309 lines
37 KiB
XML
309 lines
37 KiB
XML
<document id="E607EDFBEF4883D5992FBB0233EAF26B" ID-CLB-Dataset="28693" ID-DOI="http://dx.doi.org/10.3897/zookeys.591.7731" ID-GBIF-Dataset="bcdbe5f4-2f0e-4240-8727-64b6c91db94a" ID-PMC="PMC4926649" ID-Pensoft-Pub="1313-2970-591-1" ID-PubMed="27408541" ID-ZooBank="DD65A7BB1B834CB298BA7E8E2257FA4F" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-591-1" ModsDocOrigin="ZooKeys 591" ModsDocTitle="A revision of Chilicola (Heteroediscelis), a subgenus of xeromelissine bees (Hymenoptera, Colletidae) endemic to Chile: taxonomy, phylogeny, and biogeography, with descriptions of eight new species" checkinTime="1463833188709" checkinUser="pensoft" docAuthor="Monckton, Spencer K." docDate="2016" docId="614148BC4C43B1C77BE9149999874363" docLanguage="en" docName="ZooKeys 591: 1-144" docOrigin="ZooKeys 591" docSource="http://dx.doi.org/10.3897/zookeys.591.7731" docTitle="Chilicola (Heteroediscelis) mavida Toro & Moldenke 1979" docType="treatment" docVersion="6" lastPageNumber="77" masterDocId="FFFED84DD544452EFFC446356171FFFD" masterDocTitle="A revision of Chilicola (Heteroediscelis), a subgenus of xeromelissine bees (Hymenoptera, Colletidae) endemic to Chile: taxonomy, phylogeny, and biogeography, with descriptions of eight new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="71" updateTime="1701383329186" updateUser="ExternalLinkService">
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<mods:title id="00C20A61E8A07EB3E559D850C3AE43DE">A revision of Chilicola (Heteroediscelis), a subgenus of xeromelissine bees (Hymenoptera, Colletidae) endemic to Chile: taxonomy, phylogeny, and biogeography, with descriptions of eight new species</mods:title>
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<mods:namePart id="9656AD8A27A4C5FF96366DCAA0B20A37">Monckton, Spencer K.</mods:namePart>
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<mods:typeOfResource id="12D341B224B375883D196F58E4116FC2">text</mods:typeOfResource>
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<mods:title id="5B94F79F4C325A3E92E96C25C3EDBF9E">ZooKeys</mods:title>
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<mods:part id="90714C376B4565AFA2B6E66EF2A4C24E">
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<mods:date id="107987F9F931092ED5A38ADE7AFE63DF">2016</mods:date>
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<mods:number id="B8352278DF349C21FC663B6ABC4FBC90">591</mods:number>
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<mods:url id="6373867A8E94BF4AA113778E55775260">http://dx.doi.org/10.3897/zookeys.591.7731</mods:url>
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<mods:classification id="FDB47C9469C51C95F7B85405EAFF554A">journal article</mods:classification>
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<mods:identifier id="6F3560B8BDF6D7511063FC7F6228AC06" type="DOI">http://dx.doi.org/10.3897/zookeys.591.7731</mods:identifier>
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<mods:identifier id="AC15A5B3C17434D9FA3646A66C128075" type="Pensoft-Pub">1313-2970-591-1</mods:identifier>
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</mods:mods>
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<treatment id="614148BC4C43B1C77BE9149999874363" ID-GBIF-Taxon="150886268" LSID="urn:lsid:plazi:treatment:614148BC4C43B1C77BE9149999874363" httpUri="http://treatment.plazi.org/id/614148BC4C43B1C77BE9149999874363" lastPageId="76" lastPageNumber="77" pageId="70" pageNumber="71">
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<subSubSection id="1FF1BA92A3EFF750D62386BE570580A9" pageId="70" pageNumber="71" type="multiple">
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<paragraph id="AC8BB20B67DAA4C208378460E36112AD" pageId="70" pageNumber="71">
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<pageBreakToken id="53170AF15FF67853E30A4324AE3319AB" pageId="70" pageNumber="71" start="start">Taxon</pageBreakToken>
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classification Animalia Hymenoptera Colletidae
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</paragraph>
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</subSubSection>
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<subSubSection id="477C84A5798B7CCEC30FE79A256293E1" pageId="70" pageNumber="71" type="nomenclature">
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<paragraph id="17335DFF57852C30EFBC39AD0D4D4FD9" pageId="70" pageNumber="71">
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<taxonomicName id="8E14D3A977CEDA3E9744008E6F93C76F" ID-CoL="TZW3" authority="Toro & Moldenke, 1979" authorityName="Toro & Moldenke" authorityYear="1979" class="Insecta" family="Colletidae" genus="Chilicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chilicola (Heteroediscelis) mavida" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="mavida" subGenus="Heteroediscelis">Chilicola (Heteroediscelis) mavida Toro & Moldenke, 1979</taxonomicName>
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Figs 5A; 6C; 9
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<normalizedToken id="E7A8189D798E893DC337455041B26ADA" originalValue="A–B">A-B</normalizedToken>
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; 21B; 23A; 24
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<normalizedToken id="7CA7AD2ED8C495B8DF70957CACCCC8FB" originalValue="C–D">C-D</normalizedToken>
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; 41
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<normalizedToken id="1C989F73549E2CA018AB4A85D46F3E4F" originalValue="A–J">A-J</normalizedToken>
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; 49
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</paragraph>
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</subSubSection>
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<subSubSection id="306DA3A8E87D872E530EA606AA1E9B8C" pageId="71" pageNumber="72" type="reference_group">
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<paragraph id="0E58586DFF5F5CBAC0FB78ECD7262189" pageId="71" pageNumber="72">
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<taxonomicName id="DC1A989C876469AB341F334C3A86A2A6" class="Insecta" family="Colletidae" genus="Chilicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chilicola mavida" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="mavida">
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<pageBreakToken id="54EFEFF09D3D1D521CEC414C8EDE80C9" pageId="71" pageNumber="72" start="start">Chilicola</pageBreakToken>
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mavida
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</taxonomicName>
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Toro & Moldenke, 1979: 122-124 (male holotype, female allotype, AMNH [examined]).
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<bibRefCitation id="EA864831291373FE7079FAA75306A508" author="Michener, CD" journalOrPublisher="Scientific Papers, Natural History Museum, The University of Kansas" pageId="124" pageNumber="125" pagination="1 - 47" title="The Bee Genus Chilicola in the Tropical Andes, with Observations on Nesting Biology and a Phylogenetic Analysis of the Subgenera (Hymenoptera: Colletidae, Xeromelissinae)." url="10.5962/bhl.title.8449" volume="26" year="2002">Michener 2002</bibRefCitation>
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: 5-7 (phylogeny).
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<bibRefCitation id="AF332A0147154482494FF2B46F4C3EC9" author="Packer, L" journalOrPublisher="Systematic Entomology" pageId="125" pageNumber="126" pagination="72 - 96" title="Phylogeny and classification of the Xeromelissinae (Hymenoptera: Apoidea, Colletidae) with special emphasis on the genus Chilicola." url="10.1111/j.1365-3113.2007.00398.x" volume="33" year="2008">Packer 2008</bibRefCitation>
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: 77. Montalva and Ruz 2010: 28 (checklist).
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<bibRefCitation id="7514EFAC606585DF1712998234479011" author="Moure, JS" journalOrPublisher="Zootaxa" pageId="125" pageNumber="126" title="Catalogue of Bees (Hymenoptera, Apoidea) in the Neotropical Region - online version" url="http://www.moure.cria.org.br/catalogue" year="2012">Moure et al. 2012</bibRefCitation>
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(catalogue).
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<bibRefCitation id="71755D28A67EA7BCB5B8EEE84775BD31" author="Monckton, SK" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="124" pageNumber="125" pagination="234 - 236" title="Case 3670: Chilicolavicugna Toro & Moldenke, 1979 (Insecta, Hymenoptera, COLLETIDAE): proposed replacement of the holotype by a neotype." volume="71" year="2014">Monckton 2014</bibRefCitation>
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: 234.
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<bibRefCitation id="267D58D63F55C9F59FA2462785C3BAEA" author="Ascher, JS" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="122" pageNumber="123" title="Discover Life Bee Species Guide and World Checklist (Hymenoptera: Apoidea: Anthophila)" url="http://www.discoverlife.org/mp/20q?guide=Apoidea_species" year="2015">Ascher and Pickering 2015</bibRefCitation>
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(checklist).
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</paragraph>
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</subSubSection>
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<subSubSection id="4AE316FB27E9451AFF975A813283FF23" pageId="71" pageNumber="72" type="diagnosis">
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<paragraph id="4C6867AA19D997977200EEAA05E306BB" pageId="71" pageNumber="72">Diagnosis.</paragraph>
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<paragraph id="A619880C3CFB287DCE69782403E1418D" pageId="71" pageNumber="72">
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Males are diagnosable by the combination of first flagellomere longer than wide (≥1.1
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<normalizedToken id="6E4E2D7BB7F0370775DC20B5342F61FD" originalValue="×">x</normalizedToken>
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), malar space longer than clypeal lateral (1.05
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<normalizedToken id="CB6B46E1EFC5324B548F2393301DEEA8" originalValue="–1.1×">-1.1x</normalizedToken>
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), and S1 process with anterior apical angle acute in lateral view. Females are diagnosable by the combination of malar space subequal in length to the clypeal lateral (0.9
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<normalizedToken id="2AABA157BCDC466130746ADC9430324F" originalValue="–1×">-1x</normalizedToken>
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) and leg colouration as follows: apicodorsal spot on mesotibia yellow-brown, apicodorsal rim of metafemur yellow, wide basal ring of metatibia yellow, posterior apical rim of metatibia yellow-brown. Females of
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<taxonomicName id="834F180CADF290F29244978444CE358A" class="Insecta" family="Colletidae" genus="Chilicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chilicola curvapeligrosa" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="curvapeligrosa">Chilicola curvapeligrosa</taxonomicName>
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are most similar, but have darker leg colouration (following parts brown: apicodorsal spot on mesotibia, apicodorsal rim of metafemur, basidorsal spot on metatibia; apex of metatibia black-brown).
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</paragraph>
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</subSubSection>
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<subSubSection id="3E9A747D25DCA639131AEEB89DD27CEA" lastPageId="75" lastPageNumber="76" pageId="71" pageNumber="72" type="description">
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<paragraph id="698419E50132E3D065089F0F25FFBC24" pageId="71" pageNumber="72">Description.</paragraph>
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<paragraph id="9A175FEEA6425EB9C9A882FCBCADDCE4" pageId="71" pageNumber="72">Male. Length 5.3-5.8mm, forewing length 3.4-3.5mm, head width 1.1-1.2mm, thorax width 1.1-1.2mm, median ocellar diameter (OD) 0.13mm.</paragraph>
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<paragraph id="D58A922500407475F8D8EA786941B92E" pageId="71" pageNumber="72">Colouration: Black-brown, following parts yellow: labrum; clypeus except broadly dark along epistomal suture; lower paraocular area, extending ~1OD above transverse portion of epistomal suture and not reaching lower tangent of antennal socket; apicoventral spot on scape; apicoventral surface of pedicel; protrochanter apical rim posteriorly; apex of profemur broadly on anterior surface, narrowly on posterior surface; protibia except dark medioventrally; probasitarsus dorsally, at least in basal half; apex of mesofemur broadly on anterior surface, narrowly on posterior surface; broad basal and apical rings on mesotibia, the latter widened to apical half anteriorly; at least basoventral spot on mesobasitarsus; apicoventral rim of metacoxa and metatrochanter, the latter sometimes yellow-brown; narrow apical ring on metafemur; metatibia in basal one-third, in apical quarter of anterior surface, apical one-third of posterior surface and along ventral varina, yellow colouration continuous along ventral surface, sometimes continuous or nearly so along dorsal surface; metabasitarsus in basal one-third dorsally, basal two-thirds ventrally; anterior spot on tegula; apex of T7. Ventral surface of flagellum yellow-brown, flagellomeres suffused with brown basally. Apical half of prodistitarsus and apical one-third of mesodistitarsus brown. Posterior metatibial carina black. Metasoma black, T1-T6 marginal zones translucent yellow.</paragraph>
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<paragraph id="DFE565855DE0048E932DD733214B52EC" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
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Pubescence: White, hairs generally short (0.5OD) and sparse, not especially plumose; on lower paraocular area and around antennal base moderately long and moderately dense (1OD); genal beard long and dense (0.5-3OD) longest at midlength; discs of mesoscutum, scutellum, and metanotum mostly bare, few short hairs (0.5OD), longer and denser toward lateral margin as follows: mesoscutum (0.5-1OD), scutellum (1-2OD) also on posterior margin, longest posteromedially, metanotum (≤2OD);
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<pageBreakToken id="491FB547E416D31E1AD5FFF2E12D1434" pageId="72" pageNumber="73" start="start">propodeal</pageBreakToken>
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hairs moderately long and dense dorsolaterally (0.5-1.5OD); T1-T3 apicolateral patches of tomentum (0.5-1OD); S2 hairs long and dense (1.5OD basolaterad, 0.5OD distilaterad, shorter mesad).
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</paragraph>
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<paragraph id="48064D0C66D2EB092EC6A9D86736DFA8" pageId="73" pageNumber="74">
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<pageBreakToken id="5BECE17B30C730CC212498873A4741FC" pageId="73" pageNumber="74" start="start">Surface</pageBreakToken>
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sculpture: Microsculpture imbricate, integument generally dull; punctures generally small and deep. Clypeus and lower paraocular area moderately densely punctate (i=1-2d) except clypeus irregularly punctate in upper half (i=1-3d); supraclypeal area irregularly punctate (i=1-4d) densely punctate toward lateral margin (i≤d); frontal area very densely punctate (i≤0.5d), densely punctate around ocelli (i≤d); ocellocular space densely punctate adjacent to compound eye (i≤d) becoming impunctate adjacent to lateral ocellus; vertexal area rugose and very densely punctate (i≤0.5d); scape shallowly and moderately densely punctate (i=1-2d) densely punctate apicoventrally (i≤d); genal area microstriate and moderately densely punctate (i=1-2d); hypostomal area weakly imbricate to glossy apically and densely punctate (i=d); pronotum densely punctate (i=0.5-1d), lateral surface coarsely imbricate; mesoscutum densely punctate (i=0.5-1d) except moderately densely punctate laterally (i=1-2d); scutellum moderately densely punctate (i=1-2d) punctures crowded around median (i≤d); mesepisternum irregularly punctate, sparse below scrobe (i=2-3d), sparser anterior of episternal groove (i≥3d), impunctate just dorsad of scrobe, otherwise moderately dense on hypoepimeral area (i=1-2d); metanotum moderately densely punctate (i=1-2d) punctures crowded around median and anteriorly (i≤0.5d); metepisternum densely punctate (i≤d) distinctly longitudinally striate anterodorsally; metapostnotum rugose, fewer transverse lineations posteriorly; propodeum coarsely imbricate, rugulose posteriorly, moderately densely punctate throughout (i=1-2d); T1-T5 coarsely imbricate; T1 sparsely punctate basally (i≥2d) moderately densely punctate apically (i=1-2d); T2-T5 moderately densely punctate (i=1-2d); T6 moderately densely punctate (i=1-2d) and coarsely imbricate posteriorly; T7 sparsely puncticulate; marginal zones of terga weakly imbricate and shiny, punctate to impunctate at margins.
