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<mods:title id="87A51CFD5F87076A3D0B6A5FF4645659">Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species</mods:title>
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<mods:namePart id="2430845E42A241AFD6D69A8F8CD0FD1C">Zhou, Yu-Lingzi</mods:namePart>
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<taxonomicName id="EEA482ADC341374225FEFE861874FEDB" ID-CoL="3V87M" authority="Scheerpeltz, 1965" authorityName="Scheerpeltz" authorityYear="1965" box="[493,889,327,351]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="2893" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="1BD0253CC341374225FEFE8619A3FEDB" bold="true" box="[493,686,327,351]" italics="true" pageId="26" pageNumber="2893">Liotesba malaisei</emphasis>
<bibRefCitation id="4D3584DFC341374226A5FE861874FEDB" author="Scheerpeltz O" box="[694,889,327,351]" pageId="26" pageNumber="2893" pagination="93 - 371" refId="ref17958" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except. Megalopsidiinae et Steninae). (5. Beitrag zur Kenntnis der orientalischen Staphyliniden. Arkiv Zool Uppsala. 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965</bibRefCitation>
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<treatmentCitation id="A805DF3FC341374224C0FE461ADDFE1B" author="Scheerpeltz O" box="[211,464,391,415]" page="201" pageId="26" pageNumber="2893" year="1965">
<bibRefCitation id="4D3584DFC341374224C0FE461ADDFE1B" author="Scheerpeltz O" box="[211,464,391,415]" pageId="26" pageNumber="2893" pagination="93 - 371" refId="ref17958" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except. Megalopsidiinae et Steninae). (5. Beitrag zur Kenntnis der orientalischen Staphyliniden. Arkiv Zool Uppsala. 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965: 201</bibRefCitation>
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[
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locality: northeast
<collectingCountry id="51B3B9BEC341374226FCFE46184DFE1A" box="[751,832,391,414]" name="Myanmar" pageId="26" pageNumber="2893">Burma</collectingCountry>
, Kambaiti,
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(
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)];
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(catalogue);
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(characters;
<collectingCountry id="51B3B9BEC341374227AEFE671F2AFE3A" box="[957,1063,422,446]" name="Thailand" pageId="26" pageNumber="2893">Thailand</collectingCountry>
,
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,
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,
<collectingRegion id="EB6037CCC341374225B2FE061994FE5A" box="[417,665,454,479]" country="Thailand" name="Mae Hong Son" pageId="26" pageNumber="2893">Mae Hong Son Prov.</collectingRegion>
, Soppong);
<treatmentCitation id="A805DF3FC34137422736FE071F0BFE5A" author="Smetana A" box="[805,1030,454,478]" page="691" pageId="26" pageNumber="2893" year="2004">
<bibRefCitation id="4D3584DFC34137422736FE071F0BFE5A" author="Smetana A" box="[805,1030,454,478]" pageId="26" pageNumber="2893" pagination="687 - 698" refId="ref18057" refString="Smetana A. 2004. Staphylinidae, tribe Xantholinini. In: Lobl I, Smetana A, editors. Catalogue of Palaearctic Coleoptera, Vol. 2. Stenstrup: Apollo Books; p. 687 - 698." type="book chapter" year="2004">Smetana, 2004: 691</bibRefCitation>
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(catalogue);
<treatmentCitation id="A805DF3FC341374224C0FE271A93FE7A" author="Bordoni A" box="[211,414,486,510]" page="9" pageId="26" pageNumber="2893" year="2005">
<bibRefCitation id="4D3584DFC341374224C0FE271A93FE7A" author="Bordoni A" box="[211,414,486,510]" pageId="26" pageNumber="2893" pagination="9 - 13" refId="ref17220" refString="Bordoni A. 2005 a. Contribution to the knowledge on the Xantholini from China. IX. Description of two new species (Coleoptera, Staphylinidae). Onychium. 2: 9 - 13." type="journal article" year="2005">Bordoni, 2005a: 9</bibRefCitation>
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(characters; first record in
<collectingCountry id="51B3B9BEC341374226C1FE271817FE7A" box="[722,794,486,510]" name="China" pageId="26" pageNumber="2893">China</collectingCountry>
,
<collectingRegion id="EB6037CCC34137422736FE26188FFE7A" box="[805,898,487,510]" country="China" name="Yunnan" pageId="26" pageNumber="2893">Yunnan</collectingRegion>
,
<quantity id="EE5C54CBC3413742279DFE2718D8FE7A" box="[910,981,486,510]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="26" pageNumber="2893" unit="km" value="65.0">65 km</quantity>
east Tengchong, humid forest,
<quantity id="EE5C54CBC34137422563FDC7190AFD9A" box="[368,519,518,542]" metricMagnitude="3" metricUnit="m" metricValue="2.275" metricValueMax="2.35" metricValueMin="2.2" pageId="26" pageNumber="2893" unit="m" value="2275.0" valueMax="2350.0" valueMin="2200.0">22002350 m</quantity>
);
<bibRefCitation id="4D3584DFC34137422608FDC719FEFD9A" author="Bordoni A" box="[539,755,518,542]" pageId="26" pageNumber="2893" pagination="67 - 78" refId="ref17403" refString="Bordoni A. 2007 b. Xantholinini (Coleoptera staphylinidae). Supplement to the catalogue Palaearctic coleoptera, volume 2. Bollettino della Societa Entomologica Italiana. 139: 67 - 78." type="journal article" year="2007">Bordoni, 2007b: 71</bibRefCitation>
(catalogue).
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</paragraph>
</subSubSection>
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<emphasis id="1BD0253CC341374224C0FD871ABCFDDA" bold="true" box="[211,433,582,606]" pageId="26" pageNumber="2893">Material examined.</emphasis>
<materialsCitation id="99CCF373C341374225AEFD871F51FD39" ID-GBIF-Occurrence="3757895305" collectingDate="1934-06-04" collectingDateMax="1934-06-08" collectingDateMin="1934-06-04" collectionCode="R, NHRS" country="Myanmar" determinerName="Scheerplftz. &amp; Bordoni" elevation="2134" location="Kambaiti" pageId="26" pageNumber="2893" specimenCount="1" typeStatus="holotype">
female, with labels as follows: “
<typeStatus id="F61F478CC34137422727FD8718BCFDDA" box="[820,945,582,606]" pageId="26" pageNumber="2893" type="holotype">Holotypus</typeStatus>
”/“N. E.
<collectingCountry id="51B3B9BEC34137422031FD871F82FDDA" box="[1058,1167,582,606]" name="Myanmar" pageId="26" pageNumber="2893">BURMA</collectingCountry>
,
<location id="2C7BAFF5C341374224C0FDA71A4FFDFA" LSID="urn:lsid:plazi:treatment:A10D4838C341374325FEFE861F41FB5F:2C7BAFF5C341374224C0FDA71A4FFDFA" box="[211,322,614,638]" country="Myanmar" name="Kambaiti" pageId="26" pageNumber="2893">Kambaiti</location>
,
<quantity id="EE5C54CBC34137422541FDA71AAAFDFA" box="[338,423,614,638]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="26" pageNumber="2893" unit="ft" value="7000.0">
<elevation id="A2891E1DC34137422541FDA71AAAFDFA" box="[338,423,614,638]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="26" pageNumber="2893" unit="ft" value="7000.0">7000 ft</elevation>
</quantity>
,
<date id="5D1ADFEEC341374225A4FDA7194EFDFA" box="[439,579,614,638]" pageId="26" pageNumber="2893" value="1934-06-04" valueMax="1934-06-08" valueMin="1934-06-04">
<collectingDate id="4D5E2606C341374225A4FDA7194EFDFA" box="[439,579,614,638]" pageId="26" pageNumber="2893" value="1934-06-04" valueMax="1934-06-08" valueMin="1934-06-04">48.VI.1934</collectingDate>
</date>
,
<collectionCode id="4FB561EBC34137422641FDA61967FDFA" box="[594,618,615,638]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="2893">R</collectionCode>
. Malaise”/“
<taxonomicName id="EEA482ADC341374226E9FDA718C8FDFA" authority="Malaisei" authorityName="Malaisei" box="[762,965,614,638]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="2893" phylum="Arthropoda" rank="genus">
<emphasis id="1BD0253CC341374226E9FDA718C8FDFA" box="[762,965,614,638]" italics="true" pageId="26" pageNumber="2893">Liotesba Malaisei</emphasis>
</taxonomicName>
, det.
<determinerName id="DCC715EAC3413742201DFDA71F99FDFA" box="[1038,1172,614,638]" pageId="26" pageNumber="2893">Scheerplftz.</determinerName>
1941”/“Naturhistoriska Riksmuseet, Stockholm Loan no 261/93”/“4473 E91”/ “
<taxonomicName id="EEA482ADC341374224F2FD671AACFD3A" authorityName="Scheerpeltz" authorityYear="1965" box="[225,417,678,702]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="2893" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="1BD0253CC341374224F2FD671AACFD3A" box="[225,417,678,702]" italics="true" pageId="26" pageNumber="2893">Liotesba malaisei</emphasis>
</taxonomicName>
, det.
<determinerName id="DCC715EAC341374225F3FD64194DFD39" box="[480,576,677,701]" pageId="26" pageNumber="2893">Bordoni</determinerName>
, 1993.”/“NHRS-COLE 000008104”/. (
<collectionCode id="4FB561EBC34137422013FD641F59FD39" box="[1024,1108,677,701]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="26" pageNumber="2893">NHRS</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="61BEAAA5C341374324C0FCC21F41FB5F" lastPageId="27" lastPageNumber="2894" pageId="26" pageNumber="2893" type="description">
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<emphasis id="1BD0253CC341374224C0FCC21A5FFC9E" box="[211,338,771,794]" italics="true" pageId="26" pageNumber="2893">Description</emphasis>
</heading>
</paragraph>
<paragraph id="291BF92EC341374224C0FCEB18A3FCE6" blockId="26.[211,1172,771,993]" pageId="26" pageNumber="2893">
<emphasis id="1BD0253CC341374224C0FCEB1A7DFCC6" bold="true" box="[211,368,810,834]" pageId="26" pageNumber="2893">Measurement.</emphasis>
BL =
<quantity id="EE5C54CBC341374225DAFCEB193CFCC6" box="[457,561,810,834]" metricMagnitude="-3" metricUnit="m" metricValue="8.94" pageId="26" pageNumber="2893" unit="mm" value="8.94">8.94 mm</quantity>
, FL =
<quantity id="EE5C54CBC3413742269EFCEB19FBFCC6" box="[653,758,810,834]" metricMagnitude="-3" metricUnit="m" metricValue="5.19" pageId="26" pageNumber="2893" unit="mm" value="5.19">5.19 mm</quantity>
, HL =
<quantity id="EE5C54CBC34137422745FCEB18B3FCC6" box="[854,958,810,834]" metricMagnitude="-3" metricUnit="m" metricValue="1.47" pageId="26" pageNumber="2893" unit="mm" value="1.47">1.47 mm</quantity>
, HW =
<quantity id="EE5C54CBC34137422034FCEB1F82FCC6" box="[1063,1167,810,834]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="26" pageNumber="2893" unit="mm" value="1.14">1.14 mm</quantity>
, PL =
<quantity id="EE5C54CBC34137422504FC8B1A71FCE6" box="[279,380,842,866]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="26" pageNumber="2893" unit="mm" value="1.65">1.65 mm</quantity>
, PW =
<quantity id="EE5C54CBC341374225C0FC8B193AFCE6" box="[467,567,842,866]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="26" pageNumber="2893" unit="mm" value="1.11">1.11 mm</quantity>
, EL =
<quantity id="EE5C54CBC34137422694FC8B19E1FCE6" box="[647,748,842,866]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="2893" unit="mm" value="1.8">1.80 mm</quantity>
, EW =
<quantity id="EE5C54CBC34137422756FC8B18A4FCE6" box="[837,937,842,866]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="26" pageNumber="2893" unit="mm" value="1.29">1.29 mm</quantity>
.
</paragraph>
<paragraph id="291BF92EC341374224EDFCAB19F5FC65" blockId="26.[211,1172,771,993]" pageId="26" pageNumber="2893">Body elongate, medium-sized. Body entirely dark brown, except elytra and posterior margin of each abdominal segment reddish. Legs reddish brown, except tarsi yellowish. Antennae reddish brown, basal three antennomeres smoother than remaining antennomeres. Maxillary and labial palpi brown.</paragraph>
<paragraph id="291BF92EC341374224C0FBC81A43FA24" blockId="26.[211,1172,1033,1440]" pageId="26" pageNumber="2893">
<emphasis id="1BD0253CC341374224C0FBC81A1AFBA5" bold="true" box="[211,279,1033,1057]" pageId="26" pageNumber="2893">Head.</emphasis>
Distinctly oblong (HL to HW ratio 1.3), tempora subparallel, posterior angles rounded. Dorsal integument shiny, rather smooth, extensively bearing micropunctures, with few sparsely scattered punctures in large size at the lateral one-third and the posterior one-quarter surface of head, without any punctures on deflexed portion of tempora. Each side of cranium with anterolateral puncture near antennal insertion, midlateral puncture approximately the distance of two punctures diameter from dorsal margin of eye, temporal puncture at the posterior one-quarter, and occipital puncture at the lateral one-third. Frontal furrows and anteocular furrows absent. Eye small, subequal to one-third times the length of tempora (eye: tempora = 0.30:
<quantity id="EE5C54CBC34137422032FAC91F85FAA4" box="[1057,1160,1288,1312]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="26" pageNumber="2893" unit="mm" value="0.83">0.83 mm</quantity>
), slightly protruding laterad; two large and one small puncture next to dorsal margin of eye. Epistoma distinctly narrow, subrectangular. Distance between antennal insertions
<quantity id="EE5C54CBC341374225D1FAA91920FA04" box="[450,557,1384,1408]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="2893" unit="mm" value="0.3">0.30 mm</quantity>
being shorter than distance from antenna to eye (
<quantity id="EE5C54CBC341374224C8FA491A4FFA24" box="[219,322,1416,1440]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="26" pageNumber="2893" unit="mm" value="0.39">0.39 mm</quantity>
).
</paragraph>
<paragraph id="291BF92EC341374224C0FA061AC7F9BB" blockId="26.[211,1172,1479,1599]" pageId="26" pageNumber="2893">
<emphasis id="1BD0253CC341374224C0FA061A4FFA5B" bold="true" box="[211,322,1479,1503]" pageId="26" pageNumber="2893">Antennae.</emphasis>
Scape stout, thickened apically, slightly longer than three subsequent antennomeres combined,
<quantity id="EE5C54CBC341374225EBFA261952FA7B" box="[504,607,1511,1535]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="26" pageNumber="2893" unit="mm" value="0.53">0.53 mm</quantity>
; antennomere II elongate,
<quantity id="EE5C54CBC34137422781FA2618F5FA7B" box="[914,1016,1511,1535]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="2893" unit="mm" value="0.15">0.15 mm</quantity>
; III distinctly elongate, slightly longer than II,
<quantity id="EE5C54CBC34137422652F9C619ABF99B" box="[577,678,1543,1567]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="26" pageNumber="2893" unit="mm" value="0.18">0.18 mm</quantity>
; IV and V subequal in length,
<quantity id="EE5C54CBC341374227EFF9C61F6CF99B" box="[1020,1121,1543,1567]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="26" pageNumber="2893" unit="mm" value="0.09">0.09 mm</quantity>
; last antennomere missing.
</paragraph>
<paragraph id="291BF92EC341374324C0F9A6182DFF5B" blockId="26.[211,1171,1639,1695]" lastBlockId="27.[143,1103,167,224]" lastPageId="27" lastPageNumber="2894" pageId="26" pageNumber="2893">
<emphasis id="1BD0253CC341374224C0F9A61A50F9FB" bold="true" box="[211,349,1639,1663]" pageId="26" pageNumber="2893">Mouthparts.</emphasis>
Labrum transverse, with a pair of median teeth and a lateral tooth on each side; median teeth distinctly protruding anteriad, and lateral teeth not distinctly protruding. Mandibles falciform, right one with a sharp tooth on inner edge. Maxillary palpus elongate, last segment fusiform, apex subtruncated.
</paragraph>
<paragraph id="291BF92EC3403743249CFEC618C6FE9B" blockId="27.[143,971,263,287]" box="[143,971,263,287]" pageId="27" pageNumber="2894">
<emphasis id="1BD0253CC3403743249CFEC61BDCFE9B" bold="true" box="[143,209,263,287]" pageId="27" pageNumber="2894">Neck.</emphasis>
Narrow (
<quantity id="EE5C54CBC3403743252EFEC61AA9FE9B" box="[317,420,263,287]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="27" pageNumber="2894" unit="mm" value="0.45">0.45 mm</quantity>
), slightly wider than one-third the width of head.
</paragraph>
<paragraph id="291BF92EC3403743249CFE86196AFE7A" blockId="27.[143,1104,327,510]" pageId="27" pageNumber="2894">
<emphasis id="1BD0253CC3403743249CFE861A09FEDB" bold="true" box="[143,260,327,351]" pageId="27" pageNumber="2894">Pronotum.</emphasis>
Subrectangular (PL to PW ratio 1.4), longer than head, but same in width. Distinctly widened anteriad, lateral margins convergent backwards, anterior angles well defined, posterior angles broadly rounded. Integument extensively covered with micropunctures, with two large punctures on each anterolateral corner, one large puncture on each posterolateral corner, and also with other irregular punctures near anterior and lateral margins of pronotum.
</paragraph>
<paragraph id="291BF92EC3403743249CFDE71890FDBA" blockId="27.[143,925,550,574]" box="[143,925,550,574]" pageId="27" pageNumber="2894">
<emphasis id="1BD0253CC3403743249CFDE71A36FDBA" bold="true" box="[143,315,550,574]" pageId="27" pageNumber="2894">Mesoscutellum.</emphasis>
Shiny, with three pairs of punctures lined in two rows.
</paragraph>
<paragraph id="291BF92EC3403743249CFDA719DBFD79" blockId="27.[143,1104,614,765]" pageId="27" pageNumber="2894">
<emphasis id="1BD0253CC3403743249CFDA71BD0FDFA" bold="true" box="[143,221,614,638]" pageId="27" pageNumber="2894">Elytra.</emphasis>
Subrectangular, distinctly elongate (EL to EW ratio 1.4), longer, wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, hind margin rounded. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of dense punctures along median suture, a row along midwidth, also with three or four rows of punctures on deflexed portion.
</paragraph>
<paragraph id="291BF92EC3403743249CFCE418F6FCD9" blockId="27.[143,1104,805,862]" pageId="27" pageNumber="2894">
<emphasis id="1BD0253CC3403743249CFCE41BC1FCB9" bold="true" box="[143,204,805,829]" pageId="27" pageNumber="2894">Legs.</emphasis>
First four segments of protarsi heart-shaped, dilated, and those of mesotarsi and metatarsi slender. Last segment shorter than the IIIV combined, respectively.
</paragraph>
<paragraph id="291BF92EC3403743249CFC441877FB98" blockId="27.[143,1103,901,1052]" pageId="27" pageNumber="2894">
<emphasis id="1BD0253CC3403743249CFC441BF3FC19" bold="true" box="[143,254,901,925]" pageId="27" pageNumber="2894">Abdomen.</emphasis>
Cylindrical, broadest at segment VI. Tergites IIIVII shiny, each segment covered with distinct transverse microstriae and large scattered punctures, distance between punctures about two punctures diameter. Each tergite with a basal impression near anterior margin, rugose, with larger and denser punctures. All abdominal sternites shiny, with microstriae and punctures as those on tergites.
</paragraph>
<paragraph id="291BF92EC3403743249CFB851BD4FBF8" blockId="27.[143,1103,1092,1148]" pageId="27" pageNumber="2894">
<emphasis id="1BD0253CC3403743249CFB851A11FBD8" bold="true" box="[143,284,1092,1116]" pageId="27" pageNumber="2894">Distribution.</emphasis>
<collectingCountry id="51B3B9BEC34037432535FB851A66FBD8" box="[294,363,1092,1116]" name="China" pageId="27" pageNumber="2894">China</collectingCountry>
(
<collectingRegion id="EB6037CCC3403743256FFB851AD6FBD8" box="[380,475,1092,1116]" country="China" name="Yunnan" pageId="27" pageNumber="2894">Yunnan</collectingRegion>
);
<collectingCountry id="51B3B9BEC340374325E2FB84196CFBD9" box="[497,609,1093,1117]" name="Myanmar" pageId="27" pageNumber="2894">Myanmar</collectingCountry>
(
<collectingRegion id="EB6037CCC34037432660FB851800FBD8" box="[627,781,1092,1116]" country="Myanmar" name="Kachin" pageId="27" pageNumber="2894">Kachin State</collectingRegion>
);
<collectingCountry id="51B3B9BEC34037432730FB851887FBD8" box="[803,906,1092,1116]" name="Thailand" pageId="27" pageNumber="2894">Thailand</collectingCountry>
(
<collectingRegion id="EB6037CCC34037432788FB851BC3FBF8" country="Thailand" name="Mae Hong Son" pageId="27" pageNumber="2894">Mae Hong Son Prov.</collectingRegion>
).
</paragraph>
<paragraph id="291BF92EC3403743249CFB651F41FB5F" blockId="27.[143,1103,1188,1244]" pageId="27" pageNumber="2894">
<emphasis id="1BD0253CC3403743249CFB651BF5FB38" bold="true" box="[143,248,1188,1212]" pageId="27" pageNumber="2894">Remarks.</emphasis>
This species can be easily distinguished from its congeners by the very slender body shape, small eyes and large but few punctures on dorsal surface of head.
</paragraph>
</subSubSection>
</treatment>
</document>