treatments-xml/data/A1/0D/48/A10D4838C35C375225A0FB931F9EFBB8.xml
2024-06-21 12:46:06 +02:00

406 lines
44 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="660A96FE4925F1F42B9482E0C098843B" ID-CLB-Dataset="55793" ID-DOI="10.1080/00222933.2013.791936" ID-GBIF-Dataset="6591e69b-631a-4c4d-bf83-db8c70d1ad05" ID-ISSN="1464-5262" ID-Zenodo-Dep="5198102" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628884845165" checkinUser="felipe" docAuthor="Zhou, Yu-Lingzi &amp; Zhou, Hong-Zhang" docDate="2013" docId="A10D4838C35C375225A0FB931F9EFBB8" docLanguage="en" docName="JNatHist.47.45-46.2869-2904.pdf" docOrigin="Journal of Natural History 47 (45 - 46)" docSource="http://dx.doi.org/10.1080/00222933.2013.791936" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Liotesba Scheerpeltz 1965" docType="treatment" docVersion="9" lastPageNumber="2877" masterDocId="5D343040C35B37582413FFC11B0DFF84" masterDocTitle="Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species" masterLastPageNumber="2904" masterPageNumber="2869" pageNumber="2874" updateTime="1699079816598" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6C5A2BCE1AFAB0D9A15C8BA1C3DA81F5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E91BCA5CAF3F4A2FA203B663B5BBA845">
<mods:title id="36D88FE9B0D541AB44B30C45E6FF47A6">Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species</mods:title>
</mods:titleInfo>
<mods:name id="2306609403883DD5D1BD5AE2B443E809" type="personal">
<mods:role id="680C445845CF3850026147CE3D014983">
<mods:roleTerm id="86FD1CEB121C264841D39AD1458D552F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A0F6D4CE07CDE82C462C614373B46C6A">Zhou, Yu-Lingzi</mods:namePart>
</mods:name>
<mods:name id="27812E00CD1F5FFBC33EB72B930A58F2" type="personal">
<mods:role id="1A7CE5244CB965B18C5C3508730A3C83">
<mods:roleTerm id="1145F40E6AABF1B835983B5C3E9842F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7C4D5D094DFC3D5C28129764E18F8600">Zhou, Hong-Zhang</mods:namePart>
</mods:name>
<mods:typeOfResource id="63420E48251BC9A3284746982E5F8624">text</mods:typeOfResource>
<mods:relatedItem id="636D1B51FCD43509808B70DFFAF1D85E" type="host">
<mods:titleInfo id="D903C05CDAA3890A5D1A0B8B594A2C9C">
<mods:title id="ED7B61EDCC7630756A5CF98C65A89D2D">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="BC260121094217EB715FE775E1164B8E">
<mods:date id="DDA86B858F04921381999433F4AAAB56">2013</mods:date>
<mods:detail id="7208C35DE4017951326AB1A406EF8029" type="pubDate">
<mods:number id="3067DBB7C261B123A9FF0A5F82317FEA">2013-08-08</mods:number>
</mods:detail>
<mods:detail id="1AB149B87E8C9961628E245AC7A3BE92" type="volume">
<mods:number id="32C7FAF6E9CA3E051A012A52509D8575">47</mods:number>
</mods:detail>
<mods:detail id="F5ED96CF4C1C0E3AF38F800D2C6DFBCD" type="issue">
<mods:number id="728EEF13D150ADDA95D807DE9E6712D1">45 - 46</mods:number>
</mods:detail>
<mods:extent id="83974A6E6EB80494FB84CA119D4E7BBC" unit="page">
<mods:start id="1BFC6166D81E0664BF7B611B5A230A81">2869</mods:start>
<mods:end id="94217CE793C5D11EE92F47AAC1FFCC17">2904</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="9A4A439C9E99A327A095E80C2399438F">
<mods:url id="BE44699A5AECDFA9BCC71676BE73B56E">http://dx.doi.org/10.1080/00222933.2013.791936</mods:url>
</mods:location>
<mods:classification id="8FDBA428B54DCCD7D367721E06F824EE">journal article</mods:classification>
<mods:identifier id="EEEFF992845BF9114A3A0345D7EFAB2A" type="CLB-Dataset">55793</mods:identifier>
<mods:identifier id="7B1FC35D29B43BFFEFA24E919F141C5B" type="DOI">10.1080/00222933.2013.791936</mods:identifier>
<mods:identifier id="08667F51634C18EBC9DC0056CFB88397" type="GBIF-Dataset">6591e69b-631a-4c4d-bf83-db8c70d1ad05</mods:identifier>
<mods:identifier id="9D2ADA220DCFDFF7203444C18BBDD9CD" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="C58D34B951EC4BF93C92DBED805080EC" type="Zenodo-Dep">5198102</mods:identifier>
</mods:mods>
<treatment id="A10D4838C35C375225A0FB931F9EFBB8" ID-DOI="http://doi.org/10.5281/zenodo.6505096" ID-GBIF-Taxon="194873649" ID-Zenodo-Dep="6505096" LSID="urn:lsid:plazi:treatment:A10D4838C35C375225A0FB931F9EFBB8" httpUri="http://treatment.plazi.org/id/A10D4838C35C375225A0FB931F9EFBB8" lastPageId="10" lastPageNumber="2877" pageId="7" pageNumber="2874">
<subSubSection id="61BEAAA5C35C375F25A0FB931826FBEE" box="[435,811,1106,1130]" pageId="7" pageNumber="2874" type="nomenclature">
<paragraph id="291BF92EC35C375F25A0FB931826FBEE" blockId="7.[435,811,1106,1130]" box="[435,811,1106,1130]" pageId="7" pageNumber="2874">
<heading id="72534E42C35C375F25A0FB931826FBEE" box="[435,811,1106,1130]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="2874" reason="2">
Genus
<taxonomicName id="EEA482ADC35C375F2611FB931826FBEE" ID-CoL="8VYYF" authority="Scheerpeltz, 1965" authorityName="Scheerpeltz" authorityYear="1965" box="[514,811,1106,1130]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2874" phylum="Arthropoda" rank="genus">