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</paragraph>
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<paragraph id="EE109832833096FD41F6962DE50455A8" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
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Structure: Labrum 2.3
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<normalizedToken id="30DB4EAB4C7B698A96AF6026C7D2431D" originalValue="×">x</normalizedToken>
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wider than long (23:10); malar space subequal in length to clypeal lateral (9:8.5); LOT at level of anterior tentorial pits; clypeus subequal in length to maximum width in frontal view (31:30) extending for more than half of its length beyond LOT; median longitudinal groove present on clypeus, weak apically; subantennal sutures ~1.4
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<normalizedToken id="8803FE3973093CAD59CD0F0B4558D016" originalValue="×">x</normalizedToken>
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as long as the shortest distance between them (17:12); IAD ~2.2
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<normalizedToken id="4C509D35C0BF462DD0A5708EE4EC43AD" originalValue="×">x</normalizedToken>
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AOD (13:6); scape 2.4
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<normalizedToken id="D7E7E4AB4ACE553FB9118D6C74F16F1E" originalValue="×">x</normalizedToken>
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as long as maximum width (24:10); pedicel shorter than wide (8:9); F1 longer than wide (10:8.5); F2 1.3
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<normalizedToken id="0AE12443F52FA31367512B0EE44A25D0" originalValue="×">x</normalizedToken>
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as long as F1 and shorter than F3 (F2:F3 - 13:14); UOD ~1.4
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<normalizedToken id="6E7CABA34C781AF186FE7D7F940B851A" originalValue="×">x</normalizedToken>
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LOD, IOD ~1.1
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<normalizedToken id="CA18852BA9A425FB4244733CAFA9103E" originalValue="×">x</normalizedToken>
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UOD (UOD:IOD:LOD - 50:54:35); frontal line carinate in lower two-fifths, flat just below median ocellus for a length of ~1OD, otherwise roughly defined; MOC subequal to width of head (89:90); OOC ~0.6
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<normalizedToken id="22EAA31D28EB2B95E85FDBED34B0F17E" originalValue="×">x</normalizedToken>
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IOC (11:18); genal area ~0.5
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<normalizedToken id="5F7EE0283C96376B6C5AEAFA92FBF15B" originalValue="×">x</normalizedToken>
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as wide as compound eye in lateral view (17:33); ratio of lengths of mesoscutum: scutellum: metanotum: metapostnotum - 63:23:9:15; probasitarsus 3.25
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<normalizedToken id="07FA7450E4E076255512D7150E75F4F2" originalValue="×">x</normalizedToken>
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as long as maximum depth (26:8); length of prodistitarsus subequal to that of preceding three tarsomeres combined (from II to V - 7:5:5:16); metafemur ~1.9
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<normalizedToken id="DC24BF43D5CE79B1F78597934B57147F" originalValue="×">x</normalizedToken>
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as long as maximum depth (70:37); summit of metatibial ventral convexity rounded and near tibial apex; metatibia ~1.8
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<normalizedToken id="382C890C141A7643145500B7ADEF9B0C" originalValue="×">x</normalizedToken>
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as long as maximum depth (56:31); in apical view apical lamina of metatibia broad and thick, length 0.8
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<normalizedToken id="813D1A2ED216E8E631A2BFDD76EA8033" originalValue="×">x</normalizedToken>
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OD (8:10); ventral metatibial carina bowed ventrad and absent basally, originating at midpoint of weakly sigmoid posterior carina; metabasitarsus ~3.6
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<normalizedToken id="12C9CC1F959A58BF4A19F88BC5222D18" originalValue="×">x</normalizedToken>
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as long as maximum depth (43:12); S1 process apical surface with longitudinal median
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<pageBreakToken id="6BD9C027D69BA563771654758E7929B4" pageId="74" pageNumber="75" start="start">ridge</pageBreakToken>
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, in lateral view anterior and posterior margins convergent apically and median ridge projecting posteriorly from apex. S7 apodemal arm sclerotized margin terminating laterally at arm midlength; ventral lobe narrow, almost linear; dorsal lobe roughly triangular, wide at base and narrow apically, with row of setae long basally and shorter apically; apex of disc disinctly emarginate. S8 lateral process ~0.9
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<normalizedToken id="65A9F32BEFF023EEAA4E8360E5EC9D75" originalValue="×">x</normalizedToken>
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as wide as long, anteromedial sclerotized margin following marginal contour. Gonobase apicoventral truncate process biconvex and distinctly notched by ~1.5
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<normalizedToken id="69F142B750CC891415570FEE9CF32DD8" originalValue="×">x</normalizedToken>
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width of one convexity.
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</paragraph>
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<paragraph id="92C39B6A25BE43C0FA71A7585951BCAC" pageId="74" pageNumber="75">Female. Length 4.2-4.8mm, forewing length 2.6-3.0mm, head width 1.0mm, thorax width 1.0-1.1mm, median ocellar diameter (OD) 0.10mm.</paragraph>
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<paragraph id="E7225B47CB41F92D577C5AD43288DCBA" pageId="74" pageNumber="75">Colouration: Black to black-brown except as follows: mandible with yellow basal spot, translucent yellow in apical half except apex translucent brown; antenna variable (scape entirely dark; pedicel to F2 dark, or apicoventral rim brown; ventral surface of F3 to penultimate flagellomere yellow-brown, narrowing basally on F3, flagellomeres suffused with brown basally; terminal flagellomere yellow-brown in basal half, brown in apical half). Following parts yellow: dorsal surface of protibia; apical two-thirds of prodistitarsus, sometimes yellow-brown; narrow basal ring on mesotibia; apicodorsal rim on metafemur; wide basal ring on metatibia, narrow ventrally; anterior spot on tegula. Following parts yellow-brown: apicodorsal spot on mesotibia; posterior apical rim on metatibia. Apical half of mesodistitarsus brown. Metasoma black, T1-T5 yellow-brown beyond premarginal line to translucent yellow at margins.</paragraph>
|
||
<paragraph id="8E99AFD1BBA0112F00F72489F8A87DE1" pageId="74" pageNumber="75">Pubescence: As in male except as follows: clypeal hairs short and sparse (0.5OD); genal beard sparse (0.5-2.5OD); discs of mesoscutum and scutellum with sparse short hairs (≤0.5OD), longer and denser toward lateral margin of mesoscutum (0.5-1OD), and toward lateral and posterior margins of scutellum (1-1.5OD) longest posteromedially; scopae on metafemur and metatibia (1-1.5OD); T1-T3 apicolateral patches of tomentum (0.5-1OD); S1 hairs long and moderately dense (≤2OD); S2 scopal hairs (1-2OD).</paragraph>
|
||
<paragraph id="07648BD023FE575C17F572362485575B" pageId="74" pageNumber="75">Surface sculpture: As in male except as follows: clypeus sparsely punctate (i=2-3d) moderately densely punctate toward lateral margin (i=1-2d); supraclypeal area irregularly punctate (i=1-2d) densely punctate toward lateral margin (i≤d); lower paraocular area densely punctate (i=d); frontal area densely punctate (i≤d), moderately densely punctate around ocelli (i=1-2d); vertexal area moderately densely punctate (i=1-2d); scape shallowly and moderately densely punctate (i=1-2d); genal area weakly imbricate and moderately densely punctate (i=1-2d); hypostomal area weakly imbricate to glossy apically and moderately densely punctate (i=1-2d); mesoscutum and scutellum densely punctate; metanotum moderately densely punctate (i=0.5-2d); metepisternum densely punctate (i≤d) distinctly longitudinally striate anterodorsally (weak in some specimens); metapostnotum rugose, fewer transverse lineations posteriorly (posterior half to two-thirds longitudinally carinate in some specimens); propodeum moderately densely punctate (i=1-2d); T1-T5 moderately puncticulate; T6 moderately densely punctate (i=1-2d).</paragraph>
|
||
<paragraph id="7558FE0EB70A5AB13D1E161C5C7DD923" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75">
|
||
Structure: Labrum 2.5
|
||
<normalizedToken id="EABD03E53CCDA7C78DA5DAE63EDE904C" originalValue="×">x</normalizedToken>
|
||
wider than long (20:8); malar space subequal in length to clypeal lateral (6:6.5); LOT at level of anterior tentorial pits; clypeus ~1.1
|
||
<normalizedToken id="8A2F9D537427FD164DA002E2F0DEADCA" originalValue="×">x</normalizedToken>
|
||
as long as maximum width in frontal view (29:26) extending for approximately half of its length
|
||
<pageBreakToken id="7DB3411230E16989854212479B3FCA97" pageId="75" pageNumber="76" start="start">beyond</pageBreakToken>
|
||
LOT; median longitudinal groove weak on clypeus; subantennal sutures subequal in length to the shortest distance between them (11:11); IAD ~1.4
|
||
<normalizedToken id="724FFB5C40C43444F5C9DCADE8EF3A67" originalValue="×">x</normalizedToken>
|
||
AOD (11.5:8); scape ~3.4
|
||
<normalizedToken id="08546DFF5A4F78A2B76A68D3DD7C623D" originalValue="×">x</normalizedToken>
|
||
as long as maximum width (22:6.5); pedicel longer than wide (8:6.5); F1 longer than wide (6.5:5.5); F2 shorter than F1 and shorter than F3 (F2:F3 - 4:5); UOD ~1.3
|
||
<normalizedToken id="95078324A4D36A9C25C298181E142D0D" originalValue="×">x</normalizedToken>
|
||
LOD, IOD ~1.1
|
||
<normalizedToken id="501F0EB6D0E6AED73F9F30AD93FF6991" originalValue="×">x</normalizedToken>
|
||
as long as UOD (UOD:IOD:LOD - 44:47:34); frontal line carinate in lower third, flat above; MOC ~1.1
|
||
<normalizedToken id="2E59BD2DA6DFC9FD503FA6474A4E55F7" originalValue="×">x</normalizedToken>
|
||
as long as width of head (77:71); OOC ~0.7
|
||
<normalizedToken id="AB539173C60FBA2300696616667B967A" originalValue="×">x</normalizedToken>
|
||
IOC (11:16); genal area ~0.65
|
||
<normalizedToken id="0836EF35E0FAFED730D857AABA824B24" originalValue="×">x</normalizedToken>
|
||
as wide as eye in lateral view (15:23); ratio of lengths of mesoscutum: scutellum: metanotum: metapostnotum - 51:20:9:12.5; probasitarsus ~3.2
|
||
<normalizedToken id="F31C162F909A0A58546191A8D7EB20A1" originalValue="×">x</normalizedToken>
|
||
as long as maximum depth (21:6.5); length of prodistitarsus ~1.3
|
||
<normalizedToken id="A97FE8B352AB411C83E94D5120187849" originalValue="×">x</normalizedToken>
|
||
that of preceding three tarsomeres combined (from II to V - 4:4:3:14); metafemur ~2.9
|
||
<normalizedToken id="4F29D2A35EC652F4F238253A048C4742" originalValue="×">x</normalizedToken>
|
||
as long as maximum depth (43:15); metatibia ~4.6
|
||
<normalizedToken id="39366DA05C7CC6113DF27301F89C46CC" originalValue="×">x</normalizedToken>
|
||
as long as maximum depth (53:11.5); metabasitarsus 4.25
|
||
<normalizedToken id="862C2E10294E1DE4D6991E9298A4F46C" originalValue="×">x</normalizedToken>
|
||
as long as maximum depth (34:8).
