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<document id="98171C87DA0795D564BF6D610C482A3B" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D73922AFD95F92DC9A72EFB" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius speciosus Telnov 2020, comb. nov." docType="treatment" docVersion="10" lastPageNumber="114" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallaces Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="112" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5A7C39E3BF2A0376E6E1066C4C618746">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallaces Line</mods:title>
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<mods:namePart id="DB34C764C3A2BFD0CE021E40CD5A980D">Telnov, Dmitry</mods:namePart>
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<mods:date id="8EA61CF7C26A0156B3EBB91F841A715D">2020</mods:date>
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<treatment id="1A3787F69D73922AFD95F92DC9A72EFB" ID-DOI="http://doi.org/10.5281/zenodo.4344423" ID-GBIF-Taxon="168648127" ID-Zenodo-Dep="4344423" LSID="urn:lsid:plazi:treatment:1A3787F69D73922AFD95F92DC9A72EFB" httpUri="http://treatment.plazi.org/id/1A3787F69D73922AFD95F92DC9A72EFB" lastPageId="113" lastPageNumber="114" pageId="111" pageNumber="112">
<subSubSection id="DA84656B9D739234FD95F92DCC782ECC" box="[518,1070,1683,1710]" pageId="111" pageNumber="112" type="nomenclature">
<paragraph id="922136E09D739234FD95F92DCC782ECC" blockId="111.[427,1160,1683,1745]" box="[518,1070,1683,1710]" pageId="111" pageNumber="112">
<heading id="C969818C9D739234FD95F92DCC782ECC" box="[518,1070,1683,1710]" centered="true" fontSize="11" level="2" pageId="111" pageNumber="112" reason="2">
<taxonomicName id="559E4D639D739234FD95F92DCBF32ECC" ID-CoL="7X3XS" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[518,933,1683,1710]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="speciosus" status="comb. nov.">
<emphasis id="A0EAEAF29D739234FD95F92DCB5B2ECF" bold="true" box="[518,781,1683,1710]" italics="true" pageId="111" pageNumber="112">Maechidius speciosus</emphasis>
(
<bibRefCitation id="F60F4B119D739234FC8FF92DCBCB2ECC" author="Frey G." box="[796,925,1683,1710]" pageId="111" pageNumber="112" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D739234FC3FF92DCC782ECC" box="[940,1070,1683,1710]" pageId="111" pageNumber="112" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D739234FE38F908CCDE2EB2" box="[427,1160,1718,1745]" pageId="111" pageNumber="112" type="description">
<paragraph id="922136E09D739234FE38F908CCDE2EB2" blockId="111.[427,1160,1683,1745]" box="[427,1160,1718,1745]" pageId="111" pageNumber="112">
<figureCitation id="0AA52A659D739234FE38F908CA532EB2" box="[427,517,1718,1744]" captionStart="Figs8689" captionStartId="112.[189,241,1928,1954]" captionTargetBox="[282,1293,267,1862]" captionTargetId="figure-66@112.[278,1296,259,1877]" captionTargetPageId="112" captionText="Figs8689. PapuanandWallaceanMaechidiusMacleay, 1819,habitus, dorsalview.86.M.speciosus(Frey, 1969) comb. nov., ♀ (BMNH). 87. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 88. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 89. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122167" httpUri="https://zenodo.org/record/4122167/files/figure.png" pageId="111" pageNumber="112">Figs 86</figureCitation>
,
<figureCitation id="0AA52A659D739234FD82F908CA2D2EB2" box="[529,635,1718,1744]" captionStart="Figs 169180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="111" pageNumber="112">179180</figureCitation>
,
<figureCitation id="0AA52A659D739234FD14F908CAE02EB3" box="[647,694,1718,1745]" captionStart="Figs 242258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="111" pageNumber="112">256</figureCitation>
,
<figureCitation id="0AA52A659D739234FD52F908CAA62EB2" box="[705,752,1718,1744]" captionStart="Figs 278285" captionStartId="151.[189,241,1697,1723]" captionTargetBox="[211,1375,265,1597]" captionTargetId="figure-104@151.[211,1376,265,1614]" captionTargetPageId="151" captionText="Figs 278285. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventro-lateral view. 278. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 279. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 280. M. seriepunctatus Moser, 1920, lectotype, ♂ (SNSD). 281. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 282. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 283. M. trivialis sp. nov., holotype, ♂ (BMNH). 284. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 285. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122202" httpUri="https://zenodo.org/record/4122202/files/figure.png" pageId="111" pageNumber="112">281</figureCitation>
,
