201 lines
24 KiB
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201 lines
24 KiB
XML
<document ID-CLB-Dataset="291769" ID-DOI="http://dx.doi.org/10.3897/asp.82.e111148" ID-GBIF-Dataset="de602973-4f52-4de2-8014-342e7778d97b" ID-Pensoft-Pub="1864-8312-82-151" ID-Pensoft-UUID="8D187145BFCA55A1A73960CDE5BD124B" ID-ZooBank="43AC036E93CC4D79939A07DF54BE1A2D" ModsDocID="1864-8312-82-151" checkinTime="1711188240520" checkinUser="pensoft" docAuthor="Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent" docDate="2024" docId="982B71A3D015503EACAFCDC54F6F2963" docLanguage="en" docName="ArthropodSystPhyl 82: 151-181" docOrigin="Arthropod Systematics & amp; Phylogeny 82" docPubDate="2024-03-22" docSource="http://dx.doi.org/10.3897/asp.82.e111148" docTitle="Cretolyra noijebumensis Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024, sp. nov." docType="treatment" docUuid="5C43B26B-D68E-45EC-9E3F-47C229B98AB7" docUuidSource="ZooBank" docVersion="4" id="8D187145BFCA55A1A73960CDE5BD124B" lastPageNumber="151" masterDocId="8D187145BFCA55A1A73960CDE5BD124B" masterDocTitle="Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities" masterLastPageNumber="181" masterPageNumber="151" pageNumber="151" updateTime="1732880532958" updateUser="ExternalLinkService">
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<mods:title id="439D68CCF113DE499735775F089D90BA">Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities</mods:title>
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<mods:namePart id="1E85A7438B8565F3AED28F83BCE06447">Brazidec, Manuel</mods:namePart>
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<mods:nameIdentifier id="FE5147E8CBB262318BA5DE9B81B217BE" type="ORCID">https://orcid.org/0000-0002-0860-8972</mods:nameIdentifier>
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<mods:affiliation id="15C60C996F28AB96ECEAEEA1534DD0CB">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France & Institut de Syste ́ matique, E ́ volution, Biodiversite ́ (ISYEB), Muse ́ um national d'Histoire naturelle, CNRS, Sorbonne Universite ́, EPHE, Universite ́ des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:nameIdentifier id="E2E84459A8BF8E59B7D391A4F8C5AA2D" type="email">manuel.brazidec@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C08D1851A5AB92991E4842BA5B233FAB">Vilhelmsen, Lars</mods:namePart>
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<mods:affiliation id="0D9B4378A67CAF6D132E28FF3BAE699D">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark</mods:affiliation>
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<mods:namePart id="B262432330B99C4FCB4C5FD0467EAE1D">Boudinot, Brendon E.</mods:namePart>
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<mods:nameIdentifier id="1060CF3770E2DA331AE49555774C66A0" type="ORCID">https://orcid.org/0000-0002-4588-0430</mods:nameIdentifier>
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<mods:affiliation id="57DFDB4700659E74AFD51050C7357435">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & National Museum of Natural History, Smithsonian Institution, 10 th & Constitution Ave. NW, Washington, DC, USA & Senckenberg Gesellschaft fuer Naturforschung und Naturmuseum, Frankfurt am Main, 60325 Germany</mods:affiliation>
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<mods:namePart id="785816E1441A073A2528FD5D2DBD6036">Richter, Adrian</mods:namePart>
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<mods:nameIdentifier id="04A2E2625FC690635654FAF0A751E417" type="ORCID">https://orcid.org/0000-0001-5627-2302</mods:nameIdentifier>
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<mods:affiliation id="8CDDCE66169B06F2CA7173FD3E26F039">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & Biodiversity and Biocomplexity Unit, Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa, Japan</mods:affiliation>
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<mods:namePart id="82D90B224DBA0D755571D4EFC9DC87E0">Hammel, Joerg U.</mods:namePart>
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<mods:nameIdentifier id="2DE2F2BD84A5D34FF471A442F81B8EF7" type="ORCID">https://orcid.org/0000-0002-6744-6811</mods:nameIdentifier>