<emphasis id="1BD0253CC35C375F2611FB93196CFBEE" bold="true" box="[514,609,1106,1130]" italics="true" pageId="7" pageNumber="2874">Liotesba</emphasis>
<bibRefCitation id="4D3584DFC35C375F267BFB931826FBEE" author="Scheerpeltz O" box="[616,811,1106,1130]" pageId="7" pageNumber="2874" pagination="93 - 371" refId="ref17958" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except. Megalopsidiinae et Steninae). (5. Beitrag zur Kenntnis der orientalischen Staphyliniden. Arkiv Zool Uppsala. 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C35C375F249CFB531A8AFA8E" pageId="7" pageNumber="2874" type="reference_group">
<paragraph id="291BF92EC35C375F249CFB531A8AFA8E" blockId="7.[143,1103,1170,1290]" pageId="7" pageNumber="2874">
<treatmentCitationGroup id="09B4DE00C35C375F249CFB531A8AFA8E" pageId="7" pageNumber="2874">
<treatmentCitation id="A805DF3FC35C375F249CFB531AFBFB2E" author="Scheerpeltz O" box="[143,502,1170,1194]" page="199" pageId="7" pageNumber="2874" year="1965">
<taxonomicName id="EEA482ADC35C375F249CFB531AFBFB2E" ID-CoL="8VYYF" authority="Scheerpeltz, 1965: 199" authorityName="Scheerpeltz" authorityPageNumber="199" authorityYear="1965" box="[143,502,1170,1194]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2874" phylum="Arthropoda" rank="genus">
<emphasis id="1BD0253CC35C375F249CFB531BE3FB2E" box="[143,238,1170,1194]" italics="true" pageId="7" pageNumber="2874">Liotesba</emphasis>
<bibRefCitation id="4D3584DFC35C375F24EBFB531AFBFB2E" author="Scheerpeltz O" box="[248,502,1170,1194]" pageId="7" pageNumber="2874" pagination="93 - 371" refId="ref17958" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except. Megalopsidiinae et Steninae). (5. Beitrag zur Kenntnis der orientalischen Staphyliniden. Arkiv Zool Uppsala. 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965: 199</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(species included:
<taxonomicName id="EEA482ADC35C375F26C3FB521841FB2E" authorityName="Scheerpeltz" authorityYear="1965" box="[720,844,1170,1194]" class="Insecta" family="Staphylinidae" genus="Indoscytalinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2874" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="1BD0253CC35C375F26C3FB521841FB2E" box="[720,844,1170,1194]" italics="true" pageId="7" pageNumber="2874">L. malaisei</emphasis>
</taxonomicName>
);
<treatmentCitation id="A805DF3FC35C375F2775FB521F42FB2E" author="Herman LH" box="[870,1103,1170,1194]" page="3692" pageId="7" pageNumber="2874" year="2001">
<bibRefCitation id="4D3584DFC35C375F2775FB521F42FB2E" author="Herman LH" box="[870,1103,1170,1194]" pageId="7" pageNumber="2874" pagination="3021 - 3839" refId="ref17732" refString="Herman LH. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3) Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini. Staphylinidae incertae sedis. Fossils, Protactinae. Bull Am Museum Nat History. 265: 3021 - 3839." type="journal article" year="2001">Herman, 2001: 3692</bibRefCitation>
</treatmentCitation>
(catalogue; species of the world);
<treatmentCitation id="A805DF3FC35C375F25EEFB7319DDFB4E" author="Bordoni A" box="[509,720,1202,1226]" page="315" pageId="7" pageNumber="2874" year="2002">
<bibRefCitation id="4D3584DFC35C375F25EEFB7319DDFB4E" author="Bordoni A" box="[509,720,1202,1226]" pageId="7" pageNumber="2874" pagination="1 - 998" refId="ref17034" refString="Bordoni A. 2002. Xantholinini of the Oriental Region (Coleoptera: Staphylinidae). Classification, phylogeny and taxonomic revision. Museo Regionale di Scienze Naturali Monografie (Turin). 33: 1 - 998." type="journal article" year="2002">Bordoni, 2002: 315</bibRefCitation>
</treatmentCitation>
(revision of Oriental species; characters; key to species);
<treatmentCitation id="A805DF3FC35C375F2585FB131978FB6E" author="Smetana A" box="[406,629,1234,1258]" page="691" pageId="7" pageNumber="2874" year="2004">
<bibRefCitation id="4D3584DFC35C375F2585FB131978FB6E" author="Smetana A" box="[406,629,1234,1258]" pageId="7" pageNumber="2874" pagination="687 - 698" refId="ref18057" refString="Smetana A. 2004. Staphylinidae, tribe Xantholinini. In: Lobl I, Smetana A, editors. Catalogue of Palaearctic Coleoptera, Vol. 2. Stenstrup: Apollo Books; p. 687 - 698." type="book chapter" year="2004">Smetana, 2004: 691</bibRefCitation>
</treatmentCitation>
(catalogue; Palaearctic species);
<treatmentCitation id="A805DF3FC35C375F27F8FB131BF1FA8E" author="Bordoni A" page="71" pageId="7" pageNumber="2874" year="2007">
<bibRefCitation id="4D3584DFC35C375F27F8FB131BF1FA8E" author="Bordoni A" pageId="7" pageNumber="2874" pagination="67 - 78" refId="ref17403" refString="Bordoni A. 2007 b. Xantholinini (Coleoptera staphylinidae). Supplement to the catalogue Palaearctic coleoptera, volume 2. Bollettino della Societa Entomologica Italiana. 139: 67 - 78." type="journal article" year="2007">Bordoni, 2007b: 71</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C35C375F249CFAF01A34FAEE" pageId="7" pageNumber="2874" type="type_taxon">
<paragraph id="291BF92EC35C375F249CFAF01A34FAEE" blockId="7.[143,1104,1329,1386]" pageId="7" pageNumber="2874">
<emphasis id="1BD0253CC35C375F249CFAF01A2EFACD" bold="true" box="[143,291,1329,1353]" pageId="7" pageNumber="2874">
<typeStatus id="F61F478CC35C375F249CFAF01BCBFACD" box="[143,198,1329,1353]" pageId="7" pageNumber="2874">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="EEA482ADC35C375F253FFAF319F6FACD" ID-CoL="3V87M" authority="Scheerpeltz, 1965: 201" authorityName="Scheerpeltz" authorityPageNumber="201" authorityYear="1965" box="[300,763,1329,1354]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2874" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="1BD0253CC35C375F253FFAF31AE2FACE" box="[300,495,1330,1354]" italics="true" pageId="7" pageNumber="2874">Liotesba malaisei</emphasis>
<bibRefCitation id="4D3584DFC35C375F25EAFAF019F6FACD" author="Scheerpeltz O" box="[505,763,1329,1353]" pageId="7" pageNumber="2874" pagination="93 - 371" refId="ref17958" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except. Megalopsidiinae et Steninae). (5. Beitrag zur Kenntnis der orientalischen Staphyliniden. Arkiv Zool Uppsala. 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965: 201</bibRefCitation>
</taxonomicName>
, fixed by original designation and monotypy.
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C35C3750249CFA5D1F38FD59" lastPageId="8" lastPageNumber="2875" pageId="7" pageNumber="2874" type="diagnosis">
<paragraph id="291BF92EC35C375F249CFA5D1BF6FA37" blockId="7.[143,1103,1436,1691]" box="[143,251,1436,1459]" pageId="7" pageNumber="2874">
<heading id="72534E42C35C375F249CFA5D1BF6FA37" box="[143,251,1436,1459]" fontSize="10" level="3" pageId="7" pageNumber="2874" reason="5">
<emphasis id="1BD0253CC35C375F249CFA5D1BF6FA37" box="[143,251,1436,1459]" italics="true" pageId="7" pageNumber="2874">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="291BF92EC35C3750249CFA021A66FE5A" blockId="7.[143,1103,1436,1691]" lastBlockId="8.[211,1172,167,733]" lastPageId="8" lastPageNumber="2875" pageId="7" pageNumber="2874">
<taxonomicName id="EEA482ADC35C375F249CFA021BE3FA5F" box="[143,238,1475,1499]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2874" phylum="Arthropoda" rank="genus">
<emphasis id="1BD0253CC35C375F249CFA021BE3FA5F" box="[143,238,1475,1499]" italics="true" pageId="7" pageNumber="2874">Liotesba</emphasis>
</taxonomicName>
can be distinguished from all other genera within the tribe
<taxonomicName id="EEA482ADC35C375F27ADFA031F42FA5E" authorityName="Erichson" authorityYear="1839" box="[958,1103,1474,1498]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2874" phylum="Arthropoda" rank="tribe" tribe="Xantholinini">Xantholinini</taxonomicName>
by the following characters: (a) body subcylindrical, usually medium to large in size (
<quantity id="EE5C54CBC35C375F2484F9C31A0BF99E" box="[151,262,1538,1562]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="7" pageNumber="2874" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">712 mm</quantity>
), more rarely larger (
<quantity id="EE5C54CBC35C375F25E5F9C3197FF99E" box="[502,626,1538,1562]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="7" pageNumber="2874" unit="mm" value="14.5" valueMax="17.0" valueMin="12.0">1217 mm</quantity>
); (b) head subquadrate or subrectangular, with sparse-scattered large punctures, almost without microsculpture; (c) frontal furrow often short or not obvious, ocular furrows usually missing; (d) antennomere II shorter than III; (e) labrum with four teeth; (f) second segment of maxillary palpi and last segment of labial palpi usually longest; (g) gular sutures not fused together, separated by the deeply excavated gular plate; (h) neck wide, about half of head width; (i) pronotum with two large anterolateral punctures, and one posterolateral puncture (
<figureCitation id="B19FE5ABC353375024CFFF261A5FFF7B" box="[220,338,231,255]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="8" pageNumber="2875">Figure 1A</figureCitation>
:e, f) on each side; (j) antesternal plate with a longitudinal medium furrow; (k) superior line of hypomeron bending toward prosternum before anterior angle of pronotum, but not joining inferior line; (l) mesoventrite with distinctly projecting, longitudinal median ridge; (m) protarsi dilated; tibiae with apical ctenidium, without subapical ctenidia; (n) aedeagus subelliptical or subovoid at base, with a pair of black sclerites near apex of median lobe; (o) female genital segments comprising a distinct oblong sternite, and a pair of subtriangular sclerites in ventral side (cf.
