137 lines
19 KiB
XML
137 lines
19 KiB
XML
<document id="2263465CE366BF136D987259C7FB61B5" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D709237FDCDFEB6CB442D8F" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius simplex Frey 1969" docType="treatment" docVersion="10" lastPageNumber="109" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="109" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7D80FE902AAC3B70A3B8527F535C6817">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:namePart id="201B2F5453725E155A070311C43CF6CF">Telnov, Dmitry</mods:namePart>
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<mods:title id="318E9DC15F2C32D39A0D16DA6DB4BBBA">European Journal of Taxonomy</mods:title>
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<mods:part id="A8DEBCBF69B7127CB89D20B39E87B3CE">
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<mods:date id="97CDE0652C85E35D0591CB15D8CCB14B">2020</mods:date>
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<treatment id="1A3787F69D709237FDCDFEB6CB442D8F" ID-DOI="http://doi.org/10.5281/zenodo.4344537" ID-GBIF-Taxon="168648121" ID-Zenodo-Dep="4344537" LSID="urn:lsid:plazi:treatment:1A3787F69D709237FDCDFEB6CB442D8F" httpUri="http://treatment.plazi.org/id/1A3787F69D709237FDCDFEB6CB442D8F" lastPageNumber="109" pageId="108" pageNumber="109">
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<subSubSection id="DA84656B9D709237FDCDFEB6CB832941" box="[606,981,264,291]" pageId="108" pageNumber="109" type="nomenclature">
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<paragraph id="922136E09D709237FDCDFEB6CB832941" blockId="108.[606,981,264,328]" box="[606,981,264,291]" pageId="108" pageNumber="109">
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<heading id="C969818C9D709237FDCDFEB6CB832941" box="[606,981,264,291]" centered="true" fontSize="11" level="2" pageId="108" pageNumber="109" reason="2">
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<taxonomicName id="559E4D639D709237FDCDFEB6CB832941" ID-CoL="3XFLF" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[606,981,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="simplex">
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||
<emphasis id="A0EAEAF29D709237FDCDFEB6CB072941" bold="true" box="[606,849,264,291]" italics="true" pageId="108" pageNumber="109">Maechidius simplex</emphasis>
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<bibRefCitation id="F60F4B119D709237FCCBFEB7CB832941" author="Frey G." box="[856,981,265,291]" pageId="108" pageNumber="109" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
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</taxonomicName>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="DA84656B9D709237FD42FE93CB352925" box="[721,867,301,328]" pageId="108" pageNumber="109" type="description">
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<paragraph id="922136E09D709237FD42FE93CB352925" blockId="108.[606,981,264,328]" box="[721,867,301,328]" pageId="108" pageNumber="109">
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<figureCitation id="0AA52A659D709237FD42FE93CB7D292A" box="[721,811,301,328]" captionStart="Figs 82–85" captionStartId="106.[189,241,1894,1920]" captionTargetBox="[330,1255,264,1870]" captionTargetId="figure-67@106.[330,1258,252,1883]" captionTargetPageId="106" captionText="Figs 82–85. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 82. M. similis sp. nov., paratype, ♂ (BMNH). 83. M. skalei sp. nov., paratype, ♂ (NME). 84. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). 85. M. sougb sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122165" httpUri="https://zenodo.org/record/4122165/files/figure.png" pageId="108" pageNumber="109">Figs 84</figureCitation>
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,
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<figureCitation id="0AA52A659D709237FCA4FE93CB352925" box="[823,867,301,327]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="108" pageNumber="109">168</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DA84656B9D709237FF2EFECECAC02A4A" pageId="108" pageNumber="109" type="materials_examined">
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<paragraph id="922136E09D709237FF2EFECEC93A29E8" blockId="108.[189,364,368,394]" box="[189,364,368,394]" pageId="108" pageNumber="109">
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<heading id="C969818C9D709237FF2EFECEC93A29E8" bold="true" box="[189,364,368,394]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
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<emphasis id="A0EAEAF29D709237FF2EFECEC93A29E8" bold="true" box="[189,364,368,394]" pageId="108" pageNumber="109">Type material</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="922136E09D709237FF79FE1EC90A29D8" blockId="108.[189,1399,416,552]" box="[234,348,416,442]" pageId="108" pageNumber="109">
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<emphasis id="A0EAEAF29D709237FF79FE1EC90A29D8" bold="true" box="[234,348,416,442]" pageId="108" pageNumber="109">
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||
<typeStatus id="4D2588429D709237FF79FE1EC90A29D8" box="[234,348,416,442]" pageId="108" pageNumber="109" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
</paragraph>
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<paragraph id="922136E09D709237FF2EFE7BCAC02A4A" blockId="108.[189,1399,416,552]" pageId="108" pageNumber="109">
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<materialsCitation id="22F63CBD9D709237FF2EFE7BCAC02A4A" ID-GBIF-Occurrence="2898878498" collectionCode="WEST, NEW, TYPE, BPBM" collectorName="S. Quate & L. Quate Collectors" country="Indonesia" determinerName="G. Frey" elevation="1245" location="Star Mts." municipality="Mts. Sibil Val." pageId="108" pageNumber="109" specimenCount="2" typeStatus="holotype">
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<collectingCountry id="EA8976709D709237FF2EFE7BC90D29BD" box="[189,347,453,479]" name="Indonesia" pageId="108" pageNumber="109">INDONESIA</collectingCountry>
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•
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<specimenCount id="8498FD699D709237FEE6FE7BC9DB29BD" box="[373,397,453,479]" pageId="108" pageNumber="109">♂</specimenCount>
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; “
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<collectionCode id="F48FAE259D709237FE35FE7BC9AE29BD" box="[422,504,453,479]" pageId="108" pageNumber="109">WEST</collectionCode>
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<collectionCode id="F48FAE259D709237FD93FE7BCA1429BD" box="[512,578,453,479]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="108" pageNumber="109">NEW</collectionCode>
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<collectingCountry id="EA8976709D709237FDD9FE7BCAEF29BD" box="[586,697,453,479]" name="Guinea-Bissau" pageId="108" pageNumber="109">GUINEA</collectingCountry>
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<location id="9741603B9D709237FD52FE7BCB7A29BD" LSID="urn:lsid:plazi:treatment:1A3787F69D709237FDCDFEB6CB442D8F:9741603B9D709237FD52FE7BCB7A29BD" box="[705,812,453,479]" country="Indonesia" municipality="Mts. Sibil Val." name="Star Mts." pageId="108" pageNumber="109">
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Star
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<collectingMunicipality id="7245AC9A9D709237FD6AFE7BCB7A29BD" box="[761,812,453,479]" pageId="108" pageNumber="109">Mts.</collectingMunicipality>
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</location>
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Sibil Val.
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<quantity id="55669B059D709237FC30FE7ACBAE29BD" box="[931,1016,452,479]" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" pageId="108" pageNumber="109" unit="m" value="1245.0">
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<elevation id="19B3D1D39D709237FC30FE7ACBAE29BD" box="[931,1016,452,479]" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" pageId="108" pageNumber="109" unit="m" value="1245.0">1245m</elevation>
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</quantity>
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,
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<date id="E62010209D709237FB97FE7BCCDA29BD" box="[1028,1164,453,479]" bridgedPair="-" pageId="108" pageNumber="109">18-X-8.XI-</date>
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||
’61 [p] //
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<collectorName id="3F6B53369D709237FB61FE7BCD0129BD" box="[1266,1367,453,479]" pageId="108" pageNumber="109">S. Quate</collectorName>
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||
&
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<collectorName id="3F6B53369D709237FF2EFE54C9F02A61" box="[189,422,489,516]" pageId="108" pageNumber="109">L. Quate Collectors</collectorName>
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[p]//
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<typeStatus id="4D2588429D709237FE7BFE54CA672A66" box="[488,561,490,516]" pageId="108" pageNumber="109">
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<collectionCode id="F48FAE259D709237FE7BFE54CA672A66" box="[488,561,490,516]" pageId="108" pageNumber="109">TYPE</collectionCode>
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</typeStatus>
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[p, red label]//
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<typeStatus id="4D2588429D709237FD79FE54CB722A66" box="[746,804,490,516]" pageId="108" pageNumber="109">Type</typeStatus>
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||
<taxonomicName id="559E4D639D709237FCBEFE57CC4E2A61" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[813,1048,489,515]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="simplex" status="sp. nov.">
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<emphasis id="A0EAEAF29D709237FCBEFE57CC4E2A61" box="[813,1048,489,515]" italics="true" pageId="108" pageNumber="109">Maechidius simplex</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBD957899D709237FBB3FE54CC362A66" box="[1056,1120,490,516]" pageId="108" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
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[h] det.
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<determinerName id="67FDDA249D709237FB2EFE57CD472A61" box="[1213,1297,489,515]" pageId="108" pageNumber="109">G.Frey</determinerName>
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,1967/68 [p]”;
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<collectionCode id="F48FAE259D709237FF6FFDB0C9072A4A" box="[252,337,526,552]" collectionName="USA, Hawaii, Honolulu, Bernice P. Bishop Museum" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="108" pageNumber="109" type="Museum">BPBM</collectionCode>
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. Studied from photographs.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="DA84656B9D709237FF2EFDEEC9FD2D18" pageId="108" pageNumber="109" type="description">
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<paragraph id="922136E09D709237FF2EFDEEC9182A08" blockId="108.[189,334,592,618]" box="[189,334,592,618]" pageId="108" pageNumber="109">
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<heading id="C969818C9D709237FF2EFDEEC9182A08" bold="true" box="[189,334,592,618]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
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<emphasis id="A0EAEAF29D709237FF2EFDEEC9182A08" bold="true" box="[189,334,592,618]" pageId="108" pageNumber="109">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="922136E09D709237FF2EFD3ECB762D6A" blockId="108.[189,1399,640,1288]" pageId="108" pageNumber="109">
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Body brown, antennae and legs reddish-castaneous. Male labroclypeus subtruncate, its lateral margins slightly sinuous in dorsal view. Anterolateral angles of labroclypeus broadly rounded, not protruding (
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<figureCitation id="0AA52A659D709237FF55FD77C9642A86" box="[198,306,713,740]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="108" pageNumber="109">Fig. 167</figureCitation>
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). Canthus broadly rounded in dorsal view. Head punctures circular to irregularly ovoid, moderately deep, dense. Head setae short, scale-like, suberect, surpassing length of corresponding punctures. Pronotum broadly emarginate on anterior margin, basal margin slightly sinuous. Anterolateral angles rounded, slightly protruding. Lateral margin in dorsal view gradually widens towards obtuse postmedian angulation; constricted towards base from this point, delicately crenulate all along. Inconspicuous appressed scale-like seta present between every two crenulae. Pronotal punctures larger and deeper than those on head, in part annular. Intervening spaces subopaque, variably large, generally larger than punctures. Setation similar to on head, slightly longer. Scutellar shield narrowly rounded apically. Elytron with two rather broad flat glabrous longitudinal carinae and less distinct sutural carina. Preapical hump of elytron large but flat, somewhat Г- shaped (
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<figureCitation id="0AA52A659D709237FC7BFBACCC162C4F" box="[1000,1088,1042,1069]" captionStart="Figs 82–85" captionStartId="106.[189,241,1894,1920]" captionTargetBox="[330,1255,264,1870]" captionTargetId="figure-67@106.[330,1258,252,1883]" captionTargetPageId="106" captionText="Figs 82–85. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 82. M. similis sp. nov., paratype, ♂ (BMNH). 83. M. skalei sp. nov., paratype, ♂ (NME). 84. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). 85. M. sougb sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122165" httpUri="https://zenodo.org/record/4122165/files/figure.png" pageId="108" pageNumber="109">Fig. 84</figureCitation>
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). Elytral punctures sparse, moderately deep, irregularly ovoid. Intervening spaces subopaque, distinctly larger than punctures. Elytral setae inconspicuous, suberect; seta rises from anterior margin of each puncture, not or hardly surpassing its length. Male pygidium with long erect setae in apical part. Male protibia with nearly complete dorsal longitudinal furrow, with two large rounded distal teeth on external margin; distal tooth lobe-like prolonged anteriad. Male terminal protibial spur straight, long and acute. Male meso- and metatibia with interrupted median transverse carina.
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</paragraph>
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<paragraph id="922136E09D709237FF2EFA8EC9E62D28" blockId="108.[189,432,1328,1354]" box="[189,432,1328,1354]" pageId="108" pageNumber="109">
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||
<heading id="C969818C9D709237FF2EFA8EC9E62D28" bold="true" box="[189,432,1328,1354]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
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<emphasis id="A0EAEAF29D709237FF2EFA8EC9E62D28" bold="true" box="[189,432,1328,1354]" pageId="108" pageNumber="109">Sexual dimorphism</emphasis>
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</heading>
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</paragraph>
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<paragraph id="922136E09D709237FF2EFADEC9FD2D18" blockId="108.[189,427,1376,1402]" box="[189,427,1376,1402]" pageId="108" pageNumber="109">Female is unknown.</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D709237FF2EFA1DCB442D8F" box="[189,786,1443,1518]" pageId="108" pageNumber="109" type="discussion">
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<paragraph id="922136E09D709237FF2EFA1DC9722DDF" blockId="108.[189,292,1443,1469]" box="[189,292,1443,1469]" pageId="108" pageNumber="109">
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||
<heading id="C969818C9D709237FF2EFA1DC9722DDF" bold="true" box="[189,292,1443,1469]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
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||
<emphasis id="A0EAEAF29D709237FF2EFA1DC9722DDF" bold="true" box="[189,292,1443,1469]" pageId="108" pageNumber="109">Remark</emphasis>
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||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D709237FF2EFA6DCB442D8F" blockId="108.[189,786,1491,1518]" box="[189,786,1491,1518]" pageId="108" pageNumber="109">This species’ male genital organs were not studied.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |