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<document id="133EB8671CDE463604C4F1AD4A08E57B" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="9978976365748558D34127C9FBBA9EE1" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lapidaster Thuy 2013" docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="34" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5AA155298F5AED4FBD3BDA40D9C6BA9A">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:namePart id="6CB0C2251D02F6A778CB2579C0C9C644">Thuy, Ben</mods:namePart>
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<mods:title id="DEA2C8064EE49F809A5B08D4F28D91B5">European Journal of Taxonomy</mods:title>
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<mods:date id="E7551C6E7903B09022FB72C82F75AD01">2013</mods:date>
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<mods:number id="6342D3425042998F9D767328D8E1C48E">2013-07-02</mods:number>
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<treatment id="9978976365748558D34127C9FBBA9EE1" ID-DOI="http://doi.org/10.5281/zenodo.3844279" ID-GBIF-Taxon="164044814" ID-Zenodo-Dep="3844279" LSID="urn:lsid:plazi:treatment:9978976365748558D34127C9FBBA9EE1" httpUri="http://treatment.plazi.org/id/9978976365748558D34127C9FBBA9EE1" lastPageId="35" lastPageNumber="34" pageId="33" pageNumber="34">
<subSubSection id="59CB75FE6574855AD34127C9FC3B9D82" box="[625,963,736,763]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="116E26756574855AD34127C9FC3B9D82" blockId="33.[625,963,736,798]" box="[625,963,736,763]" pageId="33" pageNumber="34">
<heading id="4A2691196574855AD34127C9FC3B9D82" box="[625,963,736,763]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="34" reason="2">
<taxonomicName id="D6D15DF66574855AD34127C9FD0A9D83" authorityName="Thuy" authorityYear="2013" box="[625,754,736,763]" class="Ophiuroidea" family="Ophiacanthidae" genus="Lapidaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA676574855AD34127C9FD0A9D83" bold="true" box="[625,754,736,763]" italics="true" pageId="33" pageNumber="34">Lapidaster</emphasis>
</taxonomicName>
sp. nov. innom. 1
</heading>
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE6574855AD3D8262DFCB39C66" box="[744,843,772,798]" pageId="33" pageNumber="34" type="description">
<paragraph id="116E26756574855AD3D8262DFCB39C66" blockId="33.[625,963,736,798]" box="[744,843,772,798]" pageId="33" pageNumber="34">
<figureCitation id="89EA3AF06574855AD3D8262DFCC89C66" box="[744,816,772,798]" captionStart="Fig" captionStartId="34.[189,232,1754,1780]" captionTargetBox="[258,1329,265,1714]" captionTargetId="figure@34.[258,1329,265,1714]" captionTargetPageId="34" captionText="Fig. 8. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1. Lapidaster sp. nov. innom. 1 from the late Toarcian (Early Jurassic) of Seewen, Switzerland; NHMB M11208, proximal to median LAP. 2-4. Lapidaster wolfii gen. et sp. nov. from the Bajocian-Bathonian boundary (Middle Jurassic) of Touert, France. 2. GZG.INV.78508 (holotype), proximal LAP. 3. GZG. INV.78509 (paratype), median LAP. 4. GZG.INV.78510 (paratype), distal LAP. 5-7. Lapidaster varuna gen. et sp. nov. from the Callovian of Jumara, India. 5. GZG.INV.78511 (holotype), proximal LAP. 6. GZG.INV.78512 (paratype), median LAP. 7. GZG.INV.78513 (paratype), distal LAP. One common scale bar per species is given." figureDoi="http://doi.org/10.5281/zenodo.3822854" httpUri="https://zenodo.org/record/3822854/files/figure.png" pageId="33" pageNumber="34">Fig. 8</figureCitation>
: 1
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE6574855AD18D266EFD859CFD" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="116E26756574855AD18D266EFD859CFD" blockId="33.[189,1399,838,901]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="31C1015B6574855AD18D266EFD859CFD" pageId="33" pageNumber="34">
<emphasis id="23A5FA676574855AD18D266EFE729C18" box="[189,394,838,864]" italics="true" pageId="33" pageNumber="34">
p.p.
<taxonomicName id="D6D15DF66574855AD1C1266FFE729C18" authorityName="Muller &amp; Troschel" authorityYear="1842" box="[241,394,838,864]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="genus">Ophiacantha</taxonomicName>
</emphasis>
?
<taxonomicName id="D6D15DF66574855AD090266EFDE09C18" authorityName="Hess" authorityYear="1962" box="[416,536,839,864]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="species" species="toarcensis">
<emphasis id="23A5FA676574855AD090266EFDE09C18" box="[416,536,839,864]" italics="true" pageId="33" pageNumber="34">toarcensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="907000646574855AD306266FFD179C18" box="[566,751,838,864]" pageId="33" pageNumber="34">
<bibRefCitation id="75405B846574855AD306266FFD179C18" author="Hess H." box="[566,751,838,864]" pageId="33" pageNumber="34" pagination="609 - 656" refId="ref122260" refString="Hess H. 1962. Mikropalaontologische Untersuchungen an Ophiuren II: Die Ophiuren aus dem Lias (Pliensbachien-Toarcien) von Seewen (Kt. Solothurn). Eclogae geologicae Helvetiae 55: 609 - 656." type="journal article" year="1962">Hess 1962: 649</bibRefCitation>
</treatmentCitation>
(Schicht C) (original specimen incorrectly assigned to
<taxonomicName id="D6D15DF66574855AD18D2643FEAE9CFC" authorityName="Muller &amp; Troschel" authorityYear="1842" box="[189,342,874,900]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA676574855AD18D2643FEAE9CFC" box="[189,342,874,900]" italics="true" pageId="33" pageNumber="34">Ophiacantha</emphasis>
</taxonomicName>
?
<taxonomicName id="D6D15DF66574855AD05A2642FD889CFD" authority="Hess, 1962" authorityName="Hess" authorityYear="1962" box="[362,624,874,901]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="species" species="toarcensis">
<emphasis id="23A5FA676574855AD05A2642FE1A9CFC" box="[362,482,875,900]" italics="true" pageId="33" pageNumber="34">toarcensis</emphasis>
<bibRefCitation id="75405B846574855AD0D92643FD889CFD" author="Hess H." box="[489,624,874,901]" pageId="33" pageNumber="34" pagination="609 - 656" refId="ref122260" refString="Hess H. 1962. Mikropalaontologische Untersuchungen an Ophiuren II: Die Ophiuren aus dem Lias (Pliensbachien-Toarcien) von Seewen (Kt. Solothurn). Eclogae geologicae Helvetiae 55: 609 - 656." type="journal article" year="1962">Hess, 1962</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE6574855AD18D2685FE8B9B62" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="116E26756574855AD18D2685FE529CBE" blockId="33.[189,426,940,966]" box="[189,426,940,966]" pageId="33" pageNumber="34">
<heading id="4A2691196574855AD18D2685FE529CBE" bold="true" box="[189,426,940,966]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="23A5FA676574855AD18D2685FE529CBE" bold="true" box="[189,426,940,966]" pageId="33" pageNumber="34">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="116E26756574855AD18D26F2FE8B9B62" blockId="33.[189,1398,987,1050]" pageId="33" pageNumber="34">
<collectionCode id="77C0BEB06574855AD18D26F2FEED9C8D" box="[189,277,987,1013]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="33" pageNumber="34">NHMB</collectionCode>
<collectionCode id="77C0BEB06574855AD02C26F2FECE9C8D" box="[284,310,987,1013]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="33" pageNumber="34" type="Herbarium">M</collectionCode>
11208 (dissociated LAP) from the late Toarcian of Seewen,
<collectingCountry id="69C666E56574855AD2C026F2FB879C8D" box="[1008,1151,987,1013]" name="Switzerland" pageId="33" pageNumber="34">Switzerland</collectingCountry>
, the original material of
<bibRefCitation id="75405B846574855AD1ED26D6FE979B62" author="Hess H." box="[221,367,1023,1050]" pageId="33" pageNumber="34" pagination="609 - 656" refId="ref122260" refString="Hess H. 1962. Mikropalaontologische Untersuchungen an Ophiuren II: Die Ophiuren aus dem Lias (Pliensbachien-Toarcien) von Seewen (Kt. Solothurn). Eclogae geologicae Helvetiae 55: 609 - 656." type="journal article" year="1962">Hess (1962)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE6574855AD18D2168FABF9992" pageId="33" pageNumber="34" type="description">
<paragraph id="116E26756574855AD18D2168FEB69B23" blockId="33.[189,334,1089,1115]" box="[189,334,1089,1115]" pageId="33" pageNumber="34">
<heading id="4A2691196574855AD18D2168FEB69B23" bold="true" box="[189,334,1089,1115]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="23A5FA676574855AD18D2168FEB69B23" bold="true" box="[189,334,1089,1115]" pageId="33" pageNumber="34">Description</emphasis>
</heading>
</paragraph>
<paragraph id="116E26756574855AD18D2159FB8E996B" blockId="33.[189,1399,1136,1771]" pageId="33" pageNumber="34">Single dissociated, median to proximal LAP, slightly wider than high, dorsal edge nearly straight, distal edge strongly convex; ventral fifth protruding ventro-proximalwards; ventro-distal tip of LAP large, rounded, protruding; proximal edge concave, with large, strongly protruding, sharply defined, horizontally elongate, lens-shaped, prominent spur in the middle, and with second horizontally elongated, lens-shaped but slightly smaller, less well-defined, less prominent and protruding spur close to ventroproximal tip of LAP; outer surface with very slight constriction, and with coarsely meshed stereom; trabeculae of outer surface stereom showing weak tendency to merge into short, irregular vertical stripes close to row of spine articulations. Four small spine articulations, ventralmost slightly smaller than other three; dorsal and ventral lobes separated by small, round knob; strong dorsal increase in size of gaps separating spine articulations; two ventral spine articulations separated from coarsely meshed outer surface stereom by faint, irregular, confluent areoles; two dorsal spine articulations embedded in outer surface stereom. Ventral edge of LAP with large, moderately deep tentacle notch.</paragraph>
<paragraph id="116E26756574855AD18D2337FABF9992" blockId="33.[189,1399,1136,1771]" pageId="33" pageNumber="34">Inner side of LAP with sharply defined, dorsal portion of which straight to slightly bent, not widened dorsally and not approaching ventral or dorsal edges of LAP; ventral portion of LAP separated from dorsal one by rounded kink, very short and sharply defined, not merging with ventral part of LAP; inner distal edge with two large, well-defined, horizontally elongate, prominent spurs composed of dense stereom, ventral one of which slightly protruding. Inner side of tentacle notch moderately large and with coarsely meshed, slightly horizontally stretched stereom. No perforations discernible on inner side.</paragraph>
</subSubSection>
<subSubSection id="59CB75FE65748558D18D223AFBBA9EE1" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="discussion">
<paragraph id="116E26756574855AD18D223AFED79855" blockId="33.[189,303,1811,1837]" box="[189,303,1811,1837]" pageId="33" pageNumber="34">
<heading id="4A2691196574855AD18D223AFED79855" bold="true" box="[189,303,1811,1837]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="23A5FA676574855AD18D223AFED79855" bold="true" box="[189,303,1811,1837]" pageId="33" pageNumber="34">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="116E267565748558D18D226BFBBA9EE1" blockId="33.[189,1399,1857,2027]" lastBlockId="35.[189,1398,267,410]" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
Among the original material of
<taxonomicName id="D6D15DF66574855AD3032268FD349823" authorityName="Muller &amp; Troschel" authorityYear="1842" box="[563,716,1857,1883]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA676574855AD3032268FD349823" box="[563,716,1857,1883]" italics="true" pageId="33" pageNumber="34">Ophiacantha</emphasis>
</taxonomicName>
?
<taxonomicName id="D6D15DF66574855AD3D0226BFC1A9823" authority="Hess, 1962" authorityName="Hess" authorityYear="1962" box="[736,994,1857,1884]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="species" species="toarcensis">
<emphasis id="23A5FA676574855AD3D0226BFCA09823" box="[736,856,1858,1883]" italics="true" pageId="33" pageNumber="34">toarcensis</emphasis>
<bibRefCitation id="75405B846574855AD26F226BFC1A9823" author="Hess H." box="[863,994,1857,1884]" pageId="33" pageNumber="34" pagination="609 - 656" refId="ref122260" refString="Hess H. 1962. Mikropalaontologische Untersuchungen an Ophiuren II: Die Ophiuren aus dem Lias (Pliensbachien-Toarcien) von Seewen (Kt. Solothurn). Eclogae geologicae Helvetiae 55: 609 - 656." type="journal article" year="1962">Hess, 1962</bibRefCitation>
</taxonomicName>
from the Toarcian of
<collectingCountry id="69C666E56574855AD5D32268FA8A9823" box="[1251,1394,1857,1883]" name="Switzerland" pageId="33" pageNumber="34">Switzerland</collectingCountry>
, here reassigned to
<taxonomicName id="D6D15DF66574855AD0A3224FFD539807" authority="Hess, 1964" authorityName="Hess" authorityYear="1964" box="[403,683,1893,1920]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA676574855AD0A3224FFDDC9807" box="[403,548,1894,1919]" italics="true" pageId="33" pageNumber="34">Dermocoma</emphasis>
<bibRefCitation id="75405B846574855AD31A224FFD539807" author="Hess H." box="[554,683,1893,1920]" pageId="33" pageNumber="34" pagination="756 - 801" refId="ref122328" refString="Hess H. 1964. Die Ophiuren des englischen Jura. Eclogae geologicae Helvetiae 57: 756 - 801." type="journal article" year="1964">Hess, 1964</bibRefCitation>
</taxonomicName>
(see below), is a single LAP which obviously does not belong to that species. A detailed re-examination of that plate has revealed that it is a species of
<taxonomicName id="D6D15DF66574855AD58F22A0FAC698DB" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1215,1342,1929,1955]" class="Ophiuroidea" family="Ophiacanthidae" genus="Lapidaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA676574855AD58F22A0FAC698DB" box="[1215,1342,1929,1955]" italics="true" pageId="33" pageNumber="34">Lapidaster</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C6574855AD47522A3FF0998BF" pageId="33" pageNumber="34" rank="genus">gen. nov.</taxonomicNameLabel>
, on account of its
<taxonomicName id="D6D15DF66574855AD0FA2284FD8898BF" authorityName="H. L. Clark" authorityYear="1911" box="[458,624,1965,1991]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiologimus" kingdom="Animalia" order="Ophiurida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA676574855AD0FA2284FD8898BF" box="[458,624,1965,1991]" italics="true" pageId="33" pageNumber="34">Ophiologimus</emphasis>
</taxonomicName>
-like morphology and the presence of spurs on the outer proximal and inner distal edges. The unusually small spine articulations composed of separated dorsal and ventral lobes, as well as the presence of a second, strongly developed spur in the centre of the outer proximal and inner distal edges sets the LAP in question quite apart from other LAP
<typeStatus id="CE6A98D765768558D519241BFB9F9E34" box="[1065,1127,306,332]" pageId="35" pageNumber="36">types</typeStatus>
assigned to
<taxonomicName id="D6D15DF665768558D5C7241BFA8E9E34" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1271,1398,306,332]" class="Ophiuroidea" family="Ophiacanthidae" genus="Lapidaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="35" pageNumber="36" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA6765768558D5C7241BFA8E9E34" box="[1271,1398,306,332]" italics="true" pageId="35" pageNumber="36">Lapidaster</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65768558D18D2470FED09E0B" box="[189,296,345,371]" pageId="35" pageNumber="36" rank="genus">gen. nov.</taxonomicNameLabel>
It unquestionably represents a distinct, undescribed species. Unfortunately, only a single LAP is currently known, which precludes the formal description of a new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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