<bibRefCitation id="4D3584DFC353375027FFFE671F82FE3A" author="Bordoni A" box="[1004,1167,422,446]" pageId="8" pageNumber="2875" pagination="1 - 998" refId="ref17034" refString="Bordoni A. 2002. Xantholinini of the Oriental Region (Coleoptera: Staphylinidae). Classification, phylogeny and taxonomic revision. Museo Regionale di Scienze Naturali Monografie (Turin). 33: 1 - 998." type="journal article" year="2002">Bordoni, 2002</bibRefCitation>
, p. 67: fig. 69).
</paragraph>
<paragraph id="291BF92EC353375024EDFE271F38FD59" blockId="8.[211,1172,167,733]" pageId="8" pageNumber="2875">
According to our knowledge of the distributional ranges of all
<taxonomicName id="EEA482ADC353375027CEFE261F9EFE7A" box="[989,1171,486,511]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="1BD0253CC353375027CEFE261F31FE7B" box="[989,1084,487,511]" italics="true" pageId="8" pageNumber="2875">Liotesba</emphasis>
species
</taxonomicName>
(
<figureCitation id="B19FE5ABC353375024CFFDC71A34FD9A" box="[220,313,518,542]" captionStart="Figure 5" captionStartId="20.[211,278,961,983]" captionTargetBox="[212,1172,167,928]" captionTargetId="figure-5@20.[212,1172,167,928]" captionTargetPageId="20" captionText="Figure 5. Geographical distribution of species of the genus Liotesba Scheerpeltz. L. fengyangshana sp. nov. (); L. expansipalpis sp. nov. (); L. recticeps sp. nov. (); L. ovaticeps sp. nov. (); L. enthymema Bordoni (); L. hubeiana Bordoni (); L. itoi Bordoni (); L. oculifera Bordoni (); L. punctiventris (Sharp) (); L. malaisei Scheerpeltz (); L. rubescens Bordoni (); L. rubra Bordoni (); L. rufula Bordoni (); L. subsimilis (Cameron) (); L. siamensis Bordoni (); L. wittmeri Coiffait (); L. antonellae Bordoni ()." figureDoi="http://doi.org/10.5281/zenodo.5198116" httpUri="https://zenodo.org/record/5198116/files/figure.png" pageId="8" pageNumber="2875">Figure 5</figureCitation>
), the genus
<taxonomicName id="EEA482ADC353375025ADFDC71910FD9A" box="[446,541,518,542]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="genus">
<emphasis id="1BD0253CC353375025ADFDC71910FD9A" box="[446,541,518,542]" italics="true" pageId="8" pageNumber="2875">Liotesba</emphasis>
</taxonomicName>
should be an element of the Oriental region. The species are distributed along the south boundary of the Tibet Plateau toward the monsoon region of eastern Asia (from
<collectingCountry id="51B3B9BEC35337502608FD87198FFDDA" box="[539,642,582,606]" name="Thailand" pageId="8" pageNumber="2875">Thailand</collectingCountry>
across mainland
<collectingCountry id="51B3B9BEC3533750275AFD871883FDDA" box="[841,910,582,606]" name="China" pageId="8" pageNumber="2875">China</collectingCountry>
to
<collectingCountry id="51B3B9BEC353375027A6FD871F05FDDA" box="[949,1032,582,606]" name="Taiwan" pageId="8" pageNumber="2875">Taiwan</collectingCountry>
and
<collectingCountry id="51B3B9BEC35337502051FD871F85FDDA" box="[1090,1160,582,606]" name="Japan" pageId="8" pageNumber="2875">Japan</collectingCountry>
). Only
<taxonomicName id="EEA482ADC35337502500FDA61A93FDFA" baseAuthorityName="Cameron" baseAuthorityYear="1945" box="[275,414,614,638]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="1BD0253CC35337502500FDA61A93FDFA" box="[275,414,614,638]" italics="true" pageId="8" pageNumber="2875">L. subsimilis</emphasis>
</taxonomicName>
,
<taxonomicName id="EEA482ADC353375025BDFDA61956FDFA" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[430,603,615,638]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="1BD0253CC353375025BDFDA61956FDFA" box="[430,603,615,638]" italics="true" pageId="8" pageNumber="2875">L. punctiventris</emphasis>
</taxonomicName>
and
<taxonomicName id="EEA482ADC35337502685FDA6181FFDFA" authorityName="Scheerpeltz" authorityYear="1965" box="[662,786,614,638]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="1BD0253CC35337502685FDA6181FFDFA" box="[662,786,614,638]" italics="true" pageId="8" pageNumber="2875">L. malaisei</emphasis>
</taxonomicName>
are widespread. There are apparently two diversity hotspots. First, the southwest boundary of the Himalaya, where seven endemic species and two widespread species were found. Second, the Chinese mainland from where six endemic and two widespread species were recorded.
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C353375224C0FCE41F9EFBB8" lastPageId="10" lastPageNumber="2877" pageId="8" pageNumber="2875" type="key">
<paragraph id="291BF92EC353375024C0FCE41993FCB9" blockId="8.[211,670,805,829]" box="[211,670,805,829]" pageId="8" pageNumber="2875">
<heading id="72534E42C353375024C0FCE41993FCB9" box="[211,670,805,829]" fontSize="10" level="3" pageId="8" pageNumber="2875" reason="5">
<emphasis id="1BD0253CC353375024C0FCE41993FCB9" bold="true" box="[211,670,805,829]" pageId="8" pageNumber="2875">
Key to the species of
<taxonomicName id="EEA482ADC353375025AAFCE41993FCB9" ID-CoL="8VYYF" authority="Scheerpeltz" authorityName="Scheerpeltz" box="[441,670,805,829]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="genus">
<emphasis id="1BD0253CC353375025AAFCE41915FCB9" bold="true" box="[441,536,805,829]" italics="true" pageId="8" pageNumber="2875">Liotesba</emphasis>
Scheerpeltz
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<key id="5D35AE1FC35337522514FC941F9EFBB8" lastPageId="10" lastPageNumber="2877" pageId="8" pageNumber="2875">
<paragraph id="291BF92EC35337502514FC941F99FC09" blockId="8.[263,1172,853,909]" pageId="8" pageNumber="2875">
<keyLead id="92555A1BC35337502514FC941F99FC09" pageId="8" pageNumber="2875">
<keyStep id="9250E18BC35337502514FC941F99FC09" pageId="8" pageNumber="2875">
1. Body large, longer than
<quantity id="EE5C54CBC3533750262CFC941981FCE9" box="[575,652,853,877]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="2875" unit="mm" value="10.0">10 mm</quantity>
.......................................... 2 Body small, or medium in size, shorter than
<quantity id="EE5C54CBC35337502731FCB41862FC09" box="[802,879,885,909]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="2875" unit="mm" value="10.0">10 mm</quantity>
...................... 11
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC35337502514FC641F9EFC58" blockId="8.[263,1171,933,988]" pageId="8" pageNumber="2875">
<keyLead id="92555A1BC35337502514FC641F9EFC58" pageId="8" pageNumber="2875">
<keyStep id="9250E18BC35337502514FC641F9EFC58" pageId="8" pageNumber="2875">2. Elytra dark brown...................................................... 3 Elytra reddish brown.................................................... 5</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC35337502514FC351F9EFBC8" blockId="8.[263,1172,1012,1100]" pageId="8" pageNumber="2875">
<keyLead id="92555A1BC35337502514FC351F9EFBC8" pageId="8" pageNumber="2875">
<keyStep id="9250E18BC35337502514FC351F9EFBC8" pageId="8" pageNumber="2875">
3. Elytra fully covered with punctures; internal sac quite stout;
<collectingRegion id="EB6037CCC353375027C4FC351F10FB88" box="[983,1053,1012,1036]" country="China" name="Hubei" pageId="8" pageNumber="2875">Hubei</collectingRegion>
............................................................
<taxonomicName id="EEA482ADC35337502795FBD41F34FBA9" ID-CoL="3V87G" authority="Zhou &amp; Zhou, 2013" authorityName="Zhou &amp; Zhou" authorityYear="2013" box="[902,1081,1045,1069]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="expansipalpis" status="sp. nov.">
<emphasis id="1BD0253CC35337502795FBD41F34FBA9" box="[902,1081,1045,1069]" italics="true" pageId="8" pageNumber="2875">L. expansipalpis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E39847C35337502053FBD41F99FBA8" box="[1088,1172,1045,1068]" pageId="8" pageNumber="2875" rank="species">sp. nov.</taxonomicNameLabel>
Elytra furnished with three to five rows of punctures...................... 4
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC35337502514FBA51F9EFB58" blockId="8.[263,1172,1124,1244]" pageId="8" pageNumber="2875">
<keyLead id="92555A1BC35337502514FBA51F9EFB58" pageId="8" pageNumber="2875">
<keyStep id="9250E18BC35337502514FBA51F9EFB58" pageId="8" pageNumber="2875">
4. Head oval in shape; parameres of aedeagus quite long and narrow; Sichuan.......................................................
<taxonomicName id="EEA482ADC353375027A5FB441F34FB18" ID-CoL="3V87P" authority="Zhou &amp; Zhou, 2013" authorityName="Zhou &amp; Zhou" authorityYear="2013" box="[950,1081,1157,1180]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="ovaticeps" status="sp. nov.">
<emphasis id="1BD0253CC353375027A5FB441F34FB18" box="[950,1081,1157,1180]" italics="true" pageId="8" pageNumber="2875">L. ovaticeps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E39847C35337502053FB441F99FB18" box="[1088,1172,1157,1180]" pageId="8" pageNumber="2875" rank="species">sp. nov.</taxonomicNameLabel>
Head subrectangular in shape; parameres relatively short and broad;
<collectingCountry id="51B3B9BEC35337502522FB051AACFB58" box="[305,417,1220,1244]" name="Myanmar" pageId="8" pageNumber="2875">Myanmar</collectingCountry>
.........................................
<taxonomicName id="EEA482ADC35337502781FB041F9EFB58" ID-CoL="3V87F" authority="Bordoni" authorityName="Bordoni" authorityYear="2002" box="[914,1171,1220,1244]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="enthymema">
<emphasis id="1BD0253CC35337502781FB041F22FB58" box="[914,1071,1220,1244]" italics="true" pageId="8" pageNumber="2875">L. enthymema</emphasis>
Bordoni
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC35337502514FB321F9EFACF" blockId="8.[263,1171,1267,1355]" pageId="8" pageNumber="2875">
<keyLead id="92555A1BC35337502514FB321F9EFACF" pageId="8" pageNumber="2875">
<keyStep id="9250E18BC35337502514FB321F9EFACF" pageId="8" pageNumber="2875">
5. Posterior angle of head more angular, not rounded;
<collectingCountry id="51B3B9BEC35337502764FB3218C6FA8F" box="[887,971,1267,1291]" name="Bhutan" pageId="8" pageNumber="2875">Bhutan</collectingCountry>
.......................................................................
<taxonomicName id="EEA482ADC353375027AAFAD51F9EFAAF" ID-CoL="3V87Z" authority="Coiffait" authorityName="Coiffait" authorityYear="1977" box="[953,1171,1299,1323]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="1BD0253CC353375027AAFAD51F38FAAF" box="[953,1077,1300,1323]" italics="true" pageId="8" pageNumber="2875">L. wittmeri</emphasis>
Coiffait
</taxonomicName>
Posterior angle of head rounded......................................... 6
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC35337502514FAA21F9EFA1F" blockId="8.[263,1171,1379,1435]" pageId="8" pageNumber="2875">
<keyLead id="92555A1BC35337502514FAA21F9EFA1F" pageId="8" pageNumber="2875">
<keyStep id="9250E18BC35337502514FAA21F9EFA1F" pageId="8" pageNumber="2875">6. Pronotum shorter or slightly longer than head............................ 7 Pronotum distinctly longer than head.................................... 8</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC35337502514FA751F9EF9EE" blockId="8.[263,1172,1459,1643]" pageId="8" pageNumber="2875">
<keyLead id="92555A1BC35337502514FA751F9EF9EE" pageId="8" pageNumber="2875">
<keyStep id="9250E18BC35337502514FA751F9EF9EE" pageId="8" pageNumber="2875">
7. Mesoscutellum with polygonal reticulum; abdominal segments IIIVI ferruginous or dark reddish brown, markedly brighter than subsequent segments;
<collectingCountry id="51B3B9BEC35337502522FA321AACF98F" box="[305,417,1523,1547]" name="Myanmar" pageId="8" pageNumber="2875">Myanmar</collectingCountry>
...............................................
<taxonomicName id="EEA482ADC353375027C7FA351AD1F9AE" ID-CoL="3V87T" authority="Bordoni Mesoscutellum" authorityName="Bordoni Mesoscutellum" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="1BD0253CC353375027C7FA351F3DF98F" box="[980,1072,1523,1547]" italics="true" pageId="8" pageNumber="2875">L. rubra</emphasis>
Bordoni Mesoscutellum
</taxonomicName>
without microsculpture; abdominal segments unicolorous dark brown to black brown, only posterior margins of segments narrowly brighter;
<collectingCountry id="51B3B9BEC3533750258AF9921904F9EF" box="[409,521,1619,1643]" name="Myanmar" pageId="8" pageNumber="2875">Myanmar</collectingCountry>
..................................
<taxonomicName id="EEA482ADC353375027B5F9921F9EF9EE" ID-CoL="3V87S" authority="Bordoni" authorityName="Bordoni" authorityYear="2005" box="[934,1171,1618,1642]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2875" phylum="Arthropoda" rank="species" species="rubescens">
<emphasis id="1BD0253CC353375027B5F9921F22F9EE" box="[934,1071,1618,1642]" italics="true" pageId="8" pageNumber="2875">L. rubescens</emphasis>
Bordoni
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<caption id="7DDBA9A6C3523751249CFA01194BF901" ID-DOI="http://doi.org/10.5281/zenodo.5198104" ID-Zenodo-Dep="5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="9" pageNumber="2876" startId="9.[143,210,1472,1494]" targetBox="[144,1104,162,1439]" targetPageId="9">
<paragraph id="291BF92EC3523751249CFA01194BF901" blockId="9.[143,1104,1472,1669]" pageId="9" pageNumber="2876">
Figure 1.
<taxonomicName id="EEA482ADC352375124EBFA011AEBFA52" ID-CoL="3V87H" authority="Zhou &amp; Zhou, 2013" authorityName="Zhou &amp; Zhou" authorityYear="2013" box="[248,486,1472,1494]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2876" phylum="Arthropoda" rank="species" species="fengyangshana" status="sp. nov.">
<emphasis id="1BD0253CC352375124EBFA011AEBFA52" box="[248,486,1472,1494]" italics="true" pageId="9" pageNumber="2876">Liotesba fengyangshana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E39847C352375125FEFA001930FA52" box="[493,573,1473,1494]" pageId="9" pageNumber="2876" rank="species">sp. nov.</taxonomicNameLabel>
, (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm.
</paragraph>
</caption>
<paragraph id="291BF92EC35137522514FF661F9EFF7B" blockId="10.[263,1171,167,256]" pageId="10" pageNumber="2877">
<keyLead id="92555A1BC35137522514FF661F9EFF7B" pageId="10" pageNumber="2877">
<keyStep id="9250E18BC35137522514FF661F9EFF7B" pageId="10" pageNumber="2877">
8. Eye small, not more than 0.4 times the length of tempora;
<collectingRegion id="EB6037CCC351375227D3FF661F0BFF3B" box="[960,1030,167,191]" country="China" name="Hubei" pageId="10" pageNumber="2877">Hubei</collectingRegion>
.................................................................
<taxonomicName id="EEA482ADC351375227BEFF091F9EFF5B" ID-CoL="3V87J" authority="Bordoni" authorityName="Bordoni" authorityYear="2007" box="[941,1171,199,224]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="hubeiana">
<emphasis id="1BD0253CC351375227BEFF091F22FF64" box="[941,1071,200,224]" italics="true" pageId="10" pageNumber="2877">L. hubeiana</emphasis>
Bordoni
</taxonomicName>
Eye medium-sized, 0.40.5 times the length of tempora................... 9
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC35137522514FED61F9EFE0B" blockId="10.[263,1172,279,399]" pageId="10" pageNumber="2877">
<keyLead id="92555A1BC35137522514FED61F9EFE0B" pageId="10" pageNumber="2877">
<keyStep id="9250E18BC35137522514FED61F9EFE0B" pageId="10" pageNumber="2877">
9. Male tergite X angulate at lateral margin; internal sac coiled three or two times.......................................................................
<specimenCount id="3FA232A7C3513752206AFEF61A66FEEB" pageId="10" pageNumber="2877" type="male">10 Male</specimenCount>
tergite X rounded at lateral margin; internal sac coiled over four times;
<collectingCountry id="51B3B9BEC35137522522FEB61A60FE0B" box="[305,365,375,399]" name="India" pageId="10" pageNumber="2877">India</collectingCountry>
.................................................
<taxonomicName id="EEA482ADC351375227A2FEB91F9EFE0B" ID-CoL="3V87N" authority="Bordoni" authorityName="Bordoni" authorityYear="2003" box="[945,1171,375,399]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="oculifera">
<emphasis id="1BD0253CC351375227A2FEB91F3DFE0B" box="[945,1072,375,399]" italics="true" pageId="10" pageNumber="2877">L. oculifera</emphasis>
Bordoni
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC351375224E9FE671F9EFE5A" blockId="10.[250,1171,422,479]" pageId="10" pageNumber="2877">
<keyLead id="92555A1BC351375224E9FE671F9EFE5A" pageId="10" pageNumber="2877">
<keyStep id="9250E18BC351375224E9FE671F9EFE5A" pageId="10" pageNumber="2877">
10. Internal sac coiled over three times;
<collectingCountry id="51B3B9BEC351375226D5FE671814FE3A" box="[710,793,422,446]" name="Taiwan" pageId="10" pageNumber="2877">Taiwan</collectingCountry>
.................
<taxonomicName id="EEA482ADC351375227F8FE691A81FE5A" ID-CoL="3V87K" authority="Bordoni Internal" authorityName="Bordoni Internal" authorityYear="2002" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="1BD0253CC351375227F8FE691F22FE3A" box="[1003,1071,423,447]" italics="true" pageId="10" pageNumber="2877">L. itoi</emphasis>
Bordoni Internal
</taxonomicName>
sac coiled two times; Yunnan................
<taxonomicName id="EEA482ADC3513752278CFE061F9EFE5A" ID-CoL="3V87D" authority="Bordoni" authorityName="Bordoni" authorityYear="2012" box="[927,1171,454,479]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="antonellae">
<emphasis id="1BD0253CC3513752278CFE061F22FE5B" box="[927,1071,455,479]" italics="true" pageId="10" pageNumber="2877">L. antonellae</emphasis>
Bordoni
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC351375224E9FE371F99FDAA" blockId="10.[250,1172,502,558]" pageId="10" pageNumber="2877">
<keyLead id="92555A1BC351375224E9FE371F99FDAA" pageId="10" pageNumber="2877">
<keyStep id="9250E18BC351375224E9FE371F99FDAA" pageId="10" pageNumber="2877">
11. Anteocular furrow distinct;
<collectingRegion id="EB6037CCC35137522679FE3719C3FD8A" box="[618,718,502,526]" country="China" name="Zhejiang" pageId="10" pageNumber="2877">Zhejiang</collectingRegion>
.............
<taxonomicName id="EEA482ADC3513752276BFE361F34FD8A" ID-CoL="3V87H" authority="Zhou &amp; Zhou, 2013" authorityName="Zhou &amp; Zhou" authorityYear="2013" box="[888,1081,502,526]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="fengyangshana" status="sp. nov.">
<emphasis id="1BD0253CC3513752276BFE361F34FD8A" box="[888,1081,502,526]" italics="true" pageId="10" pageNumber="2877">L. fengyangshana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E39847C35137522053FE361F99FD8A" box="[1088,1172,503,526]" pageId="10" pageNumber="2877" rank="species">sp. nov.</taxonomicNameLabel>
Anteocular furrow missing............................................. 12
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC351375224E9FD871F99FDFA" blockId="10.[250,1172,582,638]" pageId="10" pageNumber="2877">
<keyLead id="92555A1BC351375224E9FD871F99FDFA" pageId="10" pageNumber="2877">
<keyStep id="9250E18BC351375224E9FD871F99FDFA" pageId="10" pageNumber="2877">
12. Head distinctly oblong; Cosmopolitan species.......
<taxonomicName id="EEA482ADC35137522781FD861F9EFDDA" ID-CoL="3V87M" authority="Scheerpeltz" authorityName="Scheerpeltz" authorityYear="1965" box="[914,1171,582,606]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="1BD0253CC35137522781FD861F06FDDA" box="[914,1035,582,606]" italics="true" pageId="10" pageNumber="2877">L. malaisei</emphasis>
Scheerpeltz
</taxonomicName>
Head subquadrate or subrectangular................................... 13
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC351375224E9FD571F99FD49" blockId="10.[250,1172,662,717]" pageId="10" pageNumber="2877">
<keyLead id="92555A1BC351375224E9FD571F99FD49" pageId="10" pageNumber="2877">
<keyStep id="9250E18BC351375224E9FD571F99FD49" pageId="10" pageNumber="2877">
13. Tempora slightly dilated; Cosmopolitan species.....
<taxonomicName id="EEA482ADC3513752279BFD561A9AFD49" ID-CoL="3V87W" authority="(Cameron) Tempora" authorityName="Tempora" baseAuthorityName="Cameron" baseAuthorityYear="1945" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="1BD0253CC3513752279BFD561F1FFD2A" box="[904,1042,662,686]" italics="true" pageId="10" pageNumber="2877">L. subsimilis</emphasis>
(Cameron) Tempora
</taxonomicName>
not dilated, usually parallel................................... 14
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC351375224E9FD241F99FC99" blockId="10.[250,1172,741,797]" pageId="10" pageNumber="2877">
<keyLead id="92555A1BC351375224E9FD241F99FC99" pageId="10" pageNumber="2877">
<keyStep id="9250E18BC351375224E9FD241F99FC99" pageId="10" pageNumber="2877">14. Frontal furrow distinct, deep and long.................................. 15 Frontal furrow indistinct............................................... 16</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC351375224E9FCF41F99FC28" blockId="10.[250,1172,821,941]" pageId="10" pageNumber="2877">
<keyLead id="92555A1BC351375224E9FCF41F99FC28" pageId="10" pageNumber="2877">
<keyStep id="9250E18BC351375224E9FCF41F99FC28" pageId="10" pageNumber="2877">
15. Distance between antennal insertions subequal to that from antenna to eye; Cosmopolitan species..............................
<taxonomicName id="EEA482ADC3513752279FFC971A99FC09" ID-CoL="3V87Q" authority="(Sharp) Distance" authorityName="Distance" baseAuthorityName="Sharp" baseAuthorityYear="1889" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="1BD0253CC3513752279FFC971F3AFCE9" box="[908,1079,854,877]" italics="true" pageId="10" pageNumber="2877">L. punctiventris</emphasis>
(Sharp) Distance
</taxonomicName>
between antennal insertions shorter than that from antenna to eye;
<collectingRegion id="EB6037CCC35137522522FC541A81FC29" box="[305,396,917,941]" country="China" name="Sichuan" pageId="10" pageNumber="2877">Sichuan</collectingRegion>
..............................................
<taxonomicName id="EEA482ADC351375227A9FC571F34FC29" ID-CoL="3V87R" authority="Zhou &amp; Zhou, 2013" authorityName="Zhou &amp; Zhou" authorityYear="2013" box="[954,1081,918,941]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="recticeps" status="sp. nov.">
<emphasis id="1BD0253CC351375227A9FC571F34FC29" box="[954,1081,918,941]" italics="true" pageId="10" pageNumber="2877">L. recticeps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E39847C35137522053FC541F99FC28" box="[1088,1172,917,940]" pageId="10" pageNumber="2877" rank="species">sp. nov.</taxonomicNameLabel>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="291BF92EC351375224E9FC051F9EFBB8" blockId="10.[250,1171,964,1084]" pageId="10" pageNumber="2877">
<keyLead id="92555A1BC351375224E9FC051F9EFBB8" pageId="10" pageNumber="2877">
<keyStep id="9250E18BC351375224E9FC051F9EFBB8" pageId="10" pageNumber="2877">
16. Lateral margins of pronotum slightly sinuate; elytra not obviously narrow;
<collectingCountry id="51B3B9BEC35137522522FC251A60FC78" box="[305,365,996,1020]" name="India" pageId="10" pageNumber="2877">India</collectingCountry>
..................................................
<taxonomicName id="EEA482ADC351375227DDFC241A8FFB98" ID-CoL="72GRF" authority="Bordoni Lateral" authorityName="Bordoni Lateral" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="rufula">
<emphasis id="1BD0253CC351375227DDFC241F22FC79" box="[974,1071,997,1021]" italics="true" pageId="10" pageNumber="2877">L. rufula</emphasis>
Bordoni Lateral
</taxonomicName>
margins of pronotum quite convergent at middle; elytra distinctly narrow;
<collectingCountry id="51B3B9BEC3513752259CFBE51AFBFBB8" box="[399,502,1060,1084]" name="Thailand" pageId="10" pageNumber="2877">Thailand</collectingCountry>
....................................
<taxonomicName id="EEA482ADC351375227BBFBE41F9EFBB8" ID-CoL="3V87V" authority="Bordoni" authorityName="Bordoni" authorityYear="2006" box="[936,1171,1060,1084]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="1BD0253CC351375227BBFBE41F22FBB8" box="[936,1071,1061,1084]" italics="true" pageId="10" pageNumber="2877">L. siamensis</emphasis>
Bordoni
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
</key>
</subSubSection>
</treatment>
</document>