), columnar base 2729 × 1517 wide; presternal area poorly sclerotised. Sternal shield (length 9095) narrowest between coxae II (9497), widest (143147), anterior margin of shield hardly conspicuous, posterior margin distinctly concave, anterolateral corners narrowly slightly extending between coxae III, gland pores
<emphasis id="6887BD20FFAB983505AEAD867603FE24" box="[1278,1318,475,501]" italics="true" pageId="11" pageNumber="497">gvb</emphasis>
indistinct; endopodal elements between coxae II and III fused with shield and endopodal elements III/IV relatively small, subtriangular, posteriorly not extended. Sternal shield with three pairs of subequal smooth setae (
<emphasis id="6887BD20FFAB9835059DAE7E7620FDEC" box="[1229,1285,547,573]" italics="true" pageId="11" pageNumber="497">st1-3</emphasis>
) (4246) and two pairs of large slit-shaped pore-like structures (
<emphasis id="6887BD20FFAB98350395AE1A71C2FDB0" box="[709,743,583,609]" italics="true" pageId="11" pageNumber="497">iv1</emphasis>
adjacent to setae
<emphasis id="6887BD20FFAB983502FAAE1A70EEFDB0" box="[938,971,583,609]" italics="true" pageId="11" pageNumber="497">st1</emphasis>
;
<emphasis id="6887BD20FFAB98350288AE1A70DEFDB0" box="[984,1019,583,609]" italics="true" pageId="11" pageNumber="497">iv2</emphasis>
between
<emphasis id="6887BD20FFAB98350534AE1A77A1FDB0" box="[1124,1156,583,609]" italics="true" pageId="11" pageNumber="497">st2</emphasis>
and
<emphasis id="6887BD20FFAB983505E9AE1A77FFFDB0" box="[1209,1242,583,609]" italics="true" pageId="11" pageNumber="497">st3</emphasis>
), setae extending slightly beyond base of following setae; shield with lineate-reticulate ornamentation on lateral surface, longitudinally median part smooth (or faintly reticulated) (
<figureCitation id="C2C87DB7FFAB983503C2AED271CEFD78" box="[658,747,655,681]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB983503AAAED2703CFD78" box="[762,793,655,681]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31</figureCitation>
). Metasternal setae
<emphasis id="6887BD20FFAB983502AFAED27705FD78" box="[1023,1056,655,681]" italics="true" pageId="11" pageNumber="497">st4</emphasis>
(4244) and metasternal poroids located on soft integument; metasternal platelets absent. Genital shield flask-shaped, slightly expanded laterally past level of setae
<emphasis id="6887BD20FFAB98350062AE8A7276FD20" box="[306,339,727,753]" italics="true" pageId="11" pageNumber="497">st5</emphasis>
, length 208212, maximum width 108111, posterior margin rounded, anterior hyaline margin of shield irregularly convex, overlapping posterior area of sternal shield (extending beyond base of setae
<emphasis id="6887BD20FFAB98350443AEA67611FCC4" box="[1299,1332,763,789]" italics="true" pageId="11" pageNumber="497">st3</emphasis>
), surface ornamented posteromedially by seven cells flanked by a median Λ-shaped ornamentation; bearing a pair of simple setae
<emphasis id="6887BD20FFAB98350188AF1E73DCFC8C" box="[216,249,835,861]" italics="true" pageId="11" pageNumber="497">st5</emphasis>
(3840) (
<figureCitation id="C2C87DB7FFAB9835003BAF1E72E6FC8C" box="[363,451,835,861]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350081AF1E7138FC8C" box="[465,541,835,861]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34</figureCitation>
). Shield flanked by two pairs of minute, oval paragenital platelets; paragenital poroids
<emphasis id="6887BD20FFAB983501A1AF3A7236FC50" box="[241,275,871,897]" italics="true" pageId="11" pageNumber="497">iv5</emphasis>
located on soft cuticle lateral to shield near seta
<emphasis id="6887BD20FFAB9835027BAF3A7069FC50" box="[811,844,871,897]" italics="true" pageId="11" pageNumber="497">st5</emphasis>
. Anal shield subtriangular, rounded anteriorly, length 9195, width 7578, anterior half faintly lineate-reticulate, bearing a pair of simple para-anal setae (2326), postanal seta (3032) thickened, sparsely barbed, apically bent (hook shaped) (see
<figureCitation id="C2C87DB7FFAB9835055FAFF27741FC18" box="[1039,1124,943,969]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350521AFF277C6FC18" box="[1137,1251,943,969]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34, 40</figureCitation>
) and with a pair of marginal pores (
<emphasis id="6887BD20FFAB9835003AAF8E72B7FC3C" box="[362,402,979,1005]" italics="true" pageId="11" pageNumber="497">gv3</emphasis>
); cribrum consisting of a terminal tuft with 34 irregular rows of spicules, anteriormost rows of spicules extending slightly beyond post-anal seta (
<figureCitation id="C2C87DB7FFAB983503A3AFAA706EFBC0" box="[755,843,1015,1041]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350209AFAA70F5FBC0" box="[857,976,1015,1041]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34, 40</figureCitation>
). Soft opisthogastric cuticle surrounding genital and anal shields with seven pairs of poroids, including one pair of paragenital poroids
<emphasis id="6887BD20FFAB983505FCA84677EBFBE4" box="[1196,1230,1051,1077]" italics="true" pageId="11" pageNumber="497">iv5</emphasis>
,
<emphasis id="6887BD20FFAB9835058CA84677DBFBE5" box="[1244,1278,1051,1076]" italics="true" pageId="11" pageNumber="497">ivp</emphasis>
and five pairs of
<emphasis id="6887BD20FFAB983501E5A86273F2FB89" box="[181,215,1087,1112]" italics="true" pageId="11" pageNumber="497">ivo</emphasis>
, inguinal gland pores
<emphasis id="6887BD20FFAB9835009FA86272D2FB88" box="[463,503,1087,1113]" italics="true" pageId="11" pageNumber="497">gv2</emphasis>
on soft cuticle close to parapodals and one pair of suboval metapodal plates (1316 long × 810 wide) present, soft cuticle with two minute paragenital platelets between metapodals and genital shield, bearing 16 pairs of thick setae (
<emphasis id="6887BD20FFAB983500A8A8DA7171FB70" box="[504,596,1159,1185]" italics="true" pageId="11" pageNumber="497">Jv1Jv5</emphasis>
,
<emphasis id="6887BD20FFAB98350331A8DA7199FB71" box="[609,700,1159,1185]" italics="true" pageId="11" pageNumber="497">Zv1 Zv</emphasis>
5,
<emphasis id="6887BD20FFAB98350388A8DA7001FB70" box="[728,804,1159,1185]" italics="true" pageId="11" pageNumber="497">R1R4</emphasis>
,
<emphasis id="6887BD20FFAB98350262A8DA7082FB70" box="[818,935,1159,1185]" italics="true" pageId="11" pageNumber="497">UR2UR3</emphasis>
); almost all setae uniform in length (2030), and arising on small sclerotised platelet, except
<emphasis id="6887BD20FFAB983503E8A8F671DFFB14" box="[696,762,1195,1221]" italics="true" pageId="11" pageNumber="497">Jv12</emphasis>
(3335) and
<emphasis id="6887BD20FFAB983502C4A8F6709BFB14" box="[916,958,1195,1221]" italics="true" pageId="11" pageNumber="497">Zv1</emphasis>
(3840) longer and setiform (
<figureCitation id="C2C87DB7FFAB98350444A8F67649FB14" box="[1300,1388,1195,1221]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 2526. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350429A8F67392FB38" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34</figureCitation>
). Peritrematal shield well developed laterad peritremes, bearing one gland pore and one poroid at level near coxae IIIII, post-stigmatic extension of shield relatively wide, not reaching posterior level of coxae IV, with two pairs of poroids and one pair of gland pores; anterior region of peritrematal shield fused to anterior margin of dorsal shield behind setae
<emphasis id="6887BD20FFAB98350021A96672AFFA84" box="[369,394,1339,1365]" italics="true" pageId="11" pageNumber="497">z1</emphasis>
. Peritremes long, extending to mid-level of coxa I (near
<emphasis id="6887BD20FFAB983502ADA9667704FA84" box="[1021,1057,1339,1365]" italics="true" pageId="11" pageNumber="497">id1</emphasis>
). Two exopodal platelets between coxae IIIII and IIIIV present, anterior platelet small and subtriangular, those between coxae IIIIV linear, narrow, more or less abutting to parapodal more or less abutting to parapodals.
</paragraph>
<paragraph id="5A4C6132FFAB98350197A9FA700CF8A0" blockId="11.[151,1437,151,2014]" pageId="11" pageNumber="497">
<taxonomicName id="9DF31AB1FFAB98350197A9FA7273FA10" authorityName="Krefft" authorityYear="1871" box="[199,342,1447,1473]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="genus">
<emphasis id="6887BD20FFAB98350197A9FA7273FA10" box="[199,342,1447,1473]" italics="true" pageId="11" pageNumber="497">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="C2C87DB7FFAB98350038A9FA7135FA10" box="[360,528,1447,1473]" captionStart="FIGURES 2729" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 2729. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Figures 2729</figureCitation>
,
<figureCitation id="C2C87DB7FFAB9835034EA9FB71B2FA10" box="[542,663,1446,1473]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">35, 36, 39</figureCitation>
). Anterior margin of epistome convex, irregularly denticulate, with smooth median spike, lineate posteriorly (
<figureCitation id="C2C87DB7FFAB983503D5A99671FAFA34" box="[645,735,1483,1509]" captionStart="FIGURES 2729" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 2729. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Fig. 28</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with 616 small denticles, except anteriormost row widest, slightly convex, with 2830 small denticles, groove with smooth posterior transverse line and progressively narrowing from anterior to posterior (
<figureCitation id="C2C87DB7FFAB9835046CAA4E76B2F9FC" box="[1340,1431,1555,1581]" captionStart="FIGURES 2729" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 2729. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Figs 27</figureCitation>
,
<figureCitation id="C2C87DB7FFAB983501C7AA6A7392F983" box="[151,183,1591,1618]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">36</figureCitation>
). Corniculi horn-like, internal malae longer than corniculi, comprising a pair of pilose median projections, and 1012 lateral, shorter, tentacle-like, smooth projections (
<figureCitation id="C2C87DB7FFAB9835024FAA067053F9A4" box="[799,886,1627,1653]" captionStart="FIGURES 2729" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 2729. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Fig. 27</figureCitation>
); labrum acuminate, longer than internal malae, fringed; hypostomal and capitular setae smooth,
<emphasis id="6887BD20FFAB983503E8AA2271F1F948" box="[696,724,1663,1689]" italics="true" pageId="11" pageNumber="497">h3</emphasis>
(4851)&gt;
<emphasis id="6887BD20FFAB98350202AA227048F948" box="[850,877,1663,1689]" italics="true" pageId="11" pageNumber="497">h1</emphasis>
(3639)&gt;
<emphasis id="6887BD20FFAB983502BBAA227720F949" box="[1003,1029,1663,1688]" italics="true" pageId="11" pageNumber="497">pc</emphasis>
(2830)&gt;
<emphasis id="6887BD20FFAB983505D3AA2277BAF948" box="[1155,1183,1663,1689]" italics="true" pageId="11" pageNumber="497">h2</emphasis>
(1720) (
<figureCitation id="C2C87DB7FFAB9835045DAA227641F948" box="[1293,1380,1663,1689]" captionStart="FIGURES 2729" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 2729. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Figs 27</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350420AA2276B5F94B" box="[1392,1424,1663,1690]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">36</figureCitation>
). Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth except palpfemur with seta
<emphasis id="6887BD20FFAB983501C7AA9A7396F930" box="[151,179,1735,1761]" italics="true" pageId="11" pageNumber="497">d3</emphasis>
thickened and inserted on small tubercles, seta
<emphasis id="6887BD20FFAB98350396AA9A71F9F930" box="[710,732,1735,1761]" italics="true" pageId="11" pageNumber="497">al</emphasis>
thickened; palpgenu with
<emphasis id="6887BD20FFAB98350554AA9A770DF930" box="[1028,1064,1735,1761]" italics="true" pageId="11" pageNumber="497">al1</emphasis>
stout,
<emphasis id="6887BD20FFAB98350522AA9A77B3F930" box="[1138,1174,1735,1761]" italics="true" pageId="11" pageNumber="497">al2</emphasis>
paddle-like; palp tarsal claw two-tined (
<figureCitation id="C2C87DB7FFAB9835001FAAB67286F8D4" box="[335,419,1770,1797]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">Fig. 35</figureCitation>
). Supralabral process indistinct. Fixed digit of chelicera with an offset and subapical tooth (gabelzhan), followed by four teeth, including two large proximal teeth posterior to a setaceous pilus dentilis, dorsal cheliceral seta prostrate, arthrodial membrane with a rounded flap and normal filaments; cheliceral lyrifissures distinct, movable digit of chelicera bidentate (
<figureCitation id="C2C87DB7FFAB983503C9AB0A71D5F8A0" box="[665,752,1879,1905]" captionStart="FIGURES 2729" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 2729. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Figs 29</figureCitation>
,
<figureCitation id="C2C87DB7FFAB983503ACAB0A7039F8A0" box="[764,796,1879,1905]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">39</figureCitation>
).
</paragraph>
<paragraph id="5A4C6132FFAB98350197AB267042F844" blockId="11.[151,1437,151,2014]" box="[199,871,1915,1941]" pageId="11" pageNumber="497">
<emphasis id="6887BD20FFAB98350197AB2672EAF845" box="[199,463,1915,1940]" italics="true" pageId="11" pageNumber="497">Insemination structures</emphasis>
. Not seen, apparently unsclerotised.
</paragraph>
<paragraph id="5A4C6132FFAB98320197ABC27288FD78" blockId="11.[151,1437,151,2014]" lastBlockId="12.[151,1437,151,789]" lastPageId="12" lastPageNumber="498" pageId="11" pageNumber="497">
<emphasis id="6887BD20FFAB98350197ABC273D9F869" box="[199,252,1951,1976]" italics="true" pageId="11" pageNumber="497">Legs</emphasis>
(
<figureCitation id="C2C87DB7FFAB9835005AABC272A8F868" box="[266,397,1951,1977]" captionStart="FIGURE 4144" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 4144. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="11" pageNumber="497">Figs 4144</figureCitation>
). Legs II and III short (352356, 343348), I and IV longer (465470) (excluding pre-tarsus). Chaetotaxy normal for free-living
<taxonomicName id="9DF31AB1FFAB9835034DAB9E71BDF80C" box="[541,664,1987,2013]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="497" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
(
<emphasis id="6887BD20FFAB983503FAAB9E71CDF80D" box="[682,744,1987,2012]" italics="true" pageId="11" pageNumber="497">sensu</emphasis>
<bibRefCitation id="3E621CC3FFAB983503A0AB9E70A8F80F" author="Evans, G. O." box="[752,909,1987,2014]" pageId="11" pageNumber="497" pagination="275 - 303" refId="ref14188" refString="Evans, G. O. (1963 a) Observations on the chaetotaxy of the legs in the free-living Gamasina (Acari: Mesostigmata). Bulletin of the British Museum (Natural History) Zoology, 10 (5), 275 - 303. https: // doi. org / 10.5962 / bhl. part. 20528" type="journal article" year="1963">Evans, 1963a</bibRefCitation>
): Leg I (
<figureCitation id="C2C87DB7FFAB983502BFAB9E7761F80C" box="[1007,1092,1987,2013]" captionStart="FIGURE 4144" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 4144. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="11" pageNumber="497">Fig. 41</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-1 (
<emphasis id="6887BD20FFAC98320079ACCA721AFF60" box="[297,319,151,177]" italics="true" pageId="12" pageNumber="498">al</emphasis>
and
<emphasis id="6887BD20FFAC98320022ACCA72ABFF60" box="[370,398,151,177]" italics="true" pageId="12" pageNumber="498">ad</emphasis>
thickened and inserted on small tubercles), femur 2-3/1, 2/3-2 (all setae on segment thickened except
<emphasis id="6887BD20FFAC983201B9ACE67229FF04" box="[233,268,187,213]" italics="true" pageId="12" pageNumber="498">al2</emphasis>
,
<emphasis id="6887BD20FFAC9832004CACE67213FF05" box="[284,310,187,212]" italics="true" pageId="12" pageNumber="498">av</emphasis>
,
<emphasis id="6887BD20FFAC98320016ACE6724FFF04" box="[326,362,187,213]" italics="true" pageId="12" pageNumber="498">pl1</emphasis>
,
<emphasis id="6887BD20FFAC98320029ACE672B8FF04" box="[377,413,187,213]" italics="true" pageId="12" pageNumber="498">pl2</emphasis>
,
<emphasis id="6887BD20FFAC983200FCACE672F2FF04" box="[428,471,187,213]" italics="true" pageId="12" pageNumber="498">ad2</emphasis>
and
<emphasis id="6887BD20FFAC98320340ACE67111FF04" box="[528,564,187,213]" italics="true" pageId="12" pageNumber="498">al1</emphasis>
,
<emphasis id="6887BD20FFAC98320313ACE67148FF04" box="[579,621,187,213]" italics="true" pageId="12" pageNumber="498">ad1</emphasis>
,
<emphasis id="6887BD20FFAC9832032DACE67182FF04" box="[637,679,187,213]" italics="true" pageId="12" pageNumber="498">ad3</emphasis>
inserted on small tubercles, see
<figureCitation id="C2C87DB7FFAC9832054BACE677E7FF04" box="[1051,1218,187,213]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="12" pageNumber="498">Figures 37, 38</figureCitation>
), genu 2-3/2, 3/1-2 (all setae on segment thickened and dorsals inserted on small tubercles, see
<figureCitation id="C2C87DB7FFAC9832055AAC8277E7FF28" box="[1034,1218,223,249]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="12" pageNumber="498">Figures 37&amp; 38</figureCitation>
), tibia 2-3/2, 3/1-2 (all setae on segment thickened and dorsals inserted on small tubercles, see
<figureCitation id="C2C87DB7FFAC983202BEAD5E77B1FECC" box="[1006,1172,259,285]" captionStart="FIGURES 3040" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 3040. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="12" pageNumber="498">Figures 37, 38</figureCitation>
). Leg II (
<figureCitation id="C2C87DB7FFAC983205AEAD5E7671FECC" box="[1278,1364,259,285]" captionStart="FIGURE 4144" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 4144. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="12" pageNumber="498">Fig. 42</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/1, 0/2-1 (
<emphasis id="6887BD20FFAC9832036BAD7A7174FE90" box="[571,593,295,321]" italics="true" pageId="12" pageNumber="498">al</emphasis>
thickened), femur 2-3/1, 2/2-1 (all setae on segment thickened except
<emphasis id="6887BD20FFAC98320421AD7A76B0FE90" box="[1393,1429,295,321]" italics="true" pageId="12" pageNumber="498">al2</emphasis>
, and
<emphasis id="6887BD20FFAC98320197AD1673D4FEB4" box="[199,241,331,357]" italics="true" pageId="12" pageNumber="498">ad1</emphasis>
,
<emphasis id="6887BD20FFAC98320050AD16720EFEB4" box="[256,299,331,357]" italics="true" pageId="12" pageNumber="498">pd1</emphasis>
,
<emphasis id="6887BD20FFAC98320069AD167246FEB4" box="[313,355,331,357]" italics="true" pageId="12" pageNumber="498">pd2</emphasis>
inserted on small tubercles,
<emphasis id="6887BD20FFAC983203F8AD1671F7FEB4" box="[680,722,331,357]" italics="true" pageId="12" pageNumber="498">pd2</emphasis>
longest), genu 2-3/1, 2/1-2 (dorsals and ventrals thickened and dorsals inserted on small tubercles), tibia 2-2/1, 2/1-2 (all setae on segment thickened except anterolaterals). Leg III (
<figureCitation id="C2C87DB7FFAC98320192ADCE7232FE7C" box="[194,279,403,429]" captionStart="FIGURE 4144" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 4144. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="12" pageNumber="498">Fig. 43</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-0 (
<emphasis id="6887BD20FFAC98320255ADCE703EFE7C" box="[773,795,403,429]" italics="true" pageId="12" pageNumber="498">al</emphasis>
and
<emphasis id="6887BD20FFAC98320201ADCE7048FE7C" box="[849,877,403,429]" italics="true" pageId="12" pageNumber="498">ad</emphasis>
thickened and inserted on small tubercles), femur 1-2/1, 1/0-1 (dorsals and anterolateral thickened,
<emphasis id="6887BD20FFAC9832039DADEA71D2FE00" box="[717,759,439,465]" italics="true" pageId="12" pageNumber="498">ad1</emphasis>
and
<emphasis id="6887BD20FFAC98320263ADEA7078FE00" box="[819,861,439,465]" italics="true" pageId="12" pageNumber="498">ad2</emphasis>
inserted on small tubercle), genu 2-2/1, 2/1-1 (all setae on segment thickened), tibia: 2-1/1, 2/1-1 (all setae on segment thickened). Leg IV (
<figureCitation id="C2C87DB7FFAC983205FDAD867620FE24" box="[1197,1285,475,501]" captionStart="FIGURE 4144" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 4144. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="12" pageNumber="498">Fig. 44</figureCitation>
): coxa 0-0/1, 0/0-0, trochanter 1-1/1, 0/2-0 (
<emphasis id="6887BD20FFAC983200A3ADA2712CFDC8" box="[499,521,511,537]" italics="true" pageId="12" pageNumber="498">al</emphasis>
and
<emphasis id="6887BD20FFAC98320312ADA2717BFDC8" box="[578,606,511,537]" italics="true" pageId="12" pageNumber="498">ad</emphasis>
thickened and inserted on small tubercles), femur 1-2/1, 1/0-1 (
<emphasis id="6887BD20FFAC98320469ADA27646FDC8" box="[1337,1379,511,537]" italics="true" pageId="12" pageNumber="498">ad1</emphasis>
,
<emphasis id="6887BD20FFAC98320422ADA276B9FDC8" box="[1394,1436,511,537]" italics="true" pageId="12" pageNumber="498">ad2</emphasis>
thickened and inserted on small tubercles), genu 2-2/1, 3/0-1 (all setae on segment slightly thickened), tibia 2-1/1, 3/1-2 (all setae on segment slightly thickened). Tarsi IIIV with 18 setae (3- 3/2, 3/2-3 +
<emphasis id="6887BD20FFAC983205D4AE1A7780FDB1" box="[1156,1189,583,608]" italics="true" pageId="12" pageNumber="498">mv</emphasis>
,
<emphasis id="6887BD20FFAC983205E4AE1A77F3FDB0" box="[1204,1238,583,609]" italics="true" pageId="12" pageNumber="498">md</emphasis>
); almost all setae on tarsus IIIV thickened, see
<figureCitation id="C2C87DB7FFAC983200A3AE3671BDFD54" box="[499,664,619,645]" captionStart="FIGURE 4144" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 4144. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="12" pageNumber="498">Figures 4244</figureCitation>
. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="5A4C6132FFAC98320197AEEE72ADFD1C" blockId="12.[151,1437,151,789]" box="[199,392,691,717]" pageId="12" pageNumber="498">
<emphasis id="6887BD20FFAC98320197AEEE722EFD1C" bold="true" box="[199,267,691,717]" pageId="12" pageNumber="498">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFAC98320197AE8A72D3FCC4" pageId="12" pageNumber="498" type="etymology">
<paragraph id="5A4C6132FFAC98320197AE8A72D3FCC4" blockId="12.[151,1437,151,789]" pageId="12" pageNumber="498">
<emphasis id="6887BD20FFAC98320197AE8A726BFD20" bold="true" box="[199,334,727,753]" pageId="12" pageNumber="498">Etymology.</emphasis>
The specific name
<taxonomicName id="9DF31AB1FFAC9832037CAE8A71A6FD20" authorityName="Joharchi" authorityYear="2022" box="[556,643,727,753]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="498" phylum="Arthropoda" rank="species" species="latisetis">
<emphasis id="6887BD20FFAC9832037CAE8A71A6FD20" box="[556,643,727,753]" italics="true" pageId="12" pageNumber="498">latisetis</emphasis>
</taxonomicName>
, from the Latin &quot;
<emphasis id="6887BD20FFAC98320219AE8A704BFD20" box="[841,878,727,753]" italics="true" pageId="12" pageNumber="498">lati</emphasis>
-&quot; meaning broad, thick and &quot;
<emphasis id="6887BD20FFAC983205EDAE8A77CAFD21" box="[1213,1263,727,752]" italics="true" pageId="12" pageNumber="498">setis</emphasis>
&quot; meaning hair, refers to the broad dorsal setae.
</paragraph>
</subSubSection>
</treatment>
</document>