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<mods:affiliation id="D497E6E1E9ABB5579FC7C8EEE81636EF">Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Str. 1, 21502 Geesthacht, Germany</mods:affiliation>
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<mods:namePart id="DDEB4D41DDAABEFCE66275EBBA36CAC9">Perkovsky, Evgeny E.</mods:namePart>
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<mods:nameIdentifier id="7C27BC6DEC4826B9D8FC009343EBE963" type="ORCID">https://orcid.org/0000-0002-7959-4379</mods:nameIdentifier>
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<mods:affiliation id="AA79B6832E40A42B6ED5F2C318C189E9">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & I. I. Schmalhausen Institut of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine</mods:affiliation>
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<mods:namePart id="827E70FE1F4812E4133F099930D52542">Fan, Yong</mods:namePart>
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<mods:affiliation id="574AFAC9498A7A47D9043863ED9AD3E5">Fushun Amber Institute, Fushun 113005, China</mods:affiliation>
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<mods:namePart id="557BE34CA3D967D967CAB328AC219FF3">Wang, Zhen</mods:namePart>
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<mods:affiliation id="9054202E7F1DB94D7682827662B67402">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="A7D8D197BC9471D8B566307F5DB0C7FE">Wu, Qiong</mods:namePart>
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<mods:affiliation id="C2A1F3B2F70F667F3DE87BD799959138">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="3A9701317C4B61DB758CAE861416A471">Wang, Bo</mods:namePart>
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<mods:affiliation id="D54BD74566D779E290BA7EC979412E40">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="11C9B261BB1F9F405CC3C835CDFEA04D">Perrichot, Vincent</mods:namePart>
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<mods:nameIdentifier id="D1A61779A0C7EC41BEF0E65CB22978DD" type="ORCID">https://orcid.org/0000-0002-7973-0430</mods:nameIdentifier>
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<mods:affiliation id="C855965BEC9987BC126313DB944085CC">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France</mods:affiliation>
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<mods:typeOfResource id="7644D5A552CD960DAF8F6194D20CEBCD">text</mods:typeOfResource>
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<mods:title id="B82489BA5087B66550D6519B65BFFFFA">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part id="3DB249DDD27E1EDCA3AF75F523BADFE1">
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<mods:date id="47B63E32AD374C11DCB56F4782800D29">2024</mods:date>
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<mods:number id="302E9C2F4BA4E33226B8C14328E8B3CA">2024-03-22</mods:number>
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</mods:detail>
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<mods:detail id="F9632165FCE2C86B44DED2797D311C2D" type="volume">
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<mods:number id="F93F6943FF50059AD48C0F8A3D264D42">82</mods:number>
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<mods:start id="9F3DBF07608C542FD8FF76B1D612DA9C">151</mods:start>
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<mods:end id="0B1628BD5F47F181237575D070A5ABAE">181</mods:end>
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<mods:url id="1EE602D15FDC0388AC5AF2539E346D2E">http://dx.doi.org/10.3897/asp.82.e111148</mods:url>
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<mods:classification id="0BB0F127757122C306684EA870B18C64">journal article</mods:classification>
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<mods:identifier id="1876EB7E24B2F2403113991DC8F0B141" type="DOI">http://dx.doi.org/10.3897/asp.82.e111148</mods:identifier>
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<treatment id="982B71A3D015503EACAFCDC54F6F2963" ID-GBIF-Taxon="224057655" LSID="urn:lsid:zoobank.org:act:5C43B26B-D68E-45EC-9E3F-47C229B98AB7" httpUri="http://treatment.plazi.org/id/982B71A3D015503EACAFCDC54F6F2963" lastPageNumber="151" pageId="0" pageNumber="151" scope_family="Dinapsidae" scope_order="Hymenoptera" scope_superFamily="Megalyroidea">
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<subSubSection id="9D3A3579DF8F686B20A4B6C800BF108E" pageId="0" pageNumber="151" type="nomenclature">
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<paragraph id="42F39D607840FD2CBB7FD9C8F283F558" pageId="0" pageNumber="151">
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3.1.1.1.1. †
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<taxonomicName id="537EBA9DA425C1C27C4C9ADB06EA9EFD" LSID="https://zoobank.org/5C43B26B-D68E-45EC-9E3F-47C229B98AB7" authority="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" class="Insecta" family="Megalyridae" genus="Cretolyra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretolyra noijebumensis" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="noijebumensis" status="sp. nov.">Cretolyra noijebumensis</taxonomicName>
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<taxonomicNameLabel id="D9D9BD3CFCE5F1737E05DA3E8E21E70B" pageId="0" pageNumber="151">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="D1DBDBD0EB0B7385D05259C42CC86E92" pageId="0" pageNumber="151" type="description">
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<paragraph id="577C175034BE53D1A1DDD9DDE7FBCD58" pageId="0" pageNumber="151">
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<figureCitation id="2AE559744C0EB0B3FEA42329E2413315" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H)." figureDoi="10.3897/asp.82.e111148.figure2" httpUri="https://binary.pensoft.net/fig/1007131" pageId="0" pageNumber="151">Figures 2A</figureCitation>
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<figureCitation id="04A4B1BC3CAEDC6F18C41BC002EF24B5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Cretolyra noijebumensis gen. et sp. nov., holotype male IGR. BU- 067, mid-Cretaceous, Kachin amber. A habitus in lateral view; B head and mesosoma in dorsal view; C fore wing; D anterior part of mesosoma in lateral view with structures outlined; E metasoma and posterior part of the mesosoma in lateral view. Scale bars: 0.5 mm (A, B, C, E); 0.25 mm (D). Abbreviations: AG = inner axillar groove; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure3" httpUri="https://binary.pensoft.net/fig/1007132" pageId="0" pageNumber="151">, 3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="ACEFFBE7B41B384723E0E4C78A84D9A0" pageId="0" pageNumber="151" type="etymology">
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<paragraph id="754166BC9DA253BE825323E3F34DD8C3" pageId="0" pageNumber="151">Etymology.</paragraph>
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<paragraph id="45DA19B9574C74F6708B58DF6064AF72" pageId="0" pageNumber="151">The specific epithet is an adjective referring to the type locality where the amber pieces were collected.</paragraph>
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</subSubSection>
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<subSubSection id="F4A93D55C6DF3CD7AC5632FCC043166F" pageId="0" pageNumber="151" type="materials_examined">
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<paragraph id="562FB72940D076B00FF89761FB77DBDB" pageId="0" pageNumber="151">Material studied.</paragraph>
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<paragraph id="B7111A358F1E4896E6BE71723980A0F6" pageId="0" pageNumber="151">
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<materialsCitation id="AEF72E47757A90CE93AF563381DF0DB0" ID-GBIF-Occurrence="4597783303" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="France" county="University of Rennes" location="Geology Department" municipality="Museum" specimenCount="1" typeStatus="Holotype">
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<emphasis id="ED1666A98025A6EF050EF515C413BBC6" bold="true" italics="true" pageId="0" pageNumber="151">
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<typeStatus id="066EEBF67CAC9193710A9BE5FD9A674D" type="holotype">Holotype</typeStatus>
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</emphasis>
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male IGR.BU-067; housed in the amber collection of the
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<location id="ACBA38BF3E04ACEAB970B162D377A41C" LSID="urn:lsid:plazi:treatment:982B71A3D015503EACAFCDC54F6F2963:ACBA38BF3E04ACEAB970B162D377A41C" country="France" county="University of Rennes" municipality="Museum" name="Geology Department">Geology Department</location>
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and
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<collectingMunicipality id="BD50BA18F5038DDF21519A7C3C69D55E">Museum</collectingMunicipality>
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of the
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<collectingCounty id="B420877B12E3AFBD3DE32E5C66CFBD0D">University of Rennes</collectingCounty>
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,
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<collectingCountry id="9F8F1B6E87AF5C2B4CB1272B06ED1C4D" name="France">France</collectingCountry>
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(IGR).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="301A6A5A27E9A8B88562B8093B9BF3A4" pageId="0" pageNumber="151" type="type locality">
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<paragraph id="FE87D737060D284F97C88DE49C2405B3" pageId="0" pageNumber="151">Type locality.</paragraph>
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<paragraph id="6FA67D07D0FF6F2B894EF6A82B6A0D28" pageId="0" pageNumber="151">Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.</paragraph>
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</subSubSection>
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<subSubSection id="B71AD2345826F0DA5C854B16CD255D80" pageId="0" pageNumber="151" type="age">
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<paragraph id="A150078F832757A3DA15B68BAB693DDA" pageId="0" pageNumber="151">Age.</paragraph>
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<paragraph id="0C8635492976BCB88C5AC8F4530141B8" pageId="0" pageNumber="151">Upper Albian to lower Cenomanian, mid-Cretaceous.</paragraph>
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</subSubSection>
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<subSubSection id="38DFC7C523ED54712704A00231403378" pageId="0" pageNumber="151" type="diagnosis">
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<paragraph id="C882C4E654D5D03572EEE0ED53CFF3DB" pageId="0" pageNumber="151">Diagnosis.</paragraph>
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<paragraph id="991E2DC2CD077526EA9ADB4619350333" pageId="0" pageNumber="151">
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Antennae inserted on edge of clypeus (Fig.
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<figureCitation id="B86B93EBA7000BAC9002F4ACD71EA218" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Cretolyra noijebumensis gen. et sp. nov., holotype male IGR. BU- 067, mid-Cretaceous, Kachin amber. A habitus in lateral view; B head and mesosoma in dorsal view; C fore wing; D anterior part of mesosoma in lateral view with structures outlined; E metasoma and posterior part of the mesosoma in lateral view. Scale bars: 0.5 mm (A, B, C, E); 0.25 mm (D). Abbreviations: AG = inner axillar groove; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure3" httpUri="https://binary.pensoft.net/fig/1007132" pageId="0" pageNumber="151">3A</figureCitation>
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; vs. inserted above clypeal margin in
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<taxonomicName id="3894100551F1CC8FF9789D408521BD20" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" class="Insecta" family="Megalyridae" genus="Cretolyra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretolyra shawi" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="shawi">
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<emphasis id="956CF4B0AB0E597528B6EC6959A698B9" italics="true" pageId="0" pageNumber="151">Cretolyra shawi</emphasis>
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</taxonomicName>
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<emphasis id="6963A61C51EF7F98BEE87248E76A3566" bold="true" pageId="0" pageNumber="151">gen. et sp. nov.</emphasis>
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); axillae contiguous medially and axillar grooves crenulated (Fig.
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<figureCitation id="B6A33820463BD3A702C29154E674803F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Cretolyra noijebumensis gen. et sp. nov., holotype male IGR. BU- 067, mid-Cretaceous, Kachin amber. A habitus in lateral view; B head and mesosoma in dorsal view; C fore wing; D anterior part of mesosoma in lateral view with structures outlined; E metasoma and posterior part of the mesosoma in lateral view. Scale bars: 0.5 mm (A, B, C, E); 0.25 mm (D). Abbreviations: AG = inner axillar groove; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure3" httpUri="https://binary.pensoft.net/fig/1007132" pageId="0" pageNumber="151">3B</figureCitation>
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; vs axillae separated medially and axillar grooves smooth in
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<taxonomicName id="5BB48C407BE1B2A990C9010BD615F771" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" class="Insecta" family="Megalyridae" genus="Cretolyra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretolyra shawi" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="shawi">
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<emphasis id="C78F418C7095736ACF4EC7895C632D69" italics="true" pageId="0" pageNumber="151">Cretolyra shawi</emphasis>
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</taxonomicName>
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<emphasis id="9758CB112BDBFCE29CAE75D6D9C32080" bold="true" pageId="0" pageNumber="151">gen. et sp. nov.</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="839E3F8AC4C394E1D1CDB5B05FA0CF18" pageId="0" pageNumber="151" type="description">
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<paragraph id="D85D5B5783A801698EF8D48AD80AEC80" pageId="0" pageNumber="151">Description.</paragraph>
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<paragraph id="9ECEA04BF18EE68297FF90CEFDA8D2A9" pageId="0" pageNumber="151">
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Body length 3.33 mm; body very pubescent, especially on metasoma where setae conceal margins of tergites; the setae have agglomerated small bubbles that make observation difficult. -
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<emphasis id="70298CC6DFB67FD5C6AB3078141F87E7" bold="true" italics="true" pageId="0" pageNumber="151">Head</emphasis>
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globular, higher than long (length 0.55 mm; height 0.75 mm); frons convex, smooth, divided by thin median sulcus; compound eye oval, higher than long; vertex convex, with smooth ocellar triangle; clypeus well-developed; torulus inserted closer to clypeus than to eye; shallow but distinct subantennal groove adjoining ventral margin of eye; antenna half as long as body, almost reaching metasoma; scape twice as long as wide (length 0.16 mm); pedicel shorter than scape (length 0.11 mm); flagellomeres cylindrical, elongate, longer than wide; flagellomere 1 shortest (length 0.15 mm); following flagellomeres longer (length ca. 0.21 mm); occipital carina crenulate. -
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<emphasis id="C53C3F61CF3A77B5A494D012B9C9FD7C" bold="true" italics="true" pageId="0" pageNumber="151">Mesosoma</emphasis>
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almost half body length (length 1.29 mm; height 0.65 mm); mesoscutum convex, shagreened, divided by smooth median mesoscutal sulcus (mesoscutum length 0.58 mm; width ~0.60 mm); parapsidal lines present, diverging anteriorly; axillae contiguous medially, axillar groove crenulate; mesoscutellum diamond-shaped; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum shorter than mesoscutellum, with incomplete pairs of median and submedian carinae, two complete lateral longitudinal carinae, anterior region of propodeum with transversal row of foveae adjacent to concealed metanotum, posterior region of propodeum with three large foveae. -
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<emphasis id="EC46E99D478C5F9F7519DF90322D3F3E" bold="true" italics="true" pageId="0" pageNumber="151">Fore wing</emphasis>
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hyaline and covered with microtrichiae, about two thirds of body length (length 2.50 mm); C, Sc+R, M+Cu, A, Rs fully pigmented; R1 extending beyond marginal cell; Rs closing marginal cell in straight line; M pigmented to apex; medial cell rectangular, narrow; Cu almost reaching posterior wing margin. -
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<emphasis id="D06116B46DC60CCF978F02EF88B18EB4" bold="true" italics="true" pageId="0" pageNumber="151">Legs</emphasis>
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covered with setae; two mesotibial and one metatibial spurs; metacoxa elongate; metafemur and metatibia swollen; metabasitarsus three times as long as following metatarsomeres and with row of short comb-like setae along ventral surface. -
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<emphasis id="1F0F576653CE146B618D8D901588E419" bold="true" italics="true" pageId="0" pageNumber="151">Metasoma</emphasis>
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half body length (length 1.48 mm), elongate and narrowed at apex; seven smooth tergites with pubescent posterior margin; first tergite largest; second tergite one quarter of metasoma length; remaining tergites shorter.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |