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<document id="AAF3E9E7C14BD86F1E03B17F02130C63" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D7B9233FDBCFE0AC9102AD4" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius seriegranosus Heller 1914" docType="treatment" docVersion="10" lastPageNumber="105" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="104" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F36757F4EFE4A5B03AB1133902CF8C9B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="72A843C4D784A8808E3FC2007393F045">
<mods:title id="824D3706E4247FCCB1B8480E851AFDD8">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="31241942F334FE40F7025009B5493954" type="personal">
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<mods:roleTerm id="51814AC51D8478A5839FB6AD3B84C648">Author</mods:roleTerm>
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<mods:namePart id="098B4183D46DFD206B0849823B8C8AD1">Telnov, Dmitry</mods:namePart>
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<mods:title id="8A8CB0856BDF8F237565BE37DFDCEC38">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="9E4E3521D48521849AD19A0BBDE415A5">
<mods:date id="30043936CFA0CE42F1D97ADD65493243">2020</mods:date>
<mods:detail id="BB08578CE21F76566F6BC72E9734B80D" type="pubDate">
<mods:number id="054185F4F93906AB105E45C4E661B20B">2020-10-19</mods:number>
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<mods:number id="FB7B861855D3C5776851497808FC7648">721</mods:number>
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<mods:classification id="754F17B1DC9BEBA85DC1CBFF2A787E46">journal article</mods:classification>
<mods:identifier id="D0013BC29EDC69909B74755BB33B225B" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="4DB2AE24CC1FB5C208B0F841092CA633" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="B828B8C7EACFE268BA4A681EC683FC3D" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="D6276331CAF3DCF0FD2180C5AB6D275E" type="Zenodo-Dep">4122118</mods:identifier>
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</mods:mods>
<treatment id="1A3787F69D7B9233FDBCFE0AC9102AD4" ID-DOI="http://doi.org/10.5281/zenodo.4344479" ID-GBIF-Taxon="168648116" ID-Zenodo-Dep="4344479" LSID="urn:lsid:plazi:treatment:1A3787F69D7B9233FDBCFE0AC9102AD4" httpUri="http://treatment.plazi.org/id/1A3787F69D7B9233FDBCFE0AC9102AD4" lastPageId="104" lastPageNumber="105" pageId="103" pageNumber="104">
<subSubSection id="DA84656B9D7B923CFDBCFE0ACC5329AD" box="[559,1029,436,463]" pageId="103" pageNumber="104" type="nomenclature">
<paragraph id="922136E09D7B923CFDBCFE0ACC5329AD" blockId="103.[515,1072,436,499]" box="[559,1029,436,463]" pageId="103" pageNumber="104">
<heading id="C969818C9D7B923CFDBCFE0ACC5329AD" box="[559,1029,436,463]" centered="true" fontSize="11" level="2" pageId="103" pageNumber="104" reason="2">
<taxonomicName id="559E4D639D7B923CFDBCFE0ACC5329AD" ID-CoL="7X3XL" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[559,1029,436,463]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="seriegranosus">
<emphasis id="A0EAEAF29D7B923CFDBCFE0ACB3A29AC" bold="true" box="[559,876,436,463]" italics="true" pageId="103" pageNumber="104">Maechidius seriegranosus</emphasis>
<bibRefCitation id="F60F4B119D7B923CFCE0FE0ACC5329AD" author="Heller K. M." box="[883,1029,436,463]" pageId="103" pageNumber="104" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7B923CFD90FE66CC662990" box="[515,1072,472,499]" pageId="103" pageNumber="104" type="description">
<paragraph id="922136E09D7B923CFD90FE66CC662990" blockId="103.[515,1072,436,499]" box="[515,1072,472,499]" pageId="103" pageNumber="104">
<figureCitation id="0AA52A659D7B923CFD90FE66CA0B2991" box="[515,605,472,499]" captionStart="Figs 78–81" captionStartId="100.[189,241,1928,1954]" captionTargetBox="[341,1237,264,1897]" captionTargetId="figure-63@100.[336,1259,251,1916]" captionTargetPageId="100" captionText="Figs 78–81. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 78. M. riedeli sp. nov., holotype, ♂ (SMNS). 79. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 80. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 81. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122163" httpUri="https://zenodo.org/record/4122163/files/figure.png" pageId="103" pageNumber="104">Figs 80</figureCitation>
,
<figureCitation id="0AA52A659D7B923CFDFAFE67CAC12991" box="[617,663,473,499]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="103" pageNumber="104">171</figureCitation>
,
<figureCitation id="0AA52A659D7B923CFD37FE66CA852991" box="[676,723,472,499]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="103" pageNumber="104">250</figureCitation>
,
<figureCitation id="0AA52A659D7B923CFD4CFE67CB582991" box="[735,782,473,499]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="103" pageNumber="104">341</figureCitation>
, 413, 515, 560, 706–708
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7B923CFF2EFDA5CB6D2BEB" pageId="103" pageNumber="104" type="materials_examined">
<paragraph id="922136E09D7B923CFF2EFDA5C93A2A56" blockId="103.[189,364,538,565]" box="[189,364,538,565]" pageId="103" pageNumber="104">
<heading id="C969818C9D7B923CFF2EFDA5C93A2A56" bold="true" box="[189,364,538,565]" fontSize="11" level="3" pageId="103" pageNumber="104" reason="3">
<emphasis id="A0EAEAF29D7B923CFF2EFDA5C93A2A56" bold="true" box="[189,364,538,565]" pageId="103" pageNumber="104">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7B923CFF79FDF4CA3A2A06" blockId="103.[189,1399,585,684]" box="[234,620,585,612]" pageId="103" pageNumber="104">
<emphasis id="A0EAEAF29D7B923CFF79FDF4C9302A06" bold="true" box="[234,358,586,612]" pageId="103" pageNumber="104">
<typeStatus id="4D2588429D7B923CFF79FDF4C9302A06" box="[234,358,586,612]" pageId="103" pageNumber="104" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D7B923CFF2EFDD0CD032ACE" blockId="103.[189,1399,585,684]" pageId="103" pageNumber="104">
<materialsCitation id="22F63CBD9D7B923CFF2EFDD0CD032ACE" ID-GBIF-Occurrence="2898878334" collectionCode="SNSD" collectorName="Br. N. Guinea Aroa Fl. E. Weiske" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="103" pageNumber="104" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="EA8976709D7B923CFF2EFDD0C9B02AEA" box="[189,486,622,648]" name="Papua New Guinea" pageId="103" pageNumber="104">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D7B923CFD9FFDD0CA722AEA" box="[524,548,622,648]" pageId="103" pageNumber="104">♂</specimenCount>
; “
<collectorName id="3F6B53369D7B923CFDD7FDD0CBBE2AEA" box="[580,1000,622,648]" pageId="103" pageNumber="104">Br. N. Guinea Aroa Fl. E.Weiske</collectorName>
[p, blue label] // 14420 [h, blue label]//
<typeStatus id="4D2588429D7B923CFE84FD2CC9332ACE" box="[279,357,658,684]" pageId="103" pageNumber="104">Typus</typeStatus>
! [p]
<taxonomicName id="559E4D639D7B923CFE0AFD2DCA682ACE" authorityName="Heller" authorityYear="1914" box="[409,574,659,684]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="seriegranosus">
<emphasis id="A0EAEAF29D7B923CFE0AFD2DCA682ACE" box="[409,574,659,684]" italics="true" pageId="103" pageNumber="104">seriegranosus</emphasis>
</taxonomicName>
[h, red label]// Staatl. Museum für Tierkunde.Dresden [p]”;
<collectionCode id="F48FAE259D7B923CFA91FD2CCD072ACE" box="[1282,1361,658,684]" collectionName="SNSD" pageId="103" pageNumber="104">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D7B923CFF2EFD6AC9302A8C" blockId="103.[189,358,724,750]" box="[189,358,724,750]" pageId="103" pageNumber="104">
<heading id="C969818C9D7B923CFF2EFD6AC9302A8C" bold="true" box="[189,358,724,750]" fontSize="11" level="3" pageId="103" pageNumber="104" reason="3">
<emphasis id="A0EAEAF29D7B923CFF2EFD6AC9302A8C" bold="true" box="[189,358,724,750]" pageId="103" pageNumber="104">New material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7B923CFF2EFCBDCB6D2BEB" blockId="103.[189,1399,771,905]" pageId="103" pageNumber="104">
<materialsCitation id="22F63CBD9D7B923CFF2EFCBDC9432B20" ID-GBIF-Occurrence="2898878563" collectingDate="1933-10" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Papua New Guinea" elevation="366" location="Papua" pageId="103" pageNumber="104" specimenCount="1">
<collectingCountry id="EA8976709D7B923CFF2EFCBDC98D2B7F" box="[189,475,771,798]" name="Papua New Guinea" pageId="103" pageNumber="104">PAPUA NEW GUINEA</collectingCountry>
• 2 specs; “
<location id="9741603B9D7B923CFDF5FCBDCAE32B7F" LSID="urn:lsid:plazi:treatment:1A3787F69D7B9233FDBCFE0AC9102AD4:9741603B9D7B923CFDF5FCBDCAE32B7F" box="[614,693,771,797]" country="Papua New Guinea" name="Papua" pageId="103" pageNumber="104">Papua</location>
: Kokoda.
<quantity id="55669B059D7B923CFCBEFCBDCBDF2B7F" box="[813,905,771,797]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="103" pageNumber="104" unit="ft" value="1200.0">
<elevation id="19B3D1D39D7B923CFCBEFCBDCBDF2B7F" box="[813,905,771,797]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="103" pageNumber="104" unit="ft" value="1200.0">1,200ft.</elevation>
</quantity>
<date id="E62010209D7B923CFC02FCBDCBB32B7F" box="[913,997,771,797]" pageId="103" pageNumber="104" value="1933-10">
<collectingDate id="F664E9C89D7B923CFC02FCBDCBB32B7F" box="[913,997,771,797]" pageId="103" pageNumber="104" value="1933-10">x.1933</collectingDate>
</date>
.
<collectorName id="3F6B53369D7B923CFC62FCBDCCF32B7F" box="[1009,1189,771,797]" pageId="103" pageNumber="104">L.E.Cheesman.</collectorName>
B.M.1934-321.”;
<collectionCode id="F48FAE259D7B923CFF2EFC96C9432B20" box="[189,277,808,834]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="103" pageNumber="104">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D7B923CFEB8FC99CD202B20" ID-GBIF-Occurrence="2898878373" box="[299,1398,806,834]" collectingDate="1934-02" collectionCode="PAPUA, BMNH" collectorName="L. E. Cheesman." country="Papua New Guinea" elevation="1524" pageId="103" pageNumber="104" specimenCount="1">
23 specs; “
<collectionCode id="F48FAE259D7B923CFE34FC99CA5B2B23" box="[423,525,807,833]" pageId="103" pageNumber="104">PAPUA</collectionCode>
:Mondo.
<quantity id="55669B059D7B923CFDE2FC99CA9B2B20" box="[625,717,807,834]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="103" pageNumber="104" unit="ft" value="5000.0">
<elevation id="19B3D1D39D7B923CFDE2FC99CA9B2B20" box="[625,717,807,834]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="103" pageNumber="104" unit="ft" value="5000.0">5,000ft.</elevation>
</quantity>
<date id="E62010209D7B923CFD40FC99CB7E2B23" box="[723,808,807,833]" pageId="103" pageNumber="104" value="1934-02">
<collectingDate id="F664E9C89D7B923CFD40FC99CB7E2B23" box="[723,808,807,833]" pageId="103" pageNumber="104" value="1934-02">ii.1934</collectingDate>
</date>
.
<collectorName id="3F6B53369D7B923CFCA1FC99CBB12B23" box="[818,999,807,833]" pageId="103" pageNumber="104">L.E.Cheesman.</collectorName>
B.M.1933-321 [&amp;] 492.”;
<collectionCode id="F48FAE259D7B923CFA8DFC96CD202B20" box="[1310,1398,808,834]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="103" pageNumber="104">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D7B923CFF45FCF5CB612BEB" ID-GBIF-Occurrence="2898878474" collectingDate="1934-02" collectionCode="PAPUA, BMNH" collectorName="L. E. Cheesman." country="Papua New Guinea" elevation="1524" pageId="103" pageNumber="104" specimenCount="1" specimenCount-female="1">
<specimenCount id="8498FD699D7B923CFF45FCF5C95D2B04" box="[214,267,843,870]" pageId="103" pageNumber="104" type="female">1 ♀</specimenCount>
; “
<collectionCode id="F48FAE259D7B923CFEB8FCF5C9C72B07" box="[299,401,843,869]" pageId="103" pageNumber="104">PAPUA</collectionCode>
:Mondo.
<quantity id="55669B059D7B923CFE6DFCF5CA0C2B04" box="[510,602,843,870]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="103" pageNumber="104" unit="ft" value="5000.0">
<elevation id="19B3D1D39D7B923CFE6DFCF5CA0C2B04" box="[510,602,843,870]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="103" pageNumber="104" unit="ft" value="5000.0">5,000ft.</elevation>
</quantity>
<date id="E62010209D7B923CFDFBFCF5CAE82B07" box="[616,702,843,869]" pageId="103" pageNumber="104" value="1934-02">
<collectingDate id="F664E9C89D7B923CFDFBFCF5CAE82B07" box="[616,702,843,869]" pageId="103" pageNumber="104" value="1934-02">ii.1934</collectingDate>
</date>
.
<collectorName id="3F6B53369D7B923CFD42FCF5CBD32B07" box="[721,901,843,869]" pageId="103" pageNumber="104">L.E.Cheesman.</collectorName>
B.M.1933-321.//
<taxonomicName id="559E4D639D7B923CFBCCFCF5C9CF2BEB" authority="Moser Determined" authorityName="Moser Determined" authorityYear="1920" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="fraterculus">
<emphasis id="A0EAEAF29D7B923CFBCCFCF5CD212B07" box="[1119,1399,843,869]" italics="true" pageId="103" pageNumber="104">Maechidius fraterculus</emphasis>
Moser Determined
</taxonomicName>
from description. G.J.A.”;
<collectionCode id="F48FAE259D7B923CFD48FCD1CB612BEB" box="[731,823,879,905]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="103" pageNumber="104">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7B923CFF2EFC0FCAE12C04" pageId="103" pageNumber="104" type="discussion">
<paragraph id="922136E09D7B923CFF2EFC0FC9792BA9" blockId="103.[189,303,945,971]" box="[189,303,945,971]" pageId="103" pageNumber="104">
<heading id="C969818C9D7B923CFF2EFC0FC9792BA9" bold="true" box="[189,303,945,971]" fontSize="11" level="3" pageId="103" pageNumber="104" reason="3">
<emphasis id="A0EAEAF29D7B923CFF2EFC0FC9792BA9" bold="true" box="[189,303,945,971]" pageId="103" pageNumber="104">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7B923CFF2EFC5ECAE12C04" blockId="103.[189,1399,992,1127]" pageId="103" pageNumber="104">
<bibRefCitation id="F60F4B119D7B923CFF2EFC5EC9C02B99" author="Heller K. M." box="[189,406,992,1019]" pageId="103" pageNumber="104" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller (1914: 628)</bibRefCitation>
likely based his description of
<taxonomicName id="559E4D639D7B923CFC91FC5FCB9B2B98" authorityName="Heller" authorityYear="1914" box="[770,973,993,1018]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="seriegranosus">
<emphasis id="A0EAEAF29D7B923CFC91FC5FCB9B2B98" box="[770,973,993,1018]" italics="true" pageId="103" pageNumber="104">M. seriegranosus</emphasis>
</taxonomicName>
on a single specimen – although not explicitly stated. The single SNSD specimen labelled “Typus♂” is herewith designated as
<typeStatus id="4D2588429D7B923CFB78FBBACD012C7C" box="[1259,1367,1028,1054]" pageId="103" pageNumber="104" type="lectotype">lectotype</typeStatus>
in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other
<typeStatus id="4D2588429D7B923CFE92FBF3C93E2C05" box="[257,360,1101,1127]" pageId="103" pageNumber="104" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7B9233FF2EFB30C9102AD4" lastPageId="104" lastPageNumber="105" pageId="103" pageNumber="104" type="description">
<paragraph id="922136E09D7B923CFF2EFB30C9182CCA" blockId="103.[189,334,1166,1192]" box="[189,334,1166,1192]" pageId="103" pageNumber="104">
<heading id="C969818C9D7B923CFF2EFB30C9182CCA" bold="true" box="[189,334,1166,1192]" fontSize="11" level="3" pageId="103" pageNumber="104" reason="3">
<emphasis id="A0EAEAF29D7B923CFF2EFB30C9182CCA" bold="true" box="[189,334,1166,1192]" pageId="103" pageNumber="104">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7B923CFF2EFB00CB112CBB" blockId="103.[189,839,1213,1241]" box="[189,839,1213,1241]" pageId="103" pageNumber="104">
MEASUREMENTS. Total body length
<quantity id="55669B059D7B923CFDC2FB03CAEF2CBA" box="[593,697,1213,1240]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="103" pageNumber="104" unit="mm" value="6.8">6.80 mm</quantity>
(
<typeStatus id="4D2588429D7B923CFD5BFB00CB6D2CBB" box="[712,827,1214,1241]" pageId="103" pageNumber="104" type="lectotype">lectotype</typeStatus>
).
</paragraph>
<paragraph id="922136E09D7B9233FF2EFABECA482A42" blockId="103.[189,1399,1280,2025]" lastBlockId="104.[189,1399,265,545]" lastPageId="104" lastPageNumber="105" pageId="103" pageNumber="104">
Dorsum and venter uniformly dark brown, appendages paler brown. Head convex dorsally between compound eyes, with inconspicuous transverse impression at front margin of either eye, glossy dorsally and ventrally. Male labroclypeus (
<figureCitation id="0AA52A659D7B923CFDC3FAF6CAE32D00" box="[592,693,1352,1378]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="103" pageNumber="104">Fig. 171</figureCitation>
) deeply U-shaped emarginate on anterior margin, its lateral margins strongly sinuous in both dorsal and lateral views. Anterolateral angles rounded, strongly protruding, raised up at angle of nearly 80–90° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Underside of labroclypeus with sparse long delicate setae along anterior and lateral margins. Canthus strongly obtusely angulate in dorsal view. Punctures of frons irregularly hexagonal, in part annular, moderately large and deep. Intervening spaces glossy, variably large. Head setae minute, appressed; each seta rises from anterior margin of corresponding puncture not surpassing its length. Antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, glossy dorsally, opaque laterally. Anterior margin of pronotum broadly emarginate with strongly protruding anterolateral angles, basal margin sinuous. Lateral margin of pronotum very broadly rounded in dorsal view, crenulate all along except denticulate prebasal area; lateral margin with 13–15 crenulae / denticles. Inconspicuous suberect seta present between every two crenulae (
<figureCitation id="0AA52A659D7B923CFD6FF96DCB372E8C" box="[764,865,1747,1774]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="103" pageNumber="104">Fig. 250</figureCitation>
). Lateral margin of pronotum sinuous in lateral view. Pronotal punctures irregularly ovoid to hexagonal, annular, dense, moderately deep. Intervening spaces glossy, flat. Setation as on head. Hypomeron separated from prosternum by moderately high nearly straight carina, with long setae on anterolateral margin opposite of compound eye. Antennal pocket moderately deep. Scutellar shield rounded apically. Elytron subopaque, with vague tracks of three longitudinal carinae, including sutural one. Elytral punctures sinuous (elongate and narrow), incision-shaped, shallow (
<figureCitation id="0AA52A659D7B923CFE7CF814CA002FA7" box="[495,598,1962,1989]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="103" pageNumber="104">Fig. 341</figureCitation>
). Intervening spaces wrinkled. Appressed to suberect seta rises from anterior margin of each puncture, not surpassing length of corresponding incision. Lateral margins of elytra in part covered with microscopical velvety pubescence. Male and female pygidium flattened dorsally, opaque, with large shallow annular punctures (
<figureCitation id="0AA52A659D749233FCCAFE93CB97292A" box="[857,961,301,328]" captionStart="Figs 504–521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504–521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="104" pageNumber="105">Fig. 515</figureCitation>
). Intervening spaces microreticulate, much smaller than punctures. Setae of pygidium suberect, surpassing length of corresponding punctures. Male protibia widened distally, with three external teeth, median one largest (Fig. 413). Male protibial terminal spur narrow. Male metatibia distally with a brush of dense long sinuous setae. Male metatibial terminal spurs long, unequal, lower spur slightly curved. Male basal metatarsomere with a brush of dense long sinuous setae on inner margin. Tarsal claws with pulvilli. Spiculum gastrale as in
<figureCitation id="0AA52A659D749233FA9FFE5CCD25299F" box="[1292,1395,482,509]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="104" pageNumber="105">Fig. 560</figureCitation>
. Aedeagus as in
<figureCitation id="0AA52A659D749233FEE5FDB8CA4C2A42" box="[374,538,518,544]" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="104" pageNumber="105">Figs 706–708</figureCitation>
.
</paragraph>
<paragraph id="922136E09D749233FF2EFDF6C9E62A00" blockId="104.[189,432,584,610]" box="[189,432,584,610]" pageId="104" pageNumber="105">
<heading id="C969818C9D749233FF2EFDF6C9E62A00" bold="true" box="[189,432,584,610]" fontSize="11" level="3" pageId="104" pageNumber="105" reason="3">
<emphasis id="A0EAEAF29D749233FF2EFDF6C9E62A00" bold="true" box="[189,432,584,610]" pageId="104" pageNumber="105">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D749233FF2EFDC6C9102AD4" blockId="104.[189,1399,632,694]" pageId="104" pageNumber="105">Female lamellae of antennal club shorter; female pygidium with inconspicuous preapical dorsal impression.</paragraph>
</subSubSection>
</treatment>
</document>
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