<figureCitation id="0AA52A659D739234FD6FF908CB7D2EB2" box="[764,811,1718,1744]" captionStart="Figs 333348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="111" pageNumber="112">346</figureCitation>
, 418, 519521, 562, 718720
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D739234FF2EF948CAEB2F73" box="[189,701,1782,1809]" pageId="111" pageNumber="112" type="reference_group">
<paragraph id="922136E09D739234FF2EF948CAEB2F73" blockId="111.[189,701,1782,1809]" box="[189,701,1782,1809]" pageId="111" pageNumber="112">
<treatmentCitationGroup id="B28E11CE9D739234FF2EF948CAEB2F73" box="[189,701,1782,1809]" pageId="111" pageNumber="112">
<treatmentCitation id="133F10F19D739234FF2EF948CAEF2F73" author="Frey G." box="[189,697,1782,1809]" page="507" pageId="111" pageNumber="112" year="1969">
<taxonomicName id="559E4D639D739234FF2EF948CAEF2F73" ID-CoL="7XWMQ" authority="Frey, 1969: 507" authorityName="Frey" authorityPageNumber="507" authorityYear="1969" box="[189,697,1782,1809]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis id="A0EAEAF29D739234FF2EF948C9A32F72" box="[189,501,1782,1808]" italics="true" pageId="111" pageNumber="112">Paramaechidius speciosus</emphasis>
<bibRefCitation id="F60F4B119D739234FE6FF949CAEF2F73" author="Frey G." box="[508,697,1782,1809]" pageId="111" pageNumber="112" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969: 507</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D73922AFF2EF889CA742AE4" lastPageId="113" lastPageNumber="114" pageId="111" pageNumber="112" type="materials_examined">
<paragraph id="922136E09D739234FF2EF889C93A2F33" blockId="111.[189,364,1847,1873]" box="[189,364,1847,1873]" pageId="111" pageNumber="112">
<heading id="C969818C9D739234FF2EF889C93A2F33" bold="true" box="[189,364,1847,1873]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
<emphasis id="A0EAEAF29D739234FF2EF889C93A2F33" bold="true" box="[189,364,1847,1873]" pageId="111" pageNumber="112">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D739234FF79F8DBCD212F8B" ID-GBIF-Occurrence="2898878336" collectingDate="1964-10-15" collectingDateMax="1964-10-24" collectingDateMin="1964-10-15" collectionCode="NEW, TYPUS, TYPE, BMNH" collectorName="M. E. Bacchus." country="Papua New Guinea" county="Mts. Budemu" determinerName="G. Frey" elevation="1219" location="Holotype" municipality="Finisterre Mts." pageId="111" pageNumber="112" specimenCount="1" typeStatus="holotype">
<paragraph id="922136E09D739234FF79F8DBC90A2F1D" blockId="111.[189,1399,1893,2025]" box="[234,348,1893,1919]" pageId="111" pageNumber="112">
<emphasis id="A0EAEAF29D739234FF79F8DBC90A2F1D" bold="true" box="[234,348,1893,1919]" pageId="111" pageNumber="112">
<typeStatus id="4D2588429D739234FF79F8DBC90A2F1D" box="[234,348,1893,1919]" pageId="111" pageNumber="112" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D739234FF2EF836CD212F8B" blockId="111.[189,1399,1893,2025]" pageId="111" pageNumber="112">
<collectingCountry id="EA8976709D739234FF2EF836C98C2FC0" box="[189,474,1928,1954]" name="Papua New Guinea" pageId="111" pageNumber="112">PAPUA NEW GUINEA</collectingCountry>
<specimenCount id="8498FD699D739234FE64F836CA592FC0" box="[503,527,1928,1954]" pageId="111" pageNumber="112"></specimenCount>
; “
<typeStatus id="4D2588429D739234FDB4F839CAC02FC3" box="[551,662,1927,1953]" pageId="111" pageNumber="112" type="holotype">
<location id="9741603B9D739234FDB4F839CAC02FC3" LSID="urn:lsid:plazi:treatment:1A3787F69D73922AFD95F92DC9A72EFB:9741603B9D739234FDB4F839CAC02FC3" box="[551,662,1927,1953]" country="Papua New Guinea" county="Mts. Budemu" municipality="Finisterre Mts." name="Holotype" pageId="111" pageNumber="112">Holotype</location>
</typeStatus>
[p, label circular, red frame] //
<collectionCode id="F48FAE259D739234FB97F836CC102FC0" box="[1028,1094,1928,1954]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="111" pageNumber="112">NEW</collectionCode>
<collectingCountry id="EA8976709D739234FBDCF836CC952FC0" box="[1103,1219,1928,1954]" name="Guinea-Bissau" pageId="111" pageNumber="112">GUINEA</collectingCountry>
:
<location id="9741603B9D739234FB5CF839CD242FC3" LSID="urn:lsid:plazi:treatment:1A3787F69D73922AFD95F92DC9A72EFB:9741603B9D739234FB5CF839CD242FC3" box="[1231,1394,1927,1953]" country="Papua New Guinea" county="Mts. Budemu" municipality="Finisterre Mts." name="Madang Dist." pageId="111" pageNumber="112">Madang Dist.</location>
,
<collectingMunicipality id="7245AC9A9D739234FF2EF814C93C2FA7" box="[189,362,1962,1989]" pageId="111" pageNumber="112">
Finisterre
<collectingCounty id="7B404E6C9D739234FEA6F815C93C2FA7" box="[309,362,1963,1989]" pageId="111" pageNumber="112">Mts.</collectingCounty>
</collectingMunicipality>
Budemu c.
<quantity id="55669B059D739234FE6EF814CA0C2FA6" box="[509,602,1962,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="111" pageNumber="112" unit="ft" value="4000.0">
<elevation id="19B3D1D39D739234FE6EF814CA0C2FA6" box="[509,602,1962,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="111" pageNumber="112" unit="ft" value="4000.0">4000 ft.</elevation>
</quantity>
<date id="E62010209D739234FDF0F814CAA82FA7" box="[611,766,1962,1989]" pageId="111" pageNumber="112" value="1964-10-15" valueMax="1964-10-24" valueMin="1964-10-15">
<collectingDate id="F664E9C89D739234FDF0F814CAA82FA7" box="[611,766,1962,1989]" pageId="111" pageNumber="112" value="1964-10-15" valueMax="1964-10-24" valueMin="1964-10-15">15-24.x.1964</collectingDate>
</date>
[p]//
<collectorName id="3F6B53369D739234FCD3F815CBB82FA6" box="[832,1006,1962,1989]" pageId="111" pageNumber="112">M.E. Bacchus.</collectorName>
B.M. 1965-120 [p]// Stn. No. 51 [p]//
<collectionCode id="F48FAE259D739234FF66F870C9072F8A" box="[245,337,1998,2024]" pageId="111" pageNumber="112">
<typeStatus id="4D2588429D739234FF66F870C9072F8A" box="[245,337,1998,2024]" pageId="111" pageNumber="112">TYPUS</typeStatus>
</collectionCode>
[p, red label]//
<collectionCode id="F48FAE259D739234FD92F871CA1C2F8B" box="[513,586,1999,2025]" pageId="111" pageNumber="112">
<typeStatus id="4D2588429D739234FD92F871CA1C2F8B" box="[513,586,1999,2025]" pageId="111" pageNumber="112">TYPE</typeStatus>
</collectionCode>
<taxonomicName id="559E4D639D739234FDC3F870CBD52F8A" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[592,899,1998,2024]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="speciosus" status="sp. nov.">
<emphasis id="A0EAEAF29D739234FDC3F870CBD52F8A" box="[592,899,1998,2024]" italics="true" pageId="111" pageNumber="112">Paramaechidius speciosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D739234FC1AF871CB902F8B" box="[905,966,1999,2025]" pageId="111" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
[h] det.
<determinerName id="67FDDA249D739234FB8DF870CC262F8A" box="[1054,1136,1998,2024]" pageId="111" pageNumber="112">G.Frey</determinerName>
,1967/68 [p]”;
<collectionCode id="F48FAE259D739234FA8BF871CD252F8B" box="[1304,1395,1999,2025]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="111" pageNumber="112">BMNH</collectionCode>
.
</paragraph>
</materialsCitation>
<caption id="C6E166689D6C922BFF2EF836CD722F8B" ID-DOI="http://doi.org/10.5281/zenodo.4122167" ID-Zenodo-Dep="4122167" httpUri="https://zenodo.org/record/4122167/files/figure.png" pageId="112" pageNumber="113" startId="112.[189,241,1928,1954]" targetBox="[282,1293,267,1862]" targetPageId="112">
<paragraph id="922136E09D6C922BFF2EF836CD722F8B" blockId="112.[189,1399,1928,2026]" pageId="112" pageNumber="113">
<emphasis id="A0EAEAF29D6C922BFF2EF836C9122FC0" bold="true" box="[189,324,1928,1954]" pageId="112" pageNumber="113">Figs8689.</emphasis>
Papuan and Wallacean
<taxonomicName id="559E4D639D6C922BFDD9F836CBD72FC0" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[586,897,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D6C922BFDD9F836CA852FC0" box="[586,723,1928,1954]" italics="true" pageId="112" pageNumber="113">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D6C922BFD45F836CBD72FC0" author="Macleay W. S." box="[726,897,1928,1954]" pageId="112" pageNumber="113" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
,habitus, dorsal view.
<emphasis id="A0EAEAF29D6C922BFBE7F836CCC72FC0" bold="true" box="[1140,1169,1928,1954]" pageId="112" pageNumber="113">86</emphasis>
.
<taxonomicName id="559E4D639D6C922BFB08F837C9572FA4" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="speciosus" status="comb. nov.">
<emphasis id="A0EAEAF29D6C922BFB08F837CD782FC0" box="[1179,1326,1929,1954]" italics="true" pageId="112" pageNumber="113">M.speciosus</emphasis>
(
<bibRefCitation id="F60F4B119D6C922BFAAAF836C8AF2FA4" author="Frey G." pageId="112" pageNumber="113" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D6C922BFE99F815C9D82FA4" box="[266,398,1963,1990]" pageId="112" pageNumber="113" rank="species">comb. nov.</taxonomicNameLabel>
, ♀ (BMNH).
<emphasis id="A0EAEAF29D6C922BFDA7F815CA072FA7" bold="true" box="[564,593,1963,1989]" pageId="112" pageNumber="113">87</emphasis>
.
<taxonomicName id="559E4D639D6C922BFDF2F812CB2B2FA4" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[609,893,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="sturnus">
<emphasis id="A0EAEAF29D6C922BFDF2F812CA892FA7" box="[609,735,1964,1989]" italics="true" pageId="112" pageNumber="113">M. sturnus</emphasis>
<bibRefCitation id="F60F4B119D6C922BFD75F812CB2B2FA4" author="Arrow G. E." box="[742,893,1964,1990]" pageId="112" pageNumber="113" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
, holotype, ♂ (BMNH).
<emphasis id="A0EAEAF29D6C922BFB0AF815CCE02FA7" bold="true" box="[1177,1206,1963,1989]" pageId="112" pageNumber="113">88</emphasis>
.
<taxonomicName id="559E4D639D6C922BFB55F812C9072F88" authority="Heller, 1895" authorityName=", Hell. Determined" authorityYear="1895" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="subcostatus">
<emphasis id="A0EAEAF29D6C922BFB55F812CD212FA7" box="[1222,1399,1963,1989]" italics="true" pageId="112" pageNumber="113">M. subcostatus</emphasis>
<bibRefCitation id="F60F4B119D6C922BFF2EF871C9072F88" author="Heller K. M." box="[189,337,1999,2026]" pageId="112" pageNumber="113" pagination="1 - 17" refId="ref98131" refString="Heller K. M. 1895. Erster Beitrag zur Papuaniscen [sic!] Kaferfauna. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden 1894 / 1895 (16): 1 - 17." type="journal article" year="1895">Heller, 1895</bibRefCitation>
</taxonomicName>
, lectotype, ♂ (SNSD).
<emphasis id="A0EAEAF29D6C922BFDF1F871CA292F8B" bold="true" box="[610,639,1999,2025]" pageId="112" pageNumber="113">89</emphasis>
.
<taxonomicName id="559E4D639D6C922BFD1DF86ECB422F8B" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[654,788,2000,2025]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="suwawa" status="sp. nov.">
<emphasis id="A0EAEAF29D6C922BFD1DF86ECB422F8B" box="[654,788,2000,2025]" italics="true" pageId="112" pageNumber="113">M. suwawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D6C922BFC88F871CB2E2F8B" box="[795,888,1999,2025]" pageId="112" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (BMNH). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D6D922AFF2EFEB7C9332940" blockId="113.[189,357,264,291]" box="[189,357,264,291]" pageId="113" pageNumber="114">
<heading id="C969818C9D6D922AFF2EFEB7C9332940" bold="true" box="[189,357,264,291]" fontSize="11" level="3" pageId="113" pageNumber="114" reason="3">
<emphasis id="A0EAEAF29D6D922AFF2EFEB7C9332940" bold="true" box="[189,357,264,291]" pageId="113" pageNumber="114">New material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6D922AFF2EFE88CA742AE4" blockId="113.[189,1399,310,646]" pageId="113" pageNumber="114">
<materialsCitation id="22F63CBD9D6D922AFF2EFE88C9BA2911" ID-GBIF-Occurrence="2898878419" collectingDate="1936-02" collectionCode="NEW, BMNH" country="Guinea-Bissau" elevation="213" location="Mt. Bougainville." municipality="Mt. Nomo." pageId="113" pageNumber="114" specimenCount="1">
<collectingCountry id="EA8976709D6D922AFF2EFE88C90D2932" box="[189,347,310,336]" name="Indonesia" pageId="113" pageNumber="114">INDONESIA</collectingCountry>
<specimenCount id="8498FD699D6D922AFEF8FE88C9C32933" box="[363,405,310,337]" pageId="113" pageNumber="114" type="male">1♂</specimenCount>
;“W.
<collectionCode id="F48FAE259D6D922AFE5BFE88CA582932" box="[456,526,310,336]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="113" pageNumber="114">NEW</collectionCode>
<collectingCountry id="EA8976709D6D922AFD82FE88CAD02932" box="[529,646,310,336]" name="Guinea-Bissau" pageId="113" pageNumber="114">GUINEA</collectingCountry>
:
<collectingMunicipality id="7245AC9A9D6D922AFD1EFE89CB512933" box="[653,775,311,337]" pageId="113" pageNumber="114">Mt.Nomo.</collectingMunicipality>
S.of
<location id="9741603B9D6D922AFCAEFE88CC502932" LSID="urn:lsid:plazi:treatment:1A3787F69D73922AFD95F92DC9A72EFB:9741603B9D6D922AFCAEFE88CC502932" box="[829,1030,310,336]" country="Guinea-Bissau" municipality="Mt. Nomo." name="Mt. Bougainville." pageId="113" pageNumber="114">Mt.Bougainville.</location>
<quantity id="55669B059D6D922AFB9AFE88CC042932" box="[1033,1106,310,336]" metricMagnitude="2" metricUnit="m" metricValue="2.1336" pageId="113" pageNumber="114" unit="ft" value="700.0">
<elevation id="19B3D1D39D6D922AFB9AFE88CC042932" box="[1033,1106,310,336]" metricMagnitude="2" metricUnit="m" metricValue="2.1336" pageId="113" pageNumber="114" unit="ft" value="700.0">700ft.</elevation>
</quantity>
<date id="E62010209D6D922AFBC1FE88CCF12932" box="[1106,1191,310,336]" pageId="113" pageNumber="114" value="1936-02">
<collectingDate id="F664E9C89D6D922AFBC1FE88CCF12932" box="[1106,1191,310,336]" pageId="113" pageNumber="114" value="1936-02">ii.1936</collectingDate>
</date>
.// L.E.Cheesman. B.M.1936-271.”;
<collectionCode id="F48FAE259D6D922AFE07FEE7C9BA2911" box="[404,492,345,371]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="113" pageNumber="114">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="22F63CBD9D6D922AFD80FEE7CCC929F7" ID-GBIF-Occurrence="2898878471" collectingDate="1936-03" collectionCode="DUTCH, NEW, BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" determinerName="G. Frey" elevation="1067" location="Mt. Lina." municipality="Cyclops Mts." pageId="113" pageNumber="114" specimenCount="1">
<specimenCount id="8498FD699D6D922AFD80FEE7CA112911" box="[531,583,345,371]" pageId="113" pageNumber="114" type="male">1 ♂</specimenCount>
; “
<specimenCount id="8498FD699D6D922AFDFAFEE7CA292911" box="[617,639,345,371]" pageId="113" pageNumber="114"></specimenCount>
//
<collectionCode id="F48FAE259D6D922AFD05FEE7CAAD2911" box="[662,763,345,371]" pageId="113" pageNumber="114">DUTCH</collectionCode>
<collectionCode id="F48FAE259D6D922AFC9AFEE7CB1D2911" box="[777,843,345,371]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="113" pageNumber="114">NEW</collectionCode>
<collectingCountry id="EA8976709D6D922AFCCAFEE7CB982911" box="[857,974,345,371]" name="Guinea-Bissau" pageId="113" pageNumber="114">GUINEA</collectingCountry>
:
<collectingMunicipality id="7245AC9A9D6D922AFC73FEE6CCD02911" box="[992,1158,344,371]" pageId="113" pageNumber="114">Cyclops Mts.</collectingMunicipality>
,
<location id="9741603B9D6D922AFB0AFEE7CD582910" LSID="urn:lsid:plazi:treatment:1A3787F69D73922AFD95F92DC9A72EFB:9741603B9D6D922AFB0AFEE7CD582910" box="[1177,1294,344,371]" country="Guinea-Bissau" municipality="Cyclops Mts." name="Mt. Lina." pageId="113" pageNumber="114">Mt. Lina.</location>
<quantity id="55669B059D6D922AFA88FEE6CD212911" box="[1307,1399,344,371]" metricMagnitude="3" metricUnit="m" metricValue="1.0668" pageId="113" pageNumber="114" unit="ft" value="3500.0">
<elevation id="19B3D1D39D6D922AFA88FEE6CD212911" box="[1307,1399,344,371]" metricMagnitude="3" metricUnit="m" metricValue="1.0668" pageId="113" pageNumber="114" unit="ft" value="3500.0">3,500ft.</elevation>
</quantity>
<date id="E62010209D6D922AFF2EFEC5C97029F7" box="[189,294,379,405]" pageId="113" pageNumber="114" value="1936-03">
<collectingDate id="F664E9C89D6D922AFF2EFEC5C97029F7" box="[189,294,379,405]" pageId="113" pageNumber="114" value="1936-03">iii. 1936</collectingDate>
</date>
.
<collectorName id="3F6B53369D6D922AFEA5FEC5C9BC29F7" box="[310,490,379,405]" pageId="113" pageNumber="114">L.E.Cheesman.</collectorName>
B.M.1936-271.//
<taxonomicName id="559E4D639D6D922AFD52FEC5CB7B29F7" authorityName="Heller" authorityYear="1910" box="[705,813,379,405]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="pauxillus">
<emphasis id="A0EAEAF29D6D922AFD52FEC5CB7B29F7" box="[705,813,379,405]" italics="true" pageId="113" pageNumber="114">pauxillus</emphasis>
</taxonomicName>
det.
<determinerName id="67FDDA249D6D922AFCF0FEC5CBE029F7" box="[867,950,379,405]" pageId="113" pageNumber="114">G.Frey</determinerName>
, 1967/68”;
<collectionCode id="F48FAE259D6D922AFBD4FEC5CCC929F7" box="[1095,1183,379,405]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="113" pageNumber="114">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="22F63CBD9D6D922AFB52FEC5C94329B8" ID-GBIF-Occurrence="2898878504" collectingDate="1938-08" collectionCode="DUTCH, NEW, BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" elevation="30" location="Japen" pageId="113" pageNumber="114" specimenCount="1">
<specimenCount id="8498FD699D6D922AFB52FEC5CCA529F7" box="[1217,1267,379,405]" pageId="113" pageNumber="114" type="male">1 ♂</specimenCount>
; “
<collectionCode id="F48FAE259D6D922AFA9DFEC5CD2029F7" box="[1294,1398,379,405]" pageId="113" pageNumber="114">DUTCH</collectionCode>
<collectionCode id="F48FAE259D6D922AFF2EFE20C8A929DA" box="[189,255,414,440]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="113" pageNumber="114">NEW</collectionCode>
<collectingCountry id="EA8976709D6D922AFE9CFE23C9D229D5" box="[271,388,413,439]" name="Guinea-Bissau" pageId="113" pageNumber="114">GUINEA</collectingCountry>
:
<location id="9741603B9D6D922AFE04FE20C98C29DA" LSID="urn:lsid:plazi:treatment:1A3787F69D73922AFD95F92DC9A72EFB:9741603B9D6D922AFE04FE20C98C29DA" box="[407,474,414,440]" country="Guinea-Bissau" name="Japen" pageId="113" pageNumber="114">Japen</location>
[sic
<specimenCount id="8498FD699D6D922AFD89FE23CA4829DA" box="[538,542,413,440]" pageId="113" pageNumber="114"></specimenCount>
] I., Mt. Baduri.
<quantity id="55669B059D6D922AFD6DFE23CB0329D5" box="[766,853,413,439]" metricMagnitude="1" metricUnit="m" metricValue="3.048" pageId="113" pageNumber="114" unit="ft" value="100.0">
<elevation id="19B3D1D39D6D922AFD6DFE23CB0329D5" box="[766,853,413,439]" metricMagnitude="1" metricUnit="m" metricValue="3.048" pageId="113" pageNumber="114" unit="ft" value="100.0">100 ft.</elevation>
</quantity>
,
<date id="E62010209D6D922AFCF9FE23CB8029D5" box="[874,982,413,439]" pageId="113" pageNumber="114" value="1938-08">
<collectingDate id="F664E9C89D6D922AFCF9FE23CB8029D5" box="[874,982,413,439]" pageId="113" pageNumber="114" value="1938-08">viii.1938</collectingDate>
</date>
.
<collectorName id="3F6B53369D6D922AFC79FE23CCC829D5" box="[1002,1182,413,439]" pageId="113" pageNumber="114">L.E.Cheesman.</collectorName>
B.M.1938-596.”;
<collectionCode id="F48FAE259D6D922AFF2EFE7EC94329B8" box="[189,277,448,474]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="113" pageNumber="114">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="22F63CBD9D6D922AFEA1FE7EC9432A7D" ID-GBIF-Occurrence="2898878308" collectingDate="1964-10-15" collectingDateMax="1964-10-24" collectingDateMin="1964-10-15" collectionCode="NEW, BMNH" collectorName="M. E. Bacchus." country="Guinea-Bissau" determinerName="G. Frey" elevation="1219" location="Finisterre Mts." municipality="Madang Dist." pageId="113" pageNumber="114" specimenCount="5" specimenCount-female="5">
<specimenCount id="8498FD699D6D922AFEA1FE7EC93729B8" box="[306,353,448,474]" pageId="113" pageNumber="114" type="male">1 ♂</specimenCount>
,
<specimenCount id="8498FD699D6D922AFEE3FE01C9E329B8" box="[368,437,447,474]" pageId="113" pageNumber="114" type="female">5 ♀♀</specimenCount>
; “Stn. No. 51. //
<collectionCode id="F48FAE259D6D922AFDE8FE7ECAEB29B8" box="[635,701,448,474]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="113" pageNumber="114">NEW</collectionCode>
<collectingCountry id="EA8976709D6D922AFD55FE7ECB6D29B8" box="[710,827,448,474]" name="Guinea-Bissau" pageId="113" pageNumber="114">GUINEA</collectingCountry>
:
<collectingMunicipality id="7245AC9A9D6D922AFCDBFE01CBBA29BB" box="[840,1004,447,473]" pageId="113" pageNumber="114">Madang Dist.</collectingMunicipality>
,
<location id="9741603B9D6D922AFC69FE01CCFF29B8" LSID="urn:lsid:plazi:treatment:1A3787F69D73922AFD95F92DC9A72EFB:9741603B9D6D922AFC69FE01CCFF29B8" box="[1018,1193,447,474]" country="Guinea-Bissau" municipality="Madang Dist." name="Finisterre Mts." pageId="113" pageNumber="114">Finisterre Mts.</location>
Budemu c.
<quantity id="55669B059D6D922AFAAFFE7EC881299E" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="113" pageNumber="114" unit="ft" value="4000.0">
<elevation id="19B3D1D39D6D922AFAAFFE7EC881299E" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="113" pageNumber="114" unit="ft" value="4000.0">4000 ft.</elevation>
</quantity>
<date id="E62010209D6D922AFF71FE5CC928299F" box="[226,382,482,509]" pageId="113" pageNumber="114" value="1964-10-15" valueMax="1964-10-24" valueMin="1964-10-15">
<collectingDate id="F664E9C89D6D922AFF71FE5CC928299F" box="[226,382,482,509]" pageId="113" pageNumber="114" value="1964-10-15" valueMax="1964-10-24" valueMin="1964-10-15">15-24.x.1964</collectingDate>
</date>
//
<collectorName id="3F6B53369D6D922AFE05FE5CCA11299E" box="[406,583,482,508]" pageId="113" pageNumber="114">M.E. Bacchus.</collectorName>
B.M. 1965-120.//
<taxonomicName id="559E4D639D6D922AFCBAFE5CCC37299E" authorityName="Heller" authorityYear="1910" box="[809,1121,482,508]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="pauxillus">
<emphasis id="A0EAEAF29D6D922AFCBAFE5CCC37299E" box="[809,1121,482,508]" italics="true" pageId="113" pageNumber="114">Paramaechidius pauxillus</emphasis>
</taxonomicName>
det.
<determinerName id="67FDDA249D6D922AFB05FE5CCCBF299E" box="[1174,1257,482,508]" pageId="113" pageNumber="114">G.Frey</determinerName>
, 1967/68.”;
<collectionCode id="F48FAE259D6D922AFF2EFDBBC9432A7D" box="[189,277,517,543]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="113" pageNumber="114">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="22F63CBD9D6D922AFEBEFDBACB122A23" ID-GBIF-Occurrence="2898878443" collectingDate="1964-10-02" collectingDateMax="1964-10-11" collectingDateMin="1964-10-02" collectionCode="NEW, BMNH" collectorName="M. E. Bacchus." country="Guinea-Bissau" determinerName="G. Frey" elevation="1082" isTailEnumeration="true" location="Finisterre Mts." municipality="Madang Dist." pageId="113" pageNumber="114" specimenCount="3" stateProvince="Madang">
<specimenCount id="8498FD699D6D922AFEBEFDBAC9972A7C" box="[301,449,516,542]" pageId="113" pageNumber="114" type="generic">3 specimens</specimenCount>
; “Stn. No. 46. //
<collectionCode id="F48FAE259D6D922AFDEDFDBBCA962A7D" box="[638,704,517,543]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="113" pageNumber="114">NEW</collectionCode>
<collectingCountry id="EA8976709D6D922AFD54FDBACB6A2A7C" box="[711,828,516,542]" name="Guinea-Bissau" pageId="113" pageNumber="114">GUINEA</collectingCountry>
:
<collectingRegion id="505AF8029D6D922AFCD4FDBACBBF2A7C" box="[839,1001,516,542]" country="Papua New Guinea" name="Madang" pageId="113" pageNumber="114">Madang Dist.</collectingRegion>
,
<location id="9741603B9D6D922AFC67FDBACCF72A7D" LSID="urn:lsid:plazi:treatment:1A3787F69D73922AFD95F92DC9A72EFB:9741603B9D6D922AFC67FDBACCF72A7D" box="[1012,1185,516,543]" country="Guinea-Bissau" name="Finisterre Mts." pageId="113" pageNumber="114" stateProvince="Madang">Finisterre Mts.</location>
Damanti
<quantity id="55669B059D6D922AFA87FDBACD212A7C" box="[1300,1399,516,543]" metricMagnitude="3" metricUnit="m" metricValue="1.08204" pageId="113" pageNumber="114" unit="ft" value="3550.0">
<elevation id="19B3D1D39D6D922AFA87FDBACD212A7C" box="[1300,1399,516,543]" metricMagnitude="3" metricUnit="m" metricValue="1.08204" pageId="113" pageNumber="114" unit="ft" value="3550.0">3,550 ft.</elevation>
</quantity>
<date id="E62010209D6D922AFF2EFD98C91C2A22" box="[189,330,550,576]" pageId="113" pageNumber="114" value="1964-10-02" valueMax="1964-10-11" valueMin="1964-10-02">
<collectingDate id="F664E9C89D6D922AFF2EFD98C91C2A22" box="[189,330,550,576]" pageId="113" pageNumber="114" value="1964-10-02" valueMax="1964-10-11" valueMin="1964-10-02">2-11.x.1964</collectingDate>
</date>
// M.E. Bacchus. B.M. 1965-120.”;
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</materialsCitation>
<materialsCitation id="22F63CBD9D6D922AFCCEFD99CA482AE4" ID-GBIF-Occurrence="2898878509" collectingDate="2000-12-16" collectionCode="NHMB" country="Papua New Guinea" county="Collection Naturhistorisches Museum Basel" latitude="-4.7" location="Salemben village" longLatPrecision="1306" longitude="145.4" municipality="Cizek" pageId="113" pageNumber="114" specimenCount="26" specimenCount-female="26" stateProvince="Madang">
17 ♂♂,
<specimenCount id="8498FD699D6D922AFC28FD98CC5B2A23" box="[955,1037,550,577]" pageId="113" pageNumber="114" type="female">26 ♀♀</specimenCount>
; “
<collectingCounty id="7B404E6C9D6D922AFBB6FD98C9382A01" pageId="113" pageNumber="114">Collection Naturhistorisches Museum Basel</collectingCounty>
//
<collectingCountry id="EA8976709D6D922AFE15FDF7CA3B2A01" box="[390,621,585,611]" name="Papua New Guinea" pageId="113" pageNumber="114">Papua New Guinea</collectingCountry>
<collectingRegion id="505AF8029D6D922AFDE5FDF7CB4B2A01" box="[630,797,585,611]" country="Papua New Guinea" name="Madang" pageId="113" pageNumber="114">Madang Prov.</collectingRegion>
L.
<collectingMunicipality id="7245AC9A9D6D922AFCDBFDF7CBDA2A01" box="[840,908,585,611]" pageId="113" pageNumber="114">Cizek</collectingMunicipality>
lgt.//
<location id="9741603B9D6D922AFC47FDF7CCF72A01" LSID="urn:lsid:plazi:treatment:1A3787F69D73922AFD95F92DC9A72EFB:9741603B9D6D922AFC47FDF7CCF72A01" box="[980,1185,585,611]" country="Papua New Guinea" county="Collection Naturhistorisches Museum Basel" latitude="-4.7" longLatPrecision="1306" longitude="145.4" municipality="Cizek" name="Salemben village" pageId="113" pageNumber="114" stateProvince="Madang">Salemben village</location>
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<date id="E62010209D6D922AFF2EFDD5C9192AE7" box="[189,335,619,645]" pageId="113" pageNumber="114" value="2000-12-16">
<collectingDate id="F664E9C89D6D922AFF2EFDD5C9192AE7" box="[189,335,619,645]" pageId="113" pageNumber="114" value="2000-12-16">
16.XII.
<quantity id="55669B059D6D922AFE9DFDD5C9192AE7" box="[270,335,619,645]" metricMagnitude="6" metricUnit="m" metricValue="2.00075" pageId="113" pageNumber="114" unit="m" value="2000750.0">2000</quantity>
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, 750 m”;
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.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6D922AFF2EFD15C9A72EFB" pageId="113" pageNumber="114" type="description">
<paragraph id="922136E09D6D922AFF2EFD15C9182AA7" blockId="113.[189,334,683,709]" box="[189,334,683,709]" pageId="113" pageNumber="114">
<heading id="C969818C9D6D922AFF2EFD15C9182AA7" bold="true" box="[189,334,683,709]" fontSize="11" level="3" pageId="113" pageNumber="114" reason="3">
<emphasis id="A0EAEAF29D6D922AFF2EFD15C9182AA7" bold="true" box="[189,334,683,709]" pageId="113" pageNumber="114">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6D922AFF2EFD64CA9C2D85" blockId="113.[189,1399,729,1511]" pageId="113" pageNumber="114">
Dorsum and venter covered with microscopical velvety pubescence. Male labroclypeus (
<figureCitation id="0AA52A659D6D922AFA97FD67CD382A91" box="[1284,1390,729,755]" captionStart="Figs 169180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="113" pageNumber="114">Fig. 179</figureCitation>
) broadly rounded on anterior margin, its anterolateral angles acute and protruding. Female labroclypeus (
<figureCitation id="0AA52A659D6D922AFF55FCA0C9782B5A" box="[198,302,798,824]" captionStart="Figs 169180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="113" pageNumber="114">Fig. 180</figureCitation>
) with less acute anterolateral angles. Head punctures ovoid, rather deep and dense. Head setae scale-like, dense, suberect, moderately long, in part clavate. Some setae not surpassing length of corresponding punctures. Pronotum broadly emarginate on anterior margin with protruding anterolateral angles. Basal margin of pronotum slightly sinuous. Lateral margin of pronotum in dorsal view very broadly rounded in anterior half, shallowly emarginate postmedially towards base (
<figureCitation id="0AA52A659D6D922AFA90FC19CD3C2BA0" box="[1283,1386,935,962]" captionStart="Figs 242258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="113" pageNumber="114">Fig. 256</figureCitation>
). Hypomeron slightly emarginate and long setose on anterior margin which is flange-like protruding, shallowly emarginate and long setose opposite compound eye (
<figureCitation id="0AA52A659D6D922AFC49FC52CC1E2C64" box="[986,1096,1004,1030]" captionStart="Figs 278285" captionStartId="151.[189,241,1697,1723]" captionTargetBox="[211,1375,265,1597]" captionTargetId="figure-104@151.[211,1376,265,1614]" captionTargetPageId="151" captionText="Figs 278285. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventro-lateral view. 278. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 279. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 280. M. seriepunctatus Moser, 1920, lectotype, ♂ (SNSD). 281. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 282. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 283. M. trivialis sp. nov., holotype, ♂ (BMNH). 284. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 285. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122202" httpUri="https://zenodo.org/record/4122202/files/figure.png" pageId="113" pageNumber="114">Fig. 281</figureCitation>
). Antennal pocket deep. Pronotal punctures of irregular oblong shape, deep, intervening spaces generally larger than punctures. Pronotal setae scale-like, dense, suberect, in part minute and not surpassing length of corresponding punctures, in part (in particular, along lateral margins and on anterior margin opposing frons) long and clavate. Elytral punctures irregular in shape, deep, somewhat irregularly clustered (
<figureCitation id="0AA52A659D6D922AFB47FBCBCD6D2CF2" box="[1236,1339,1141,1168]" captionStart="Figs 333348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="113" pageNumber="114">Fig. 346</figureCitation>
) and generally smaller than those on pronotum. Setae between puncture clusters longer, scale-like, stronger erect, arranged in several irregular longitudinal rows. Setae in puncture clusters minute, not surpassing length of corresponding punctures. Male and female pygidium with deep irregularly shaped punctures. Intervening spaces glossy, variably large. Male pygidium with two flat latero-median humps (humps very inconspicuous in some specimens), with group of large long dense scale-like setae (can be strongly reduced in some specimens) pointing obliquely to middle or posteriad (
<figureCitation id="0AA52A659D6D922AFC61FAFDCCCE2D3C" box="[1010,1176,1347,1374]" captionStart="Figs 504521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="113" pageNumber="114">Figs 519520</figureCitation>
). Female pygidium with two latero-median humps and transverse row of long scale-like whitish setae pointing obliquely medially (
<figureCitation id="0AA52A659D6D922AFEA6FA36C9CB2DC1" box="[309,413,1416,1443]" captionStart="Figs 504521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="113" pageNumber="114">Fig. 521</figureCitation>
). Male and female protibia with two rather acute terminal teeth on external margin (Fig. 418). Male terminal protibial spur curved, female one straight and pointed. Spiculum gastrale as in
<figureCitation id="0AA52A659D6D922AFF2EFA73C9752D85" box="[189,291,1484,1511]" captionStart="Figs 545569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="113" pageNumber="114">Fig. 562</figureCitation>
. Male aedeagus as in
<figureCitation id="0AA52A659D6D922AFDB1FA73CA902D85" box="[546,710,1485,1511]" captionStart="Figs 712725" captionStartId="179.[189,241,1892,1918]" captionTargetBox="[433,1144,277,1858]" captionTargetId="figure-72@179.[427,1160,265,1870]" captionTargetPageId="179" captionText="Figs 712725. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 712 714. M. skalei sp. nov., holotype, ♂ (NME). 715717. M. sougb sp. nov., holotype, ♂ (NME). 718720. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 721723. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 724725. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122252" httpUri="https://zenodo.org/record/4122252/files/figure.png" pageId="113" pageNumber="114">Figs 718720</figureCitation>
.
</paragraph>
<paragraph id="922136E09D6D922AFF2EF9B3C9E62E45" blockId="113.[189,432,1549,1575]" box="[189,432,1549,1575]" pageId="113" pageNumber="114">
<heading id="C969818C9D6D922AFF2EF9B3C9E62E45" bold="true" box="[189,432,1549,1575]" fontSize="11" level="3" pageId="113" pageNumber="114" reason="3">
<emphasis id="A0EAEAF29D6D922AFF2EF9B3C9E62E45" bold="true" box="[189,432,1549,1575]" pageId="113" pageNumber="114">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6D922AFF2EF985C9A72EFB" blockId="113.[189,1399,1595,1690]" pageId="113" pageNumber="114">Lamellae of female antennal club shorter; apex of female elytron with conspicuous, long and rather acutely protruding in this species; female pygidium with a glossy flat median longitudinal area between two latero-median humps.</paragraph>
</subSubSection>
</treatment>
</document>