|
||
</paragraph>
|
||
<caption id="EAA66234DAF4B734EA2603B7D3172345" pageId="75" pageNumber="76">
|
||
<paragraph id="8FAA211C18F55C3AD099EE7036B7EEE0" pageId="75" pageNumber="76">
|
||
Figure 41.
|
||
<taxonomicName id="2798E8A447C722367CCE6B6F908B4D4C" class="Insecta" family="Colletidae" genus="Chilicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chilicola mavida" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="mavida">Chilicola mavida</taxonomicName>
|
||
: A male habitus, lateral view, scale bar 1 mm B male head, frontal view, scale bar 0.5 mm C female habitus, lateral view, scale bar 1 mm D female head, frontal view, scale bar 0.5 mm E male metatibia, posterior view, scale bar 0.25 mm F male S1 process, lateral view†, scale bar 0.25 mm G male S1 process, apical view, scale bar 0.25 mm H male S7, scale bar 0.25 mm I male S8, scale bar 0.25 mm J male genital capsule, scale bar 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="490EBCBA79F29DE7436EE238F37B0136" pageId="75" pageNumber="76" type="materials_examined">
|
||
<paragraph id="C05B9BB6E4511D715F584B8779E13CB5" pageId="75" pageNumber="76">Material studied</paragraph>
|
||
<paragraph id="95F9D78DFF9EBB1A097E0911E4B96C3E" pageId="75" pageNumber="76">
|
||
(21 males & 19 females): Holotype (male): Region IV,
|
||
<normalizedToken id="C9F9E0277C0045AD13D5AD3EEAC45DA3" originalValue="Baños">Banos</normalizedToken>
|
||
del Toro,
|
||
<geoCoordinate id="D97A5A6CEC31177A15185843FF32AC17" direction="south" orientation="latitude" precision="55" value="-29.838">S 29.838°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="50F487637FFE306511E35ED739740A37" direction="west" orientation="longitude" precision="55" value="-70.026">W 70.026 °</geoCoordinate>
|
||
, 3202m, i.1970, Toro (AMNH); Allotype (female): same data as holotype (AMNH); Region IV: one paratype male, same data as holotype (AMNH); five paratype males, Laguna Dam,
|
||
<geoCoordinate id="7E23138686652D65BDD801A431725CED" direction="south" orientation="latitude" precision="55" value="-30.206">S 30.206°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EECF89878C63C81456F472C251A622A3" direction="west" orientation="longitude" precision="55" value="-70.042">W 70.042°</geoCoordinate>
|
||
, 3170m, 5.xii.1950, Ross & Michelbacher (AMNH); one paratype male,
|
||
<normalizedToken id="F6B8B94F4CF7F1A6DDF18DBC811B92D3" originalValue="Río">Rio</normalizedToken>
|
||
Laguna,
|
||
<geoCoordinate id="A171A2A8C6E37566DA6FB597C3E68A90" direction="south" orientation="latitude" precision="55" value="-30.191">S 30.191°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8DD8D4B0B53C079DE5A6F3DC6E47FCE0" direction="west" orientation="longitude" precision="55" value="-70.047">W 70.047°</geoCoordinate>
|
||
, 3000m, i.1970, Dazarola (AMNH); one female, same locality as holotype, 1970, collector unknown (AMNH); one male,
|
||
<normalizedToken id="3B34DF014C9AE7D6C0A8A20DE68BD241" originalValue="Río">Rio</normalizedToken>
|
||
Laguna,
|
||
<geoCoordinate id="E4937E3FA2B3F0DCE24359812B8033FC" direction="south" orientation="latitude" precision="55" value="-30.191">S 30.191°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3E231285D0099766D6B3F87B27D577F2" direction="west" orientation="longitude" precision="55" value="-70.047">W 70.047°</geoCoordinate>
|
||
, 3000m, 1970, H. Toro (AMNH); one male, same locality, 8.i.1970, H. Toro (PUCV); one male, CH-41 km 165, S of La Laguna,
|
||
<geoCoordinate id="32E8E5624E006C7873D6756F8B88CD0B" direction="south" orientation="latitude" precision="5" value="-30.1479">S 30.1479°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="14DD78E3D1ECB6CD38FB7E49DBC22679" direction="west" orientation="longitude" precision="5" value="-70.0518">W 70.0518°</geoCoordinate>
|
||
, 2767m, 15.xii.2013, L. Packer (PCYU); one male, 41-CH km km 171, N of Emb La Laguna,
|
||
<geoCoordinate id="C40E6D2C0D4941C3C7194BB3BB13EE34" direction="south" orientation="latitude" precision="5" value="-30.1787">S 30.1787°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D6DC73BD51516226BC54AE6BF26172F4" direction="west" orientation="longitude" precision="5" value="-70.0473">W 70.0473°</geoCoordinate>
|
||
, 2942m, 15.xii.2013, J. Postlethwaite, CCDB-19989 D05 (PCYU); two females, W of Junta Toro, 41-CH km 146,
|
||
<geoCoordinate id="8EF9E7238797019CB0BD698AEB700D4F" direction="south" orientation="latitude" precision="5" value="-29.9675">S 29.9675°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5433E803C8A8A1BA2D5EDA9F05DB0D02" direction="west" orientation="longitude" precision="5" value="-70.1285">W 70.1285°</geoCoordinate>
|
||
, 1977m, 8.xi.2013, S. Monckton, pan traps, CCDB-19989 D01 // PCYU 0021694, CCDB-19989 D02 // PCYU 0021695 (PCYU); one male, 41-CH km, 137.8,
|
||
<geoCoordinate id="AE00209719AD87BAB68430BC6F9FBF54" direction="south" orientation="latitude" precision="5" value="-29.967">S 29.9670°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F4C68F7F1A1381D14252B3606A211FA6" direction="west" orientation="longitude" precision="5" value="-70.1936">W 70.1936</geoCoordinate>
|
||
, 800m, 8.xi.2013, S. Monckton, pan trap, CCDB-19989 B10 // PCYU 0021684 (PCYU); one male, E of
|
||
<normalizedToken id="D03683F2F0B36186812E4840FEFBBC3D" originalValue="Vicuña">Vicuna</normalizedToken>
|
||
, 127.8 km,
|
||
<geoCoordinate id="33F4FB05AA38F731B495DA4124BD0321" direction="south" orientation="latitude" precision="5" value="-29.9777">S 29.9777°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F5D5EDB9D41833AC38D9370795DBD089" direction="west" orientation="longitude" precision="5" value="-70.2293">W 70.2293°</geoCoordinate>
|
||
, 1711m, 12-19.ix.2010, L. Packer (PCYU); one female, Elqui Prov., Pangue,
|
||
<geoCoordinate id="F22E5D7A7F031FF493CC71DDFB7C1538" direction="south" orientation="latitude" precision="5" value="-30.1539">S 30.1539°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D8975DDCB754D78CC4D9D9EFEC5E8838" direction="west" orientation="longitude" precision="5" value="-70.6639">W 70.6639°</geoCoordinate>
|
||
, 1686m, 11-30.ix.2004, A. Ugarte (PCYU); two males, Elqui: El Pangue, 24 km S of
|
||
<normalizedToken id="ED90FCB3E1D4F65B9087631F4424216E" originalValue="Vicuña">Vicuna</normalizedToken>
|
||
,
|
||
<geoCoordinate id="AFA37EF5932BCD52910F4AC822914C85" direction="south" orientation="latitude" precision="55" value="-30.168">S 30.168°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="81C4B991FD2E6FB2D0486796689D836C" direction="west" orientation="longitude" precision="55" value="-70.663">W 70.663°</geoCoordinate>
|
||
, 1711m, 31.x.1992, Snyder & Sharkov (AMNH); one male and one female, Elqui Prov., 26 km S of
|
||
<normalizedToken id="5FF46CAAD1CBC216F31323C107DC3C43" originalValue="Vicuña">Vicuna</normalizedToken>
|
||
,
|
||
<geoCoordinate id="50F509C5EE4C7D7E959B6D4E5227801B" direction="south" orientation="latitude" precision="555" value="-30.19">S 30.19°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8A2C18E8DF867C628C031CBBA9192D46" direction="west" orientation="longitude" precision="55" value="-70.661">W 70.661°</geoCoordinate>
|
||
, 1691m, 29.x.1992, Rozen, Sharkov, Snyder (AMNH); three females,
|
||
<normalizedToken id="7887EC720409B5F6766E94F724EA6687" originalValue="Limarí">Limari</normalizedToken>
|
||
Prov., Las Mollacas, E Monte Patria,
|
||
<geoCoordinate id="718797F17CB8020C205B7D6DB85275FB" direction="south" orientation="latitude" precision="55" value="-30.753">S 30.753°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="40482B37E16AC0E451967A89DCD5C295" direction="west" orientation="longitude" precision="55" value="-70.657">W 70.657°</geoCoordinate>
|
||
, 1142m, 13.x.1994, Rozen, Quinter, Ascher, on
|
||
<taxonomicName id="EFE86BB1CBE593D018BA4751002C5D55" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oxalis" order="Oxalidales" pageId="75" pageNumber="76" phylum="Tracheophyta" rank="genus">Oxalis</taxonomicName>
|
||
(AMNH); Region V: one female, Portillo Aduanas,
|
||
<geoCoordinate id="ED8D0897E8A5897840399A800680DAD4" direction="south" orientation="latitude" precision="5" value="-32.8304">S 32.8304°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="DACCA8FC58C6B184D61B52AF1145BE18" direction="west" orientation="longitude" precision="5" value="-70.0905">W 70.0905°</geoCoordinate>
|
||
, 3210m, 10.i.2009, L. Packer, on
|
||
<taxonomicName id="F8EB33619B2F2841E27352ED14010F24" class="Magnoliopsida" family="Hydrophyllaceae" genus="Phacelia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Phacelia" order="Boraginales" pageId="75" pageNumber="76" phylum="Tracheophyta" rank="genus">Phacelia</taxonomicName>
|
||
, CHI09-13-2-011 (PCYU); two males and six females, Portillo,
|
||
<geoCoordinate id="DD1C2E4CF46191A31F6CB7D81FEA42C4" direction="south" orientation="latitude" precision="5" value="-32.8372">S 32.8372°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5C365EBCB9180A9B2DFF8C8961478DBE" direction="west" orientation="longitude" precision="5" value="-70.1296">W 70.1296°</geoCoordinate>
|
||
, 2870m, 11.i.2009, L. Packer, on
|
||
<taxonomicName id="6B05A637A36A7C386F247B6265629429" class="Magnoliopsida" family="Hydrophyllaceae" genus="Phacelia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Phacelia" order="Boraginales" pageId="75" pageNumber="76" phylum="Tracheophyta" rank="genus">Phacelia</taxonomicName>
|
||
, CHI09-13-1-003, CHI09-13-1-005, CHI09-13-1-004, CHI09-13-1-009 (PCYU); two females, same data (BBSL); two females, same data (CTMI); one female, same locality, 30.xii.2008, L. Packer, CHI09-2-1-018 (PCYU); one female, Portillo,
|
||
<geoCoordinate id="80415682D585435A9FD0EE62E90BB31B" direction="south" orientation="latitude" precision="5" value="-32.8507">S 32.8507°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A76450EE1F1E108921DD2340850A10E8" direction="west" orientation="longitude" precision="5" value="-70.1339">W 70.1339°</geoCoordinate>
|
||
, 2612m, 30.xii.2009, L. Packer, CHI09-2-2-031 (PCYU); one male and one female, same data, on
|
||
<taxonomicName id="3AACC576C9FEAE6110366018CF2DABF2" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stachys" order="Lamiales" pageId="75" pageNumber="76" phylum="Tracheophyta" rank="genus">Stachys</taxonomicName>
|
||
, CHI09-2-2-039, CHI09-2-2-0398 (PCYU); one male, Guardia Vieja mountain, Aconcagua Prov.,
|
||
<geoCoordinate id="9C1229E81124B6D28834474AB809351D" direction="south" orientation="latitude" precision="55" value="-32.904">S 32.904°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="452578D29D8FF8BC544DDCEA430D5FAE" direction="west" orientation="longitude" precision="55" value="-70.272">W 70.272°</geoCoordinate>
|
||
, 1600m, 17.ix.1964, L. E.
|
||
<normalizedToken id="4F3D4AA6111A46BA4E8829A6C7599C16" originalValue="Peña">Pena</normalizedToken>
|
||
(AMNH).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1C7955EC074CA5B9718E2F7000E2C532" pageId="76" pageNumber="77" type="variation">
|
||
<paragraph id="682ACA0AF8B7C0AC22B102B959D177CA" pageId="76" pageNumber="77">
|
||
<pageBreakToken id="2DDE51002D763F9BE5C1B590CF02C46C" pageId="76" pageNumber="77" start="start">Variation</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="84EE8B525E36A6A969792DBD62ADD1C9" pageId="76" pageNumber="77">Some females have a yellow-brown or yellow spot on the lower paraocular area below the anterior tentorial pit.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3B802191F76CAA90896C2F5501C1D81C" pageId="76" pageNumber="77" type="distribution">
|
||
<paragraph id="1E2AA94E6AD79992CDEFE8B80FC85052" pageId="76" pageNumber="77">Distribution.</paragraph>
|
||
<paragraph id="B663E87120032512F73EC57AEE4E744C" pageId="76" pageNumber="77">
|
||
Central Andean Cordillera and upper Coquimban Desert, just extending into Southern Andean Cordillera, from
|
||
<normalizedToken id="90F6C802AE1DE8507307CF0435535BBF" originalValue="Baños">Banos</normalizedToken>
|
||
del Toro & Embalse La Laguna west to El Pangue (Region IV) and south to Portillo (Region V); 1142-3210m a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8AAD7EACCCEFCCBECF6093068C4A152" pageId="76" pageNumber="77" type="ecology">
|
||
<paragraph id="9B7D06D83EF6ECAFAAC1F31D2DF36DA0" pageId="76" pageNumber="77">Ecology.</paragraph>
|
||
<paragraph id="10103F04DFA86E6B03546BC6BFEF8217" pageId="76" pageNumber="77">
|
||
Collected on
|
||
<taxonomicName id="55C68BC77A15DF8B52AB7D668385891B" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oxalis" order="Oxalidales" pageId="76" pageNumber="77" phylum="Tracheophyta" rank="genus">Oxalis</taxonomicName>
|
||
L. (3 records),
|
||
<taxonomicName id="1146FA66B77706DFB437E42461D114DF" class="Magnoliopsida" family="Hydrophyllaceae" genus="Phacelia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Phacelia" order="Boraginales" pageId="76" pageNumber="77" phylum="Tracheophyta" rank="genus">Phacelia</taxonomicName>
|
||
(9 records), and
|
||
<taxonomicName id="FF2FE968D0AF8A809F6DCB74931609F2" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stachys" order="Lamiales" pageId="76" pageNumber="77" phylum="Tracheophyta" rank="genus">Stachys</taxonomicName>
|
||
L. (2 records). Recorded September to